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Introductory

SPECIES: Taraxacum officinale | Dandelion
ABBREVIATION : TAROFF SYNONYMS : NO-ENTRY SCS PLANT CODE : TAOF COMMON NAMES : dandelion common dandelion TAXONOMY : The currently accepted scientific name for dandelion is Taraxacum officinale Weber [134]. There are no recognized subspecies, varieties, or forms. LIFE FORM : Forb FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Lora L. Esser, February 1993 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Esser, Lora L. 1993. Taraxacum officinale. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Taraxacum officinale | Dandelion
GENERAL DISTRIBUTION : Dandelion is of Eurasian origin but has become naturalized throughