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Introductory

SPECIES: Antennaria racemosa | Raceme Pussytoes
ABBREVIATION : ANTRAC SYNONYMS : NO-ENTRY SCS PLANT CODE : ANRA COMMON NAMES : raceme pussytoes TAXONOMY : The currently accepted scientific name of raceme pussytoes is Antennaria racemosa Hook. [6,8,13,15]. There are no recognized subspecies, varieties, or forms. Antennaria is a complex genus due to a high degree of apomixis, polyploidy, and hybridization [11]. LIFE FORM : Forb FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Robin F. Matthews, November 1993 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Matthews, Robin F. 1993. Antennaria racemosa. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Antennaria racemosa | Raceme Pussytoes
GENERAL DISTRIBUTION : Raceme pussytoes is distributed from British Columbia south to Oregon and east to Alberta, Montana, and Wyoming [13,15]. ECOSYSTEMS : FRES20 Douglas-fir FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce FRES25 Larch FRES26 Lodgepole pine STATES : ID MT OR WA WY AB BC ADMINISTRATIVE UNITS : BIHO BICA CRLA GLAC GRTE MORA NOCA OLYM YELL BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 5 Columbia Plateau 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K002 Cedar - hemlock - Douglas-fir forest K004 Fir - hemlock forest K012 Douglas-fir forest K014 Grand fir - Douglas-fir forest K015 Western spruce - fir forest SAF COVER TYPES : 205 Mountain hemlock 206 Engelmann spruce - subalpine fir 210 Interior Douglas-fir 212 Western larch 213 Grand fir 218 Lodgepole pine 224 Western hemlock 227 Western redcedar - western hemlock 228 Western redcedar 229 Pacific Douglas-fir 230 Douglas-fir - western hemlock SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Some species commonly associated with raceme pussytoes include bearberry (Arctostaphylos uva-ursi), prince's pine (Chimaphila umbellata), twinflower (Linnaea borealis), grouse whortleberry (Vaccinium scoparium), heartleaf arnica (Arnica cordifolia), western yarrow (Achillea millefolium), fireweed (Epilobium angustifolium), fleabane (Erigeron spp.), showy aster (Aster conspicuus), dandelion (Taraxacum officinale), scarlet paintbrush (Castilleja miniata), Virginia strawberry (Fragaria virginiana), and pinegrass (Calamagrostis rubescens) [3,12,16].

VALUE AND USE

SPECIES: Antennaria racemosa | Raceme Pussytoes
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Raceme pussytoes has moderate forage value for deer [21]. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : In Douglas-fir (Pseudotsuga menziesii)/pinegrass habitats in central Idaho, raceme pussytoes cover decreased after clearcutting and shelterwood cutting followed by mechanical scarification; after clearcutting followed by broadcast burning; and after clearcutting with no subsequent site preparation [21]. It showed a similiar response to the same silvicultural treatments in Douglas-fir/ninebark (Physocarpus malvaceus) and Douglas-fir/globe huckleberry (Vaccinium globulare) habitat types in western Montana [2]. Raceme pussytoes is moderately resistant to trampling [5]. Pussytoes (Antennaria spp.) usually increase in response to grazing [25].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Antennaria racemosa | Raceme Pussytoes
GENERAL BOTANICAL CHARACTERISTICS : Raceme pussytoes is a native, perennial forb with creeping, leafy stolons. Stems are 4 to 24 inches (10-60 cm) tall. The inflorescence is a generally open cyme, but flowers may be more crowded at higher elevations [13,15]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Raceme pussytoes reproduces from seed or spreads vegetatively through stolons. Seeds are light and wind dispersed. They are generally not stored in soil seedbanks. Raceme pussytoes seeds germinate on mineral soil in partial shade; germination and seedling establishment do not increase dramatically following disturbance. Vegetative growth tends to increase following partial removal of the canopy [21]. SITE CHARACTERISTICS : Raceme pussytoes is found on moist, at least partially shaded sites in cool montane and subalpine forests [6,13,15]. In west-central Montana it is particularly abundant on north slopes, where it may be found along roads and in other disturbed places [15]. Raceme pussytoes is found at elevations of 5,600 to 11,500 feet (1,700-3,500 m) in Wyoming and 3,500 to 6,800 feet (1,050-2,050 m) in Montana [7]. SUCCESSIONAL STATUS : Facultative Seral Species Raceme pussytoes is shade tolerant but is often found on disturbed sites on north-facing slopes [15]. In central Idaho it is an early seral species in subalpine fir (Abies lasiocarpa)/beargrass (Xerophyllum tenax) habitat types [19], but is considered late seral in Douglas-fir/ninebark and Douglas-fir/pinegrass habitat types [20,21]. SEASONAL DEVELOPMENT : Raceme pussytoes flowers from May through August [13].

