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Wildlife, Animals, and Plants
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Introductory
WILDLIFE SPECIES: Scolopax minor | American Woodcock
ABBREVIATION :
SCMI
COMMON NAMES :
American woodcock
woodcock
becasse
wood snipe
TAXONOMY :
The commonly accepted scientific name for the American woodcock is
Scolopax minor Gmelin. Until recently the genus was Philohela [1].
There are no subspecies.
ORDER :
Charadriiformes
CLASS :
Bird
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
The American woodcock is listed by the U.S. Fish and Wildlife Service as
a National Species of Special Emphasis [6].
COMPILED BY AND DATE :
S. A. Snyder, December 1991
LAST REVISED BY AND DATE :
NO-ENTRY
AUTHORSHIP AND CITATION :
Snyder, S. A. 1991. Scolopax minor. In: Remainder of Citation
WILDLIFE DISTRIBUTION AND OCCURRENCE
WILDLIFE SPECIES: Scolopax minor | American Woodcock
GENERAL DISTRIBUTION :
Breeding range of the American woodcock extends from southern Manitoba
east through Prince Edward Island and Nova Scotia and south through the
eastern and central United States into Florida. Populations extend as
far west as eastern Texas and Oklahoma. Winter range of the American
woodcock is the southern portion of its breeding range, from southern
Missouri south [2,6,8]. There are two populations of American woodcock:
the Eastern and Central. Each is distinguished by its own migration
corridor, but some mixing does occur [6]. The Eastern population breeds
in New England and the Canadian Maritime Provinces and winters in the
South Atlantic States. The Central population breeds in the Great Lake
States and Canada and winters in the Gulf States.
ECOSYSTEMS :
FRES10 White-red-jack pine
FRES11 Spruce-fir
FRES12 Longleaf-slash pine
FRES13 Loblolly-shortleaf pine
FRES14 Oak-pine
FRES15 Oak-hickory
FRES16 Oak-gum-cypress
FRES17 Elm-ash-cottonwood
FRES18 Maple-beech-birch
FRES19 Aspen-birch
FRES23 Fir-spruce
FRES41 Wet grasslands
STATES :
| AL |
AR |
CT |
DE |
FL |
GA |
IL |
IN |
IA |
KS |
| KY |
LA |
ME |
MD |
MA |
MI |
MN |
MS |
MO |
NH |
| NJ |
NM |
NY |
NC |
ND |
OH |
OK |
OR |
PA |
RI |
| SC |
SD |
TN |
TX |
UT |
VT |
VA |
WA |
WV |
WI |
BLM PHYSIOGRAPHIC REGIONS :
14 Great Plains
KUCHLER PLANT ASSOCIATIONS :
K080 Marl - Everglades
K081 Oak savanna
K084 Cross Timbers
K089 Blackbelt
K090 Live oak - seas oats
K091 Cypress savanna
K092 Everglades
K093 Great Lakes spruce - fir forest
K099 Maple - basswood forest
K100 Oak - hickory forest
K101 Elm - ash forest
K102 Beech - maple forest
K103 Mixed mesophytic forest
K104 Appalachian oak forest
K106 Northern hardwoods
K107 Northern hardwoods - fir forest
K108 Northern hardwoods - spruce forest
K109 Transition between K104 and K106
K110 Northeastern oak - pine forest
K111 Oak - hickory - pine forest
K112 Southern mixed forest
K113 Southern floodplain forest
K114 Pocosin
K115 Sand pine scrub
K116 Subtropical pine forest
SAF COVER TYPES :
1 Jack pine
5 Balsam fir
12 Black spruce
13 Black spruce - tamarack
14 Northern pin oak
15 Red pine
16 Aspen
17 Pin cherry
18 Paper birch
19 Grey birch - red maple
20 White pine - northern red oak - red maple
21 Eastern white pine
22 White pine - hemlock
23 Eastern hemlock
24 Hemlock - yellow birch
25 Sugar maple - beech - yellow birch
26 Sugar maple - basswood
27 Sugar maple
28 Black cherry - maple
35 Paper birch - red spruce - balsam fir
38 Tamarack
39 Black ash - American elm - red maple
55 Northern red oak
60 Beech - sugar maple
61 River birch - sycamore
63 Cottonwood
64 Sassafras - persimmon
65 Pin oak - sweet gum
69 Sand pine
70 Longleaf pine
71 Longleaf pine - scrub oak
75 Shortleaf pine
76 Shortleaf pine - oak
80 Loblolly pine - shortleaf pine
81 Loblolly pine
82 Loblolly pine - hardwood
83 Longleaf pine - slash pine
84 Slash pine
85 Slash pine - hardwood
87 Sweet gum - yellow poplar
88 Willow oak - water oak - diamondleaf oak
89 Live oak
92 Sweetgum - willow oak
94 Sycamore - sweetgum - American elm
95 Black willow
101 Baldcypress
105 Tropical hardwoods
107 White spruce
108 Red maple
111 South Florida slash pine
SRM (RANGELAND) COVER TYPES :
NO-ENTRY
PLANT COMMUNITIES :
American woodcock inhabit both bottomland and upland plant communities.
