|
Wildlife, Animals, and Plants
|
|
Introductory
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
ABBREVIATION :
HEDALPA
SYNONYMS :
Hedysarum americanum
SCS PLANT CODE :
HEALA
COMMON NAMES :
American sweetvetch
American hedysarum
alpine sweetvetch
purple hedysarum
TAXONOMY :
The accepted scientific name of American sweetvetch is Hedysarum alpinum
var. americanum Michx. Two other varieties of Hedysarum alpinum are
recognized. Hedysarum alpinum var. grandiflorum Rollins has a similar
range as H. a. var. alpinum but has larger flowers (14-18 mm compared to
11-15 mm). Hedysarum alpinum var. philoscia (A. Nels.) Rollins occurs
to the south and southeast of the other two varieties in Wyoming, South
Dakota, and Alberta, and has pubescent fruit [10].
LIFE FORM :
Forb
FEDERAL LEGAL STATUS :
No special status
OTHER STATUS :
American sweetvetch is rare in Glacier National Park [16].
COMPILED BY AND DATE :
Tara Y. Williams, July 1990
LAST REVISED BY AND DATE :
NO-ENTRY
AUTHORSHIP AND CITATION :
Williams, T. Y. 1990. Hedysarum alpinum var. americanum. In: Remainder of Citation
DISTRIBUTION AND OCCURRENCE
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
GENERAL DISTRIBUTION :
American sweetvetch is a circumpolar species. In the western
hemisphere, it occurs from Alaska through western Canada to British
Columbia and Saskatchewan, south to north-central Montana, Wyoming, and
South Dakota. It also occurs as far east as Labrador and New England
[20,21].
ECOSYSTEMS :
FRES11 Spruce - fir
FRES23 Fir - spruce
FRES26 Lodgepole pine
FRES36 Mountain grasslands
FRES37 Mountain meadows
FRES44 Alpine
STATES :
AK MA ME MT ND SD VT WY AB BC
NT SK YT
ADMINISTRATIVE UNITS :
DENA GLAC LACL WRST
BLM PHYSIOGRAPHIC REGIONS :
8 Northern Rocky Mountains
9 Middle Rocky Mountains
16 Upper Missouri Basin and Broken Lands
KUCHLER PLANT ASSOCIATIONS :
K008 Lodgepole pine - subalpine forest
K015 Western spruce - fir forest
K050 Fescue - wheatgrass
K052 Alpine meadows and barren
SAF COVER TYPES :
201 White spruce
202 White spruce - paper birch
204 Black spruce
206 Engelmann spruce - subalpine fir
218 Lodgepole pine
251 White spruce - aspen
253 Black spruce - white spruce
SRM (RANGELAND) COVER TYPES :
NO-ENTRY
HABITAT TYPES AND PLANT COMMUNITIES :
American sweetvetch has been listed as a dominant in a plant association
in the following classification:
The vegetation of Alberta [16]
Associated spceies: American sweetvetch commonly occurs wih American
green alder (Alnus viridis ssp. crispa), bog birch (Betula glandulosa),
tufted hairgrass (Deschampsia caespitosa), Bering hairgrass (D.
beringensis), rough fescue (Festuca scabrella), northern bedstraw
(Galium boreale), shrubby cinquefoil (Potentilla fruticosa), balsam
poplar (Populus balsamifera), lodgepole pine (Pinus contorta), white
spruce (Picea glauca), and willows (Salix spp.) [3,4,7,11,16,17].
VALUE AND USE
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
WOOD PRODUCTS VALUE :
NO-ENTRY
IMPORTANCE TO LIVESTOCK AND WILDLIFE :
Roots of another sweetvetch, Hedysarum sulphurescens, are an important
early spring food for grizzly bears [23]. However, American sweetvetch
is probably not abundant enough in Glacier National Park to be an
important forage for grizzlies [1,15].
PALATABILITY :
Other members of the genus Hedysarum are rated as palatable to sheep and
fairly so for cattle during midsummer and early fall [1]. Roots of
Hedysarym sulphurescens are highly palatable to grizzly bears. American
sweetvetch, however, is high in tannins, which may lower its
palatability [1].
NUTRITIONAL VALUE :
A nutritive analysis of American sweetvetch showed that the mean percent
digestible protein was 2.4 [3]. On a nutritive value index, where
clipped dried alfalfa (Medicago sativa) had the top score of 100,
American sweetvetch rated a mean score of 12.3, which was 80 percent
lower than the overall mean of five other forbs analyzed in the study
[3]. Another study showed very different results. Protein and fiber
quantities were similar to those of alfalfa, but American sweetvetch was
much higher in tannins than other forage legumes [1].
COVER VALUE :
An American sweetvetch community in Alaska was the primary nesting site
for a population of dusky Canada geese. It was also the nesting site
for several other waterfowl and birds. A population of voles was
restricted to the area and nested in and fed on the sweetvetch.
VALUE FOR REHABILITATION OF DISTURBED SITES :
American sweetvetch has been recommended for use in revegetation
projects in Alaska. After one growing season at a subalpine site at
2,483 feet (757 m), sweetvetch seedlings showed 73 percent survival with
added topsoil and 87 percent survival without added topsoil.
Transplanting was much more successful using book-style containers than
tube-style containers [5]. In the tundra zone of the Northwest
Territories, Canada, American sweetvetch is a prominent species (about
10 percent cover) on many sites in borrow pits alongside an abandoned
highway [11].
OTHER USES AND VALUES :
American sweetvetch forms nitrogen-fixing root nodules [6]. It is also
useful as bee fodder, as it is freely visited by honey bees [1].
MANAGEMENT CONSIDERATIONS :
American sweetvetch has some potential for use as a forage species for
roadsides and small spring pastures or as a hay crop [1]. For
cultivation purposes, agronomic selection for a strain lower in tannins
is suggested. Specimens from seeds collected at different locations
showed highly variable tannin quantities. Strains with lower tannin
content would presumably be more palatable. American sweetvetch is also
susceptible to a leaf spotting fungus and verticillium wilt, so
resistant strains should be sought [1].
American sweetvetch showed similar percent cover and frequency following
clearcutting as in a control. Abundance was slightly lower after
shelterwood cutting [22].
American sweetvetch recovers slowly after cutting and produces little
regrowth compared to alfalfa [1].
BOTANICAL AND ECOLOGICAL CHARACTERISTICS
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
GENERAL BOTANICAL CHARACTERISTICS :
American sweetvetch is a native, perennial forb with a woody taproot.
It grows in tufts, 8 to 30 inches (20-75 cm) high, with long, erect
stems. The perfect flowers are papilionaceous and vary from pink to
purple. The fruit is a flat loment, constricted into more or less oval,
indehiscent segments with four to six seeds [1,10,15,20].
RAUNKIAER LIFE FORM :
Chamaephyte
REGENERATION PROCESSES :
American sweetvetch reproduces sexually. The flowers are pollinated by
honey bees. The erect stems aid in natural seed dispersal by gravity
and wind. Field seed has shown a germination rate of 97 to 100 percent.
Scarification increases germination rate [1]. Plants grow in groups of
a few to hundreds of individuals [1,11].
SITE CHARACTERISTICS :
American sweetvetch grows in meadows and on slopes from low to fairly
high elevations [15]. It occurs on shale slides, roadside verges, and
forest fringes [1,20], and often occurs near water, such as on sand
bars, point bars, limey rivershores, lakeshores, and near seashores
between beach and forest [4,17,20,21]. It can grow on shallow, poorly
developed, rocky, dry, calcareous soils or in cold, wet, acidic, organic
substrates [4,7,16,17]. It does not do well in poorly drained, bog-type
woodlands [17]. American sweetvetch may occur in wetlands, although it
usually does not [19]. It survives in areas that occasionally flood
[17].
American sweetvetch is found at the following elevations [5,6,7,13].
1,720 to 2,480 feet (525-757 m) in AK
3,770 to 4,260 feet (1150-1300 m) in AB
5,000 feet (1525 m) in MT
8,100 to 8,200 feet (2470-2500 m) in WY
SUCCESSIONAL STATUS :
NO-ENTRY
SEASONAL DEVELOPMENT :
American sweetvetch begins to grow in May, earlier than other perennial
legumes. It produces buds in June and flowers in June and July [1].
The seeds ripen in July and August; the pods ripen unevenly. The
seedlings and mature plants are moderately tolerant to spring and fall
frosts, suffering some winterkill [1].
FIRE ECOLOGY
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
FIRE ECOLOGY OR ADAPTATIONS :
American sweetvetch establishes on a burned site via wind-carried seed
[1]. Often it is present on moist sites where fire is not a great
hazard [17].
POSTFIRE REGENERATION STRATEGY :
Initial-offsite colonizer (off-site, initial community)
FIRE EFFECTS
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
IMMEDIATE FIRE EFFECT ON PLANT :
American sweetvetch plants are killed by fire [22].
DISCUSSION AND QUALIFICATION OF FIRE EFFECT :
NO-ENTRY
PLANT RESPONSE TO FIRE :
On Willow Island (near Fairbanks, Alaska) in a white spruce/thinleaf
alder (Alnus incana ssp. tenuifolia)/feathermoss (Hylocomium splendens)
forest type, American sweetvetch showed much lower percent cover and
frequency following clearcutting and burning than in a control. There
was not such a reduction after clearcutting alone. The species was
present in the stand within two growing seasons after the burn treatment
[22].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE :
NO-ENTRY
FIRE MANAGEMENT CONSIDERATIONS :
NO-ENTRY
REFERENCES
SPECIES: Hedysarum alpinum var. americanum | American Sweetvetch
REFERENCES :
1. Bassendowski, K. A.; Smith, J. Drew; Howarth, R. E. 1989. The potential
value of Hedysarum alpinum var. americanum as a forage legume for the
northern Canadian prairies. Canadian Journal of Plant Science. 69:
815-822. [11487]
2. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals,
reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's
associations for the eleven western states. Tech. Note 301. Denver, CO:
U.S. Department of the Interior, Bureau of Land Management. 169 p.
