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Introductory

SPECIES: Convolvulus arvensis | Field Bindweed
ABBREVIATION : CONARV SYNONYMS : Convolvulus ambigens House Convolvulus incanus Vahl Strophocaulos arvensis (L.) Small SCS PLANT CODE : COAR4 COMMON NAMES : field bindweed bindweed European bindweed creeping jenny wild morning-glory orchard morning-glory TAXONOMY : The currently accepted name of field bindweed is Convolvulus arvensis L., in the morning-glory family (Convolvulaceae). There are no recognized subspecies or varieties. Three forms based on leaf shape are used infrequently [11]: C. a. f. arvensis C. a. f. cordifolius Lasch C. a. f. auriculatus Desr. LIFE FORM : Forb FEDERAL LEGAL STATUS : No special status OTHER STATUS : Field bindweed has been declared a noxious weed in 48 states, the District of Columbia, and the southern provinces of Canada [28,40]. COMPILED BY AND DATE : Diane S. Pavek, October 1992. LAST REVISED BY AND DATE : AUTHORSHIP AND CITATION : Pavek, Diane S. 1992. Convolvulus arvensis. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Convolvulus arvensis | Field Bindweed
GENERAL DISTRIBUTION : Field bindweed is a native of southern Europe and naturalized throughout the United States and Canada [11]. In the United States, field bindweed's range extends from California [26]; throughout the Pacific Northwest [16], Intermountain region [8,9,14,41], and Great Plains [13,21]; south into Arizona [19] and Texas [4]; and in all states east of the Mississippi River [11,17,29,34,36,38]. It is adventitious is Hawaii [43]. Field bindweed occurs in the southern provinces of Canada from the East Coast across the plains to the West Coast [33]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES12 Longleaf - slash pine FRES13 Loblolly - shortleaf pine FRES14 Oak - pine FRES15 Oak - hickory FRES16 Oak - gum - cypress FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES19 Aspen - birch FRES20 Douglas-fir FRES21 Ponderosa pine FRES22 Western white pine FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce FRES25 Larch FRES26 Lodgepole pine FRES27 Redwood FRES28 Western hardwoods FRES29 Sagebrush FRES30 Desert shrub FRES31 Shinnery FRES32 Texas savanna FRES33 Southwestern shrubsteppe FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES36 Mountain grasslands FRES37 Mountain meadows FRES38 Plains grasslands FRES39 Prairie FRES40 Desert grasslands FRES41 Wet grasslands FRES42 Annual grasslands STATES : AL AZ AR CA CO CT DE FL GA HI ID IL IN IA KS KY LA ME MD MA MI MN MS MO MT NE NV NH NJ NM NY NC ND OH OK OR PA RI SC SD TN TX UT VT VA WA WV WI WY AB BC MB NB NS ON PE PQ SK MEXICO ADMINISTRATIVE UNITS : ACAD BADL BITH BICA BLCA BRCA CACH CARE CATO COLO COLM CODA CUVA DETO FLFO GATE GWCA GLAC GLCA GRCA GRTE GRKO GRBA GUMO INDU JECA JODA LAME LAMR LAVO LABE MANA MEVE MOCA MORU NABR NERI NOCA OLYM PEFO PINN PORE REDW ROCR SAMO SCBL SLBE THRO TICA VAFO WICR WICA YELL ZION BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K025 Alder - ash forest K026 Oregon oakwoods K028 Mosaic of K002 and K026 K029 California mixed evergreen forest K030 California oakwoods K031 Oak - juniper woodlands K047 Fescue - oatgrass K048 California steppe K050 Fescue - wheatgrass K051 Wheatgrass - bluegrass K053 Grama - galleta steppe K054 Grama - tobosa prairie K055 Sagebrush steppe K056 Wheatgrass - needlegrass shrubsteppe K057 Galleta - three-awn shrubsteppe K058 Grama - tobosa shrubsteppe K059 Trans-Pecos shrub savanna K060 Mesquite savanna K061 Mesquite - acacia savanna K062 Mesquite - live oak savanna K063 Foothills prairie K064 Grama - needlegrass - wheatgrass K065 Grama - buffalograss K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem - needlegrass K068 Wheatgrass - grama - buffalograss K069 Bluestem - grama prairie K070 Sandsage - bluestem prairie K071 Shinnery K074 Bluestem prairie K075 Nebraska Sandhills prairie K076 Blackland prairie K077 Bluestem - sacahuista prairie K078 Southern cordgrass prairie K079 Palmetto prairie K081 Oak savanna K082 Mosaic of K074 and K100 K084 Cross Timbers K088 Fayette prairie K089 Black Belt K090 Live oak - sea oats K098 Northern floodplain forest K099 Maple - basswood forest K100 Oak - hickory forest K101 Elm - ash forest K102 Beech - maple forest K103 Mixed mesophytic forest K104 Appalachian oak forest K106 Northern hardwoods K107 Northern hardwoods - fir forest K108 Northern hardwoods - spruce forest K109 Transition between K104 and K106 K110 Northeastern oak - pine forest K111 Oak - hickory - pine forest K112 Southern mixed forest K113 Southern floodplain forest SAF COVER TYPES : 1 Jack pine 15 Red pine 16 Aspen 17 Pin cherry 18 Paper birch 20 White pine - northern red oak - red maple 21 Eastern white pine 22 White pine - hemlock 23 Eastern hemlock 24 Hemlock - yellow birch 25 Sugar maple - beech - yellow birch 26 Sugar maple - basswood 27 Sugar maple 28 Black cherry - maple 30 Red spruce - yellow birch 31 Red spruce - sugar maple - beech 32 Red spruce 33 Red spruce - balsam fir 34 Red spruce - Fraser fir 35 Paper birch - red spruce - balsam fir 37 Northern white-cedar 39 Black ash - American elm - red maple 40 Post oak - blackjack oak 42 Bur oak 43 Bear oak 44 Chestnut oak 45 Pitch pine 50 Black locust 51 White pine - chestnut oak 52 White oak - black oak - northern red oak 53 White oak 55 Northern red oak 57 Yellow-poplar 58 Yellow-poplar - eastern hemlock 59 Yellow-poplar - white oak - northern red oak 60 Beech - sugar maple 61 River birch - sycamore 62 Silver maple - American elm 63 Cottonwood 64 Sassafras - persimmon 70 Longleaf pine 71 Longleaf pine - scrub oak 73 Southern redcedar 76 Shortleaf pine - oak 78 Virginia pine - oak 80 Loblolly pine - shortleaf pine 81 Loblolly pine 82 Loblolly pine - hardwood 83 Longleaf pine - slash pine 84 Slash pine 85 Slash pine - hardwood 88 Willow oak - water oak - diamondleaf oak 89 Live oak 93 Sugarberry - American elm - green ash 98 Pond pine 108 Red maple 110 Black oak 111 South Florida slash pine 210 Interior Douglas-fir 211 White fir 212 Western larch 213 Grand fir 215 Western white pine 216 Blue spruce 217 Aspen 218 Lodgepole pine 219 Limber pine 220 Rocky Mountain juniper 229 Pacific Douglas-fir 230 Douglas-fir - western hemlock 232 Redwood 233 Oregon white oak 234 Douglas-fir - tanoak - Pacific madrone 235 Cottonwood - willow 236 Bur oak 237 Interior ponderosa pine 238 Western juniper 239 Pinyon - juniper 240 Arizona cypress 241 Western live oak 243 Sierra Nevada mixed conifer 244 Pacific ponderosa pine - Douglas-fir 245 Pacific ponderosa pine 246 California black oak 247 Jeffrey pine 248 Knobcone pine 249 Canyon live oak 250 Blue oak - Digger pine 255 California coast live oak SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Field bindweed is more indicative of disturbed areas than natural systems. Field bindweed is listed as a dominant forb in 12.4 percent of the sample sites in a riverine United States Fish and Wildlife Service deep water-wetland classification. The main factors listed as influencing field bindweed occurrence were livestock and wildlife grazing and recreation use [27]. In Wyoming it is a dominanat herbaceous plant in the blue spruce (Picea pugens)-lodgepole pine (Pinus contorta)-white fir (Abies concolor) subtype of the coniferous overstory habitat type. The publication that lists field bindweed as a dominant is: A physical and biological characterization of riparian habitat and its importance to wildlife in Wyoming [27].