These include second-growth hardwood forests with aspen (Populus spp.)
or alder (Alnus spp.), oak (Quercus spp;), hickory (Carya spp.), and
maple (Acer spp.). They also inhabit spruce (Picea spp.)-fir (Abies
spp.)-hardwood forests [9]. Southern pine plantations of loblolly pine
(Pinus taeda), shortleaf pine (P. echinata), and longleaf pine (P.
palustris) also provide habitat for the American woodcock [6].
REFERENCES :
NO-ENTRY
BIOLOGICAL DATA AND HABITAT REQUIREMENTS
WILDLIFE SPECIES: Scolopax minor | American Woodcock
TIMING OF MAJOR LIFE HISTORY EVENTS :
Mating season - late February through early May after male courtship
display in January and February.
Eggs - laid in ground nest shortly after mating; three to five per clutch;
incubation is about 20 days.
Maturity - chicks almost fully grown in 28 days; sexually mature in 10
to 12 months.
Life Expectancy - about 8 years in captivity.
Migration - in spring can begin as early as late February; in
autumn can begin in September, usually determined
by the first frost.
[6,16,23,25]
PREFERRED HABITAT :
American woodcocks are classified as shorebirds and as such tend most
often to frequent mesic woodlands, bottomlands, and wet meadows. They
also inhabit recently logged or burned areas, agricultural land, and
riparian woodlands [8]. American woodcocks show a high preference for
aspen and alder stands and damp fields with little ground cover. The
abundance of earthworms, the main staple of woodcock diets, strongly
dictates American woodcock distribution [6]. Although hardwood forest
soils support a greater abundance of earthworms, conifer forests do
support shallow-burrowing worms which woodcocks will pursue,
particularly if ground vegetation is sparse.
American woodcocks nest in a variety of habitats such as open fields,
mixed forests, bushfields, and conifer plantations but appear to prefer
areas with high shrub-stem density [7]. Greg and Hale [13], however,
found that most nests in Wisconsin were located in woodlands dominated
by aspen.
In Nova Scotia, American woodcocks seemed to prefer pastures with woody
vegetation to those without for singing grounds. Feeding areas with
cover or near forest edges were preferred over large open areas [26].
COVER REQUIREMENTS :
American woodcocks need only moderate cover. Because woodcocks probe
through soil for food, ground vegetation in foraging areas must be
sparse [6]. Thick vegetation is more beneficial as hiding cover for
predators than as escape cover for the woodcock. Open woodlands or
brushy fields that provide patchy cover are used for roosting,
courtship, nesting, and brood rearing. Open areas with patches of
shrubs are used for singing grounds, where males begin courtship and
claim territories. These areas must be open enough to enable the birds
to carry-out their flight pattern displays [23].
FOOD HABITS :
Earthworms (Lumbricus spp.; Aporrectodea spp.; Diplocardia spp.) make up
50 to 90 percent of the American woodcock's diet. Other food items
include beetles (Coleoptera), flies (Diptera), centipedes (Chilopoda),
butterfly and moth larvae (Lepidoptera), crustaceans, and other insect
larvae [6,8]. Seeds of raspberry and blackberry (Rubus spp.), alder,
elderberry (Sambucus spp.), cinquefoil (Potentilla spp.), dogwood
(Cornus spp.), smartweed (Polygonum spp.), ragweed (Ambrosia spp.),
sedge (Carex spp.), bedstraw (Galium spp.), and other plants are
occasionally eaten [23].