[434]
3. Bezeau, L. M.; Johnston, A. 1962. In vitro digestibility of range forage
plants of the Festuca scabrella association. Canadian Journal of Plant
Science. 42: 692-697. [441]
4. Chapman, Kim Alan. 1986. Alpine hedysarum (Hedysarum alpinum) discovered
in Michigan. Michigan Botanist. 25(4): 45-46. [11493]
5. Densmore, R. V.; Holmes, K. W. 1987. Assisted revegetation in Denali
National Park, Alaska, U.S.A. Arctic and Alpine Research. 19(4):
544-548. [6078]
6. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information
network (PIN) data base: Colorado, Montana, North Dakota, Utah, and
Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior,
Fish and Wildlife Service. 786 p. [806]
7. Edwards, M. E.; Armbruster, W. S. 1989. A tundra-steppe transition on
Kathul Mountain, Alaska, U.S.A. Arctic and Alpine Research. 21(3):
296-304. [9673]
8. Eyre, F. H., ed. 1980. Forest cover types of the United States and
Canada. Washington, DC: Society of American Foresters. 148 p. [905]
9. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others].
1977. Vegetation and environmental features of forest and range
ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of
Agriculture, Forest Service. 68 p. [998]
10. Hitchcock, C. Leo; Cronquist, Arthur. 1961. Vascular plants of the
Pacific Northwest. Part 3: Saxifragaceae to Ericaceae. Seattle, WA:
University of Washington Press. 614 p. [1167]
11. Kershaw, G. Peter; Kershaw, Linda J. 1987. Successful plant colonizers
on disturbances in tundra areas of northwestern Canada. Arctic and
Alpine Research. 19(4): 451-460. [6115]
12. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation
of the conterminous United States. Special Publication No. 36. New York:
American Geographical Society. 77 p. [1384]
13. La Roi, George H.; Hnatiuk, Roger J. 1980. The Pinus contorta forests of
Banff and Jasper National Parks: a study in comparative synecology and
syntaxonomy. Ecological Monographs. 50(1): 1-29. [8347]
14. Stickney, Peter F. 1989. Seral origin of species originating in northern
Rocky Mountain forests. Unpublished draft on file at: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station, Fire
Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090]
15. Lesica, Peter. 1984. Rare vascular plants of Glacier National Park,
Montana. Missoula, MT: University of Montana, Department of Botany. 27
p. [12049]
16. Moss, E. H. 1955. The vegetation of Alberta. Botanical Review. 21(9):
493-567. [6878]
17. Pearce, C. M.; McLennan, D.; Cordes, L. D. 1988. The evolution and
maintenance of white spruce woodlands on the Mackenzie Delta, N. W. T.,
Canada. Holarctic Ecology. 11(4): 248-258. [10472]
18. Raunkiaer, C. 1934. The life forms of plants and statistical plant
geography. Oxford: Clarendon Press. 632 p. [2843]
19. Reed, Porter B., Jr. 1988. National list of plant species that occur in
wetlands: Alaska (Region A). Biological Report 88(26.11). Washington,
DC: U.S Department of the Interior, Fish and Wildlife Service. In
cooperation with: National and Regional Interagency Review Panels. 86 p.
[9328]
20. Standley, Paul C. 1921. Flora of Glacier National Park, Montana.
Contributions from the United States National Herbarium. Vol. 22, Part
5. Washington, DC: United States National Museum, Smithsonian
Institution: 235-438. [12318]
21. Seymour, Frank Conkling. 1982. The flora of New England. 2d ed.
Phytologia Memoirs 5. Plainfield, NJ: Harold N. Moldenke and Alma L.
Moldenke. 611 p. [7604]
22. Dyrness, C. T.; Viereck, L. A.; Foote, M. J.; Zasada, J. C. 1988. The
effect on vegetation and soil temperature of logging flood-plain white
spruce. Res. Pap. PNW-RP-392. Portland, OR: U.S. Department of
Agriculture, Forest Service, Pacific Northwest Research Station. 45 p.
[7471]
23. McLellan, B. N. 1986. Use-availability analysis and timber selection by
grizzly bears. In: Contreras, Glen P.; Evans, Keith E., compilers.
Proceedings--grizzly bear habitat symposium; 1985 April 30 - May 2;
Missoula, MT. Gen. Tech. Rep. INT-207. Ogden, UT: U.S. Department of
Agriculture, Forest Service, Intermountain Research Station: 163-166.
[14527]
24. U.S. Department of Agriculture, Soil Conservation Service. 1982.
National list of scientific plant names. Vol. 1. List of plant names.
SCS-TP-159. Washington, DC. 416 p. [11573]
Index
Related categories for Species: Hedysarum alpinum var. americanum
| American Sweetvetch
|
 |