VALUE AND USE

SPECIES: Convolvulus arvensis | Field Bindweed
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Field bindweed is consumed by ungulates, small mammals, and birds [1,5,18]. It comprised 1.7 percent of mule deer summer diet on a central Colorado ponderosa pine (Pinus ponderosa)-bunchgrass range [5]. Analysis of rumen samples showed that field bindweed was consumed by 30 percent of the individuals in one white-tailed deer population in Montana [1]. PALATABILITY : The palatability of field bindweed for wildlife species in Montana and Utah is rated as follows: MT UT Pronghorn fair good Elk -- good Mule deer -- good White-tailed deer poor good Small mammals poor fair Small non-game birds poor fair Upland game birds poor fair Waterfowl -- fair NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : Field bindweed provides poor environmental protection during one or more seasons for wild ungulates, game and small nongame birds, and small mammals in Utah. VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : Field bindweed is one of the ten worst weeds in the world [6,7,18,37]. Efforts must be made to prevent field bindweed from establishing on disturbed sites. Soil pasteurization is recommended to prepare a clean seedbed for native plant production [2]. Long-lasting control of field bindweed can be achieved with an integrated pest management program utilizing biological, mechanical, and chemical methods [28]. Tillage has been less successful as a control method and should be combined with fall cover crop seedings [28]. Biological Control: Research is progressing on various fungal pathogens (Altenaria brassicae, Bartalinia robillardoides, Myrothecium roridum) from India for field bindweed control [32,35]. In one study, the fungus Phomopsis convolvulus killed 95 percent of field bindweed seedlings [24]. Insects (e.g., Galeruca rufa, Eriophyes sp., Spermophagus sericeus, Erysiphe convolvuli) from Europe have been introduced with mixed levels of control [31]. Chemical Control: Chemical control has been used successfully against mature field bindweed plants [28]. Chemical choice depends upon (1) extent of infestation, (2) use of land where growing, and (3) proximity to water or likelihood of irrigation. Some effective chemicals used are 2,4-D, dicamba, picloram, or chlorates [28]. Seeds are impervious to water and to water-soluble chemicals (i.e., herbicides). After herbaceous growth is killed, seed bank reserves will continue to establish infestations [37]. Therefore, seedlings must be controlled; they are susceptible to 2,4-D or glyphosate [37].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Convolvulus arvensis | Field Bindweed
GENERAL BOTANICAL CHARACTERISTICS : Field bindweed has prostrate to climbing stems that grow from 7 to 80 inches (20-200 cm) long [16,34]. The main root system can penetrate 20 feet (6 m) deep [7,28]. It has extensive and deeply descending rhizomes. Leaves are variable in form but are more or less arrow shaped and 1 to 2 inches (2-6 cm) long [11]. Fruits are egg-shaped capsules that split open when mature, each yielding two to four dark brown seeds [16,28,29]. RAUNKIAER LIFE FORM : Geophyte REGENERATION PROCESSES : Field bindweed spreads by seeds and rhizomes. Seeds may live in the soil for 30 to 50 years before germinating [7,28,37]. Seeds of field bindweed are not readily cleaned from crop seeds and are spread through unclean seed or feed [28]. Additionally, seeds are spread when livestock move from infested grazing areas to weed-free land [28]. The hard seed coats must be scarified before germination occurs; seeds remain viable after passing through animal digestive tracts [18]. While vernalization at 41 degrees Fahrenheit (5 deg C) increased germination rates, it was not necessary for germination to occur in laboratory trials [18]. Seeds optimally germinated at 84 degrees Fahrenheit (29 deg C) with 24 hours of darkness; however, the seeds will germinate at tempertures ranging from freezing (34 degrees Fahrenheit [1 deg C]) to above 100 degrees Fahrenheit (38 deg C) [37]. Rhizomes grow rapidly, creating dense colonies [28]. SITE CHARACTERISTICS : Field bindweed occurs and is competitive in disturbed ground: waste places, roadsides, railroad right-of-ways, fields, gardens, and open river banks [4,11,14,23,26]. Field bindweed often occurs on well-drained soils and also is found on shallow, less productive soils [23,28,42]. It occurs on a wide variety of soil textural classes, from loams to clays [23]. Field bindweed is found from sea level to 7,500 feet (2,286 m) [26,27]. Some associated species found in disturbed riparian areas were mugwort (Artemisia vulgaris), common dandelion (Taraxacum officinale), and bird vetch (Vicia cracca) [6]. A dominant species found with field bindweed in degraded, wet meadow margins was medusahead (Taeniatherum asperum) [42]. SUCCESSIONAL STATUS : Obligate Initial Community Species Field bindweed occurs as a component of ruderal flora in disturbed areas that are rich in introduced species [23]. Dall'Armellina and Zimdahl [6] reported that reductions in light intensity resulted in significant reductions in shoot and root dry matter. Field bindweed cannot tolerate shade and uses its viney stems to move into sunlight. Therefore, depending upon community physiognomy, it is unlikely that field bindweed persists in later stages of community succession. SEASONAL DEVELOPMENT : Field bindweed flowers from late April to October throughout its range [4,13,16,29]. Seeds mature within 2 weeks after pollination during hot summer days; germination occurs in the fall and spring [37]. Foliage dies with the first heavy frost. Rhizomes overwinter and readily sprout in the spring [28].