FIRE ECOLOGY

SPECIES: Antennaria racemosa | Raceme Pussytoes
FIRE ECOLOGY OR ADAPTATIONS : Raceme pussytoes colonizes bare mineral soil from light, wind-dispersed seed [19,21]. POSTFIRE REGENERATION STRATEGY : Initial-offsite colonizer (off-site, initial community) Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Antennaria racemosa | Raceme Pussytoes
IMMEDIATE FIRE EFFECT ON PLANT : Raceme pussytoes is probably killed by most fires. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Raceme pussytoes response to fire varies. It decreased in cover in the first years following stand-replacing wildfires in Douglas-fir/pinegrass, Douglas-fir/ninebark, and Douglas-fir/globe huckleberry habitat types in central Idaho and western Montana [2,21]. However, it increased after fires in subalpine fir/beargrass habitats and was often found in the understory of open lodgepole pine (Pinus contorta) stands, especially where one or more light surface fires had recently occurred [19]. Raceme pussytoes was present in increasing numbers in the first 10 years following the Pattee Canyon Fire in western Montana [24]. Raceme pussytoes was also present within 7 to 9 years following a broadcast burn in a western larch (Larix occidentalis)-Douglas-fir forest in Montana, although it was not a component of the prefire community [22]. Reese [18] stated that raceme pussytoes is sometimes present on recently burned sites in the Teton Wilderness, Wyoming. Anderson [1] reported that raceme pussytoes was present "via vegetative regrowth" in postfire year 1 following moderately severe fires in lodgepole pine stands in Yellowstone National Park. No other information was found on the ability of raceme pussytoes to sprout after fire. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Antennaria racemosa | Raceme Pussytoes
REFERENCES : 1. Anderson, Jay E.; Romme, William H. 1991. Initial floristics in lodgepole pine (Pinus contorta) forests following the 1988 Yellowstone fires. International Journal of Wildland Fire. 1(2): 119-124. [16008] 2. Arno, Stephen F.; Simmerman, Dennis G.; Keane, Robert E. 1985. Forest succession on four habitat types in western Montana. Gen. Tech. Rep. INT-177. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 74 p. [349] 3. Basile, Joseph V.; Jensen, Chester E. 1971. Grazing potential on lodgepole pine clearcuts in Montana. Res. Pap. INT-98. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 11 p. [8280] 4. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 5. Cole, David N. 1988. Disturbance and recovery of trampled montane grassland and forests in Montana. Res. Pap. INT-389. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 37 p. [3622] 6. Cronquist, Arthur. 1955. Vascular plants of the Pacific Northwest: Part 5: Compositae. Seattle: University of Washington Press. 343 p. [716] 7. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 8. Dorn, Robert D. 1988. Vascular plants of Wyoming. Cheyenne, WY: Mountain West Publishing. 340 p. [6129] 9. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 10. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 11. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 12. Hann, Wendel John. 1982. A taxonomy for classification of seral vegetation of selected habitat types in western Montana. Moscow, ID: University of Idaho. 235 p. Dissertation. [1073] 13. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 14. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 15. Lackschewitz, Klaus. 1991. Vascular plants of west-central Montana--identification guidebook. Gen. Tech. Rep. INT-227. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 648 p. [13798] 16. La Roi, George H.; Hnatiuk, Roger J. 1980. The Pinus contorta forests of Banff and Jasper National Parks: a study in comparative synecology and syntaxonomy. Ecological Monographs. 50(1): 1-29. [8347] 17. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 18. Reese, Jerry B.; Mohr, Francis R.; Dean, Ronald E.; Klabunde, Thomas. 1976. Teton Wilderness fire management plan. Part I: Ecological and resource description of the Teton Wilderness. Jackson, WY: U.S. Department of Agriculture, Forest Service, Bridger-Teton National Forest. 123 p. [21064] 19. Simpson, Michael L. 1990. The subalpine fir/beargrass habitat type: Succession and management. Moscow, ID: University of Idaho. 134 p. Thesis. [13464] 20. Steele, Robert; Geier-Hayes, Kathleen. 1989. The Douglas-fir/ninebark habitat type in central Idaho: succession and management. Gen. Tech. Rep. INT-252. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 65 p. [8136] 21. Steele, Robert; Geier-Hayes, Kathleen. 1993. The Douglas-fir/pinegrass habitat type in central Idaho: succession and management. Gen. Tech. Rep. INT-298. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 83 p. [21512] 22. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 23. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 24. Toth, Barbara L. 1991. Factors affecting conifer regeneration and community structure after a wildfire in western Montana. Corvallis, OR: Oregon State University. 124 p. Thesis. [14425] 25. Tweit, Susan J.; Houston, Kent E. 1980. Grassland and shrubland habitat types of the Shoshone National Forest. Cody, WY: U.S. Department of Agriculture, Forest Service, Shoshone National Forest. 143 p. [2377] 26. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573]

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