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Introductory

WILDLIFE SPECIES: Scolopax minor | American Woodcock
ABBREVIATION : SCMI COMMON NAMES : American woodcock woodcock becasse wood snipe TAXONOMY : The commonly accepted scientific name for the American woodcock is Scolopax minor Gmelin. Until recently the genus was Philohela [1]. There are no subspecies. ORDER : Charadriiformes CLASS : Bird FEDERAL LEGAL STATUS : No special status OTHER STATUS : The American woodcock is listed by the U.S. Fish and Wildlife Service as a National Species of Special Emphasis [6]. COMPILED BY AND DATE : S. A. Snyder, December 1991 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Snyder, S. A. 1991. Scolopax minor. In: Remainder of Citation

WILDLIFE DISTRIBUTION AND OCCURRENCE

WILDLIFE SPECIES: Scolopax minor | American Woodcock
GENERAL DISTRIBUTION : Breeding range of the American woodcock extends from southern Manitoba east through Prince Edward Island and Nova Scotia and south through the eastern and central United States into Florida. Populations extend as far west as eastern Texas and Oklahoma. Winter range of the American woodcock is the southern portion of its breeding range, from southern Missouri south [2,6,8]. There are two populations of American woodcock: the Eastern and Central. Each is distinguished by its own migration corridor, but some mixing does occur [6]. The Eastern population breeds in New England and the Canadian Maritime Provinces and winters in the South Atlantic States. The Central population breeds in the Great Lake States and Canada and winters in the Gulf States. ECOSYSTEMS : FRES10 White-red-jack pine FRES11 Spruce-fir FRES12 Longleaf-slash pine FRES13 Loblolly-shortleaf pine FRES14 Oak-pine FRES15 Oak-hickory FRES16 Oak-gum-cypress FRES17 Elm-ash-cottonwood FRES18 Maple-beech-birch FRES19 Aspen-birch FRES23 Fir-spruce FRES41 Wet grasslands STATES :
AL AR CT DE FL GA IL IN IA KS
KY LA ME MD MA MI MN MS MO NH
NJ NM NY NC ND OH OK OR PA RI
SC SD TN TX UT VT VA WA WV WI

MB NB NF NS ON PE PQ
BLM PHYSIOGRAPHIC REGIONS : 14 Great Plains KUCHLER PLANT ASSOCIATIONS : K080 Marl - Everglades K081 Oak savanna K084 Cross Timbers K089 Blackbelt K090 Live oak - seas oats K091 Cypress savanna K092 Everglades K093 Great Lakes spruce - fir forest K099 Maple - basswood forest K100 Oak - hickory forest K101 Elm - ash forest K102 Beech - maple forest K103 Mixed mesophytic forest K104 Appalachian oak forest K106 Northern hardwoods K107 Northern hardwoods - fir forest K108 Northern hardwoods - spruce forest K109 Transition between K104 and K106 K110 Northeastern oak - pine forest K111 Oak - hickory - pine forest K112 Southern mixed forest K113 Southern floodplain forest K114 Pocosin K115 Sand pine scrub K116 Subtropical pine forest SAF COVER TYPES : 1 Jack pine 5 Balsam fir 12 Black spruce 13 Black spruce - tamarack 14 Northern pin oak 15 Red pine 16 Aspen 17 Pin cherry 18 Paper birch 19 Grey birch - red maple 20 White pine - northern red oak - red maple 21 Eastern white pine 22 White pine - hemlock 23 Eastern hemlock 24 Hemlock - yellow birch 25 Sugar maple - beech - yellow birch 26 Sugar maple - basswood 27 Sugar maple 28 Black cherry - maple 35 Paper birch - red spruce - balsam fir 38 Tamarack 39 Black ash - American elm - red maple 55 Northern red oak 60 Beech - sugar maple 61 River birch - sycamore 63 Cottonwood 64 Sassafras - persimmon 65 Pin oak - sweet gum 69 Sand pine 70 Longleaf pine 71 Longleaf pine - scrub oak 75 Shortleaf pine 76 Shortleaf pine - oak 80 Loblolly pine - shortleaf pine 81 Loblolly pine 82 Loblolly pine - hardwood 83 Longleaf pine - slash pine 84 Slash pine 85 Slash pine - hardwood 87 Sweet gum - yellow poplar 88 Willow oak - water oak - diamondleaf oak 89 Live oak 92 Sweetgum - willow oak 94 Sycamore - sweetgum - American elm 95 Black willow 101 Baldcypress 105 Tropical hardwoods 107 White spruce 108 Red maple 111 South Florida slash pine SRM (RANGELAND) COVER TYPES : NO-ENTRY PLANT COMMUNITIES : American woodcock inhabit both bottomland and upland plant communities. These include second-growth hardwood forests with aspen (Populus spp.) or alder (Alnus spp.), oak (Quercus spp;), hickory (Carya spp.), and maple (Acer spp.). They also inhabit spruce (Picea spp.)-fir (Abies spp.)-hardwood forests [9]. Southern pine plantations of loblolly pine (Pinus taeda), shortleaf pine (P. echinata), and longleaf pine (P. palustris) also provide habitat for the American woodcock [6]. REFERENCES : NO-ENTRY