PREDATORS :
Predators include humans and hawks (Accipitridae) [14]. Although no
specific information on other predators was found, many animals can prey
on woodcock eggs, such as fox (Vulpes vulpes), skunks and weasels
(Mustelidae), racoon (Procyon lotor), opossum (Didelphis virginiana),
snakes, and birds of prey [23].
MANAGEMENT CONSIDERATIONS :
American woodcocks are a popular game species in the East [2]. Much
habitat manipulation is conducted to improve woodcock populations.
Clearcutting aspen stands is a common and successful practice, usually
leading to an influx of woodcocks within the first 2 years after harvest
[3]. When these stands are cut, bare mineral soil is exposed and
woodcocks can easily forage for earthworms [6]. However, because
succession can take place rapidly following harvest of aspen stands, the
benefits of cutting may be short-lived [15]. Aspen stand management
strategies for improving American woodcock habitat have been outlined
[22].
Wetland draining and pesticide use can both have a negative impact on
woodcocks [6]. Draining wetlands reduces the water table and
subsequently earthworm abundance. The American woodcock is extremely
sensitive to pesticides, since the majority of their diet includes
insects [5,24].
REFERENCES :
NO-ENTRY
FIRE EFFECTS AND USE
WILDLIFE SPECIES: Scolopax minor | American Woodcock
DIRECT FIRE EFFECTS ON ANIMALS :
Depending on season, fire could destroy American woodcock nests.
HABITAT RELATED FIRE EFFECTS :
Fires appear to have a beneficial effect on woodcock habitat. The most
notable benefit is the reduction of ground vegetation, which facilitates
foraging beneath the soil surface [6]. Fires can also raise the pH of
soils, which can favor earthworm abundance [20]. One negative impact of
fire may be the drying out of soil, which would drive earthworms deeper
into the ground. Fire can also destroy important hiding and roosting
cover [6,17].
Woodcock behavior studies have shown that woodcocks prefer burned to
unburned areas for foraging, and often times are attracted to areas even
as fires still burn [10,19,22].
FIRE USE :
Fires can be used to clear vegetation for better foraging opportunities,
as well as raise soil pH, which may promote earthworm abundance.
However, adequate cover must be maintained for nesting, brood rearing,
and roosting.
REFERENCES :
NO-ENTRY
REFERENCES
WILDLIFE SPECIES: Scolopax minor | American Woodcock
REFERENCES :
1. Donohoe, Robert W. 1974. American hornbeam Carpinus caroliniana Walt.
In: Gill, John D.; Healy, William M., eds. Shrubs and vines for
northeastern wildlife. Gen. Tech. Rep. NE-9. Upper Darby, PA: U.S.
Department of Agriculture, Forest Service, Northeastern Forest
Experiment Station: 86-88. [13714]
2. Artmann, Joseph W. 1977. Woodcock status report, 1975. Special
Scientific Report - Wildlife No. 210. Washington, DC: U.S. Department of
the Interior, Fish and Wildlife Service. 36 p. [16610]
3. Bennett, Carl L.; Rabe, Dale L.; Prince, Harold, H. 1982. Response of
several species, with emphasis on woodcock, to extensive habitat
manipulations. In: Dwyer, Thomas J.; Storm, Gerald L., technical
coordinators. Wildlife Research Report 14. Washington, DC: U.S.
Department of the Interior, Fish and Wildlife Service: 97-105. [16614]
4. Arno, Stephen F.; Gruell, George E. 1986. Douglas-fir encroachment into
mountain grasslands in southwestern Montana. Journal of Range
Management. 39(3): 272-276. [343]
5. Besadny, C. D. 1978. Upland game and waterfowl. In: Proceedings, 67th
convention of international fish and wildlife agencies; 1977 September
12-14; Vancouver, BC. Washington, DC: International Association of Fish
and Wildlife Agencies: 13-35. [16623]
6. Cade, Brian S. 1985. Habitat suitability index models: American woodcock
(wintering). Biological Report 82 (10.105). Washington, DC: U.S.
Department of the Interior, Fish and Wildlife Service. 23 p. [16611]
7. Johnson, Randall C.; Causey, M. Keith. 1982. Use of longleaf pine stands
by woodcock in southern Alabama following prescribed burning. In: Dwyer,
Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research
Report 14. Washington, DC: U.S. Department of the Interior, Fish and
Wildlife Service: 120-125. [16616]
8. DeGraaf, Richard M.; Scott, Virgil E.; Hamre, R. H.; [and others]. 1991.
Forest and rangeland birds of the United States: Natural history and
habitat use. Agric. Handb. 688. Washington, DC: U.S. Department of
Agriculture, Forest Service. 625 p. [15856]
9. Dwyer, Thomas J.; Derlech, Eric L.; McAuley, Daniel G. 1982. Woodcock
brood ecology in Maine. In: Dwyer, Thomas J.; Storm, Gerald L.,
technical coordinators. Wildlife Research Report 14. Washington, DC:
U.S. Department of the Interior, Fish and Wildlife Service: 63-70.
[16613]
10. Edwards, William R.; Ellis, J. A. 1958. Responses of three avian species
to burning. Wilson Bulletin. 81(3): 338-339. [16619]
11. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. [905]
12. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. [998]
13. Gregg, Larry E.; Hale, James, B. 1977. Woodcock nesting habitat in
northern Wisconsin. Auk. 94: 489-493. [16618]
14. Grzybowski, Joseph A.; Eaton, Stephen W. 1976. Prey items of goshawks in
southwestern New York. Wilson Bulletin. 88: 669-670. [16625]
15. Hale, James B.; Gregg, Larry E. 1976. Woodcock use of clearcut aspen
areas in Wisconsin. Wildlife Society Bulletin. 4(3): 111-115. [16622]
16. Hamel, Paul B.; LeGrand, Harry E., Jr.; Lennartz, Michael R.;
Gauthreaux, Sidney A., Jr. 1982. Bird-habitat relationships on
southeastern forest lands. Gen. Tech. Rep. SE-22. Asheville, NC: U.S.
Department of Agriculture, Forest Service, Southeastern Forest
Experiment Station. 417 p. [15423]
17. Irving, Frank D., Jr. 1950. Some possible effects on wildlife of
controlled burning in the pine types of Minnesota. Minneapolis, MN:
University of Minnesota. 43 p. Thesis. [15913]
18. Johnsgard, Paul A. 1981. The plovers, sandpipers, and snipes of the
world. Lincoln, NE: University of Nebraska. 493 p. [16617]
19. Johnson, Randall C.; Causey, M. Keith. 1982. Use of longleaf pine stands
by woodcock in southern Alabama following prescribed burning. In: Dwyer,
Thomas J.; Storm, Gerald L., technical coordinators. Wildlife Research
Report 14. Washington, DC: U.S. Department of the Interior, Fish and
Wildlife Service: 120-125. [16616]
20. Kozlowski, T. T.; Ahlgren, C. E., eds. 1974. Fire and ecosystems. New
York: Academic Press. 542 p. [1374]
21. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. [1384]
22. Sepik, Greg F.; Dwyer, Thomas J. 1982. Woodcock in response to habitat
management in Maine. In: Dwyer, Thomas J.; Storm, Gerald L., technical
coordinators. Wildlife Research Report 14. Washington, DC: U.S.
Department of the Interior, Fish and Wildlife Service: 106-113. [16615]
23. Sheldon, William G. 1967. The book of the American woodcock. Amherst,
MA: University of Massachusetts. 227 p. [16624]
24. Stickel, William H.; Hayne, D. W.; Stickel, Lucille F. 1965. Effects of
heptachlor-contaminated earthworms on woodcocks. Journal of Wildlife
Management. 29: 132-146. [16621]
25. Terres, John K. 1980. The Audubon Society encyclopedia of North American
birds. New York: Alfred A. Knopf. 1109 p. [16195]
26. Wishart, Richard A.; Bider, J. Roger. 1976. Habitat preferences of
woodcock in southwestern Quebec. Journal of Wildlife Management. 40:
523-531. [16620]
Index
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