FIRE ECOLOGY

SPECIES: Convolvulus arvensis | Field Bindweed
FIRE ECOLOGY OR ADAPTATIONS : Field bindweed has deeply growing rhizomes that would escape injury or death from fire. Vegetative growth will rapidly sprout from these protected rhizomes [15]. POSTFIRE REGENERATION STRATEGY : Rhizomatous herb, rhizome in soil Initial-offsite colonizer (off-site, initial community) Geophyte, growing points deep in soil

FIRE EFFECTS

SPECIES: Convolvulus arvensis | Field Bindweed
IMMEDIATE FIRE EFFECT ON PLANT : While prescribed fires are used in agriculture for weed control and seedbed preparation, no studies have been done exclusively on field bindweed. Fire top-kills field bindweed [15]. Its deep rhizomes would survive most fires; however, Degennaro and Weller [7] found 70 percent of the roots of field bindweed were in the top 6 inches (15 cm) of soil. Low to moderately severe fires could set back plant growth or kill those plants without extensive rhizomes. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Field bindweed sprouts from surviving rhizomes after fire has killed the vegetative plant parts [15]. Plants would sprout after spring or summer burning and continue growing until fall frosts. After vegetative growth is restored, rhizomes extend and continue to create dense colonies in any open habitat [15]. Since field bindweed seed can germinate after several decades, buried seed that survives fire would sprout whenever it is brought to the surface in succeeding years [37]. Because seed survives digestive tracts, small mammals and birds could transport seed onto a burned site. In addition to seed scarification occurring by stomach acids, fire scarification of surviving seed could enhance or promote germination. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : In North Dakota, a 30 acre (12 ha) field was prepared for planting to native grasses by burning and summer fallowing for 2 years [15]. Within 2 months after burning, perennial weeds, including field bindweed, were very dense. Follow-up treatment with 1 year of herbicide (2,4-D) application was not successful. A second year of herbicide treatment was planned. FIRE MANAGEMENT CONSIDERATIONS : Prescribed fires do not successfully control field bindweed. An integrated approach should include herbicide applications and/or soil pasteurization to achieve control [28,37].

REFERENCES

SPECIES: Convolvulus arvensis | Field Bindweed
REFERENCES : 1. Allen, Eugene O. 1968. Range use, foods, condition, and productivity of white-tailed deer in Montana. Journal of Wildlife Management. 32(1): 130-141. [16331] 2. Bainbridge, David A. 1990. Soil solarization for restorationists. Restoration & Management Notes. 8(2): 96-98. [14160] 3. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 4. Correll, Donovan S.; Johnston, Marshall C. 1970. Manual of the vascular plants of Texas. Renner, TX: Texas Research Foundation. 1881 p. [4003] 5. Currie, P. O.; Reichert, D. W.; Malechek, J. C.; Wallmo, O. C. 1977. Forage selection comparisons for mule deer and cattle under managed ponderosa pine. Journal of Range Management. 30(5): 352-356. [4697] 6. Dall'Armellina, Armando A.; Zimdahl, Robert L. 1988. Effect of light on growth and development of field bindweed (Convolvulus arvensis) and Russian knapweed (Centaurea repens). Weed Science. 36: 779-783. [6487] 7. Degennaro, Francis P.; Weller, Stephen C. 1984. Growth and reproductive characteristics of field bindweed (Convolvulus arvensis) biotypes. Weed Science. 32: 525-528. [19136] 8. Dorn, Robert D. 1977. Willows of the Rocky Mountain States. Rhodora. 79: 390-429. [6000] 9. Dorn, Robert D. 1984. Vascular plants of Montana. Cheyenne, WY: Mountain West Publishing. 276 p. [819] 10. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 11. Fernald, Merritt Lyndon. 1950. Gray's manual of botany. [Corrections supplied by R. C. Rollins]. Portland, OR: Dioscorides Press. 1632 p. (Dudley, Theodore R., gen. ed.; Biosystematics, Floristic & Phylogeny Series; vol. 2). [14935] 12. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 13. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 14. Harrington, H. D. 1964. Manual of the plants of Colorado. 2d ed. Chicago: The Swallow Press Inc. 666 p. [6851] 15. Higgins, Kenneth F.; Duebbert, Harold F.; Jacobson, Erling T. 1983. Successful establishment of stands of cool-season native grasses (North Dakota). Restoration & Management Notes. 1(2): 12. [11647] 16. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 17. Jones, G. N.; Fuller, G. D. 1955. Vascular plants of Illinois. Urbana, IL: University of Illinois Press. 593 p. [18964] 18. Jordan, L. S.; Jordan, J. L. 1982. Effects of pre-chilling on Convolvulus arvensis L. seed coat and germination. Annals of Botany. 49: 421-423. [19137] 19. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2d ed. Berkeley, CA: University of California Press. 1085 p. [6563] 20. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 21. Lakela, O. 1965. A flora of northeastern Minnesota. Minneapolis, MN: University of Minnesota Press. 541 p. [18142] 22. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 23. Morin, Edith; Bouchard, Andre; Jutras, Pierre. 1989. Ecological analysis of disturbed riverbanks in the Montreal area of Quebec. Environmental Management. 13(2): 215-225. [13233] 24. Morin, L.; Watson, A. K.; Reeleder, R. D. 1989. Efficacy of Phomopsis convolvulus for control of field bindweed (Convolvulus arvensis). Weed Science. 37(6): 830-835. [19145] 25. Muenscher, W. C. 1940. Poisonous plants of the United States. New York: MacMillan Co. 266 p. [18141] 26. Munz, Philip A. 1973. A California flora and supplement. Berkeley, CA: University of California Press. 1905 p. [6155] 27. Olson, R. A.; Gerhart, W. A. 1982. A physical and biological characterization of riparian habitat and its importance to wildlife in Wyoming. Cheyenne, WY: Wyoming Game and Fish Department. 188 p. [6755] 28. Pacific Northwest Extension. 1974. Field bindweed. PNW 115. Corvallis, OR; Pullman, WA; Moscow, ID. 3 p. [6611] 29. Radford, Albert E.; Ahles, Harry E.; Bell, C. Ritchie. 1968. Manual of the vascular flora of the Carolinas. Chapel Hill, NC: The University of North Carolina Press. 1183 p. [7606] 30. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 31. Rosenthal, S. S. 1981. European organisms of interest for the biological control of Convolvulus arvensis in the United States. In: Del Fosse, E. S., ed. Proceedings, 5th International symposium on biological control of weeds. Melbourne, Australia: CSIRO: 537-544. [19139] 32. Saharan, G. S.; Kaushik, J. C.; Kaushik, C. D. 1982. Two new host records of Alternaria brassicae; Angallis arvensis and Convolvulus arvensis, India. Indian Phytophathology. 35(1): 172. [19140] 33. Scoggan, H. J. 1978. The flora of Canada. Ottawa, Canada: National Museums of Canada. (4 volumes). [18143] 34. Seymour, Frank Conkling. 1982. The flora of New England. 2d ed. Phytologia Memoirs 5. Plainfield, NJ: Harold N. Moldenke and Alma L. Moldenke. 611 p. [7604] 35. Srivastava, R. C.; Srivastava, H. C. 1980. Two new leaf spot diseases, Bartalinia robillardoides, Myrothecium roridum from India on Fradescantia virginis and Confolculus arvensis. Indian Journal of Mycol. Plant Path. 10(2): 172. [19141] 36. Steyermark, J. A. 1963. Flora of Missouri. Ames, IA: Iowa State University Press. 1725 p. [18144] 37. Swan, D. G. 1983. Regeneration of field bindweed (Convolvulus arvensis) seedlings. Weeds Today. 14(4): 3-4. [19138] 38. Tidestrom, I.; Kittell, T. 1941. A flora of Arizona and New Mexico. Washington, DC: The Catholic University of America Press. 897 p. [18145] 39. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 40. U.S. Department of the Interior, Bureau of Land Management, Oregon State Office. 1985. Final Northwest Area noxious weed control program environmental impact statement. Portland, OR. 295 p. [12796] 41. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 42. Young, James A.; Evans, Raymond A.; Eckert, Richard E., Jr. 1969. Wheatgrass establishment with tillage and herbicides in a mesic medusahead community. Journal of Range Management. 22: 151-155. [2666] 43. St. John, Harold. 1973. List and summary of the flowering plants in the Hawaiian islands. Hong Kong: Cathay Press Limited. 519 p. [25354]

Index

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