BIOLOGICAL DATA AND HABITAT REQUIREMENTS

WILDLIFE SPECIES: Scolopax minor | American Woodcock
TIMING OF MAJOR LIFE HISTORY EVENTS : Mating season - late February through early May after male courtship display in January and February. Eggs - laid in ground nest shortly after mating; three to five per clutch; incubation is about 20 days. Maturity - chicks almost fully grown in 28 days; sexually mature in 10 to 12 months. Life Expectancy - about 8 years in captivity. Migration - in spring can begin as early as late February; in autumn can begin in September, usually determined by the first frost. [6,16,23,25] PREFERRED HABITAT : American woodcocks are classified as shorebirds and as such tend most often to frequent mesic woodlands, bottomlands, and wet meadows. They also inhabit recently logged or burned areas, agricultural land, and riparian woodlands [8]. American woodcocks show a high preference for aspen and alder stands and damp fields with little ground cover. The abundance of earthworms, the main staple of woodcock diets, strongly dictates American woodcock distribution [6]. Although hardwood forest soils support a greater abundance of earthworms, conifer forests do support shallow-burrowing worms which woodcocks will pursue, particularly if ground vegetation is sparse. American woodcocks nest in a variety of habitats such as open fields, mixed forests, bushfields, and conifer plantations but appear to prefer areas with high shrub-stem density [7]. Greg and Hale [13], however, found that most nests in Wisconsin were located in woodlands dominated by aspen. In Nova Scotia, American woodcocks seemed to prefer pastures with woody vegetation to those without for singing grounds. Feeding areas with cover or near forest edges were preferred over large open areas [26]. COVER REQUIREMENTS : American woodcocks need only moderate cover. Because woodcocks probe through soil for food, ground vegetation in foraging areas must be sparse [6]. Thick vegetation is more beneficial as hiding cover for predators than as escape cover for the woodcock. Open woodlands or brushy fields that provide patchy cover are used for roosting, courtship, nesting, and brood rearing. Open areas with patches of shrubs are used for singing grounds, where males begin courtship and claim territories. These areas must be open enough to enable the birds to carry-out their flight pattern displays [23]. FOOD HABITS : Earthworms (Lumbricus spp.; Aporrectodea spp.; Diplocardia spp.) make up 50 to 90 percent of the American woodcock's diet. Other food items include beetles (Coleoptera), flies (Diptera), centipedes (Chilopoda), butterfly and moth larvae (Lepidoptera), crustaceans, and other insect larvae [6,8]. Seeds of raspberry and blackberry (Rubus spp.), alder, elderberry (Sambucus spp.), cinquefoil (Potentilla spp.), dogwood (Cornus spp.), smartweed (Polygonum spp.), ragweed (Ambrosia spp.), sedge (Carex spp.), bedstraw (Galium spp.), and other plants are occasionally eaten [23]. PREDATORS : Predators include humans and hawks (Accipitridae) [14]. Although no specific information on other predators was found, many animals can prey on woodcock eggs, such as fox (Vulpes vulpes), skunks and weasels (Mustelidae), racoon (Procyon lotor), opossum (Didelphis virginiana), snakes, and birds of prey [23]. MANAGEMENT CONSIDERATIONS : American woodcocks are a popular game species in the East [2]. Much habitat manipulation is conducted to improve woodcock populations. Clearcutting aspen stands is a common and successful practice, usually leading to an influx of woodcocks within the first 2 years after harvest [3]. When these stands are cut, bare mineral soil is exposed and woodcocks can easily forage for earthworms [6]. However, because succession can take place rapidly following harvest of aspen stands, the benefits of cutting may be short-lived [15]. Aspen stand management strategies for improving American woodcock habitat have been outlined [22]. Wetland draining and pesticide use can both have a negative impact on woodcocks [6]. Draining wetlands reduces the water table and subsequently earthworm abundance. The American woodcock is extremely sensitive to pesticides, since the majority of their diet includes insects [5,24]. REFERENCES : NO-ENTRY

FIRE EFFECTS AND USE

WILDLIFE SPECIES: Scolopax minor | American Woodcock
DIRECT FIRE EFFECTS ON ANIMALS : Depending on season, fire could destroy American woodcock nests. HABITAT RELATED FIRE EFFECTS : Fires appear to have a beneficial effect on woodcock habitat. The most notable benefit is the reduction of ground vegetation, which facilitates foraging beneath the soil surface [6]. Fires can also raise the pH of soils, which can favor earthworm abundance [20]. One negative impact of fire may be the drying out of soil, which would drive earthworms deeper into the ground. Fire can also destroy important hiding and roosting cover [6,17]. Woodcock behavior studies have shown that woodcocks prefer burned to unburned areas for foraging, and often times are attracted to areas even as fires still burn [10,19,22]. FIRE USE : Fires can be used to clear vegetation for better foraging opportunities, as well as raise soil pH, which may promote earthworm abundance. However, adequate cover must be maintained for nesting, brood rearing, and roosting. REFERENCES : NO-ENTRY

REFERENCES

WILDLIFE SPECIES: Scolopax minor | American Woodcock
REFERENCES : 1. Donohoe, Robert W. 1974. American hornbeam Carpinus caroliniana Walt. In: Gill, John D.; Healy, William M., eds. Shrubs and vines for northeastern wildlife. Gen. Tech. Rep. NE-9. Upper Darby, PA: U.S. Department of Agriculture, Forest Service, Northeastern Forest Experiment Station: 86-88. [13714] 2. Artmann, Joseph W. 1977. Woodcock status report, 1975. Special Scientific Report - Wildlife No. 210. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 36 p. [16610] 3. Bennett, Carl L.; Rabe, Dale L.; Prince, Harold, H. 1982. Response of several species, with emphasis on woodcock, to extensive habitat manipulations. In: Dwyer, Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research Report 14. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service: 97-105. [16614] 4. Arno, Stephen F.; Gruell, George E. 1986. Douglas-fir encroachment into mountain grasslands in southwestern Montana. Journal of Range Management. 39(3): 272-276. [343] 5. Besadny, C. D. 1978. Upland game and waterfowl. In: Proceedings, 67th convention of international fish and wildlife agencies; 1977 September 12-14; Vancouver, BC. Washington, DC: International Association of Fish and Wildlife Agencies: 13-35. [16623] 6. Cade, Brian S. 1985. Habitat suitability index models: American woodcock (wintering). Biological Report 82 (10.105). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 23 p. [16611] 7. Johnson, Randall C.; Causey, M. Keith. 1982. Use of longleaf pine stands by woodcock in southern Alabama following prescribed burning. In: Dwyer, Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research Report 14. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service: 120-125. [16616] 8. DeGraaf, Richard M.; Scott, Virgil E.; Hamre, R. H.; [and others]. 1991. Forest and rangeland birds of the United States: Natural history and habitat use. Agric. Handb. 688. Washington, DC: U.S. Department of Agriculture, Forest Service. 625 p. [15856] 9. Dwyer, Thomas J.; Derlech, Eric L.; McAuley, Daniel G. 1982. Woodcock brood ecology in Maine. In: Dwyer, Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research Report 14. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service: 63-70. [16613] 10. Edwards, William R.; Ellis, J. A. 1958. Responses of three avian species to burning. Wilson Bulletin. 81(3): 338-339. [16619] 11. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 12. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 13. Gregg, Larry E.; Hale, James, B. 1977. Woodcock nesting habitat in northern Wisconsin. Auk. 94: 489-493. [16618] 14. Grzybowski, Joseph A.; Eaton, Stephen W. 1976. Prey items of goshawks in southwestern New York. Wilson Bulletin. 88: 669-670. [16625] 15. Hale, James B.; Gregg, Larry E. 1976. Woodcock use of clearcut aspen areas in Wisconsin. Wildlife Society Bulletin. 4(3): 111-115. [16622] 16. Hamel, Paul B.; LeGrand, Harry E., Jr.; Lennartz, Michael R.; Gauthreaux, Sidney A., Jr. 1982. Bird-habitat relationships on southeastern forest lands. Gen. Tech. Rep. SE-22. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station. 417 p. [15423] 17. Irving, Frank D., Jr. 1950. Some possible effects on wildlife of controlled burning in the pine types of Minnesota. Minneapolis, MN: University of Minnesota. 43 p. Thesis. [15913] 18. Johnsgard, Paul A. 1981. The plovers, sandpipers, and snipes of the world. Lincoln, NE: University of Nebraska. 493 p. [16617] 19. Johnson, Randall C.; Causey, M. Keith. 1982. Use of longleaf pine stands by woodcock in southern Alabama following prescribed burning. In: Dwyer, Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research Report 14. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service: 120-125. [16616] 20. Kozlowski, T. T.; Ahlgren, C. E., eds. 1974. Fire and ecosystems. New York: Academic Press. 542 p. [1374] 21. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 22. Sepik, Greg F.; Dwyer, Thomas J. 1982. Woodcock in response to habitat management in Maine. In: Dwyer, Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research Report 14. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service: 106-113. [16615] 23. Sheldon, William G. 1967. The book of the American woodcock. Amherst, MA: University of Massachusetts. 227 p. [16624] 24. Stickel, William H.; Hayne, D. W.; Stickel, Lucille F. 1965. Effects of heptachlor-contaminated earthworms on woodcocks. Journal of Wildlife Management. 29: 132-146. [16621] 25. Terres, John K. 1980. The Audubon Society encyclopedia of North American birds. New York: Alfred A. Knopf. 1109 p. [16195] 26. Wishart, Richard A.; Bider, J. Roger. 1976. Habitat preferences of woodcock in southwestern Quebec. Journal of Wildlife Management. 40: 523-531. [16620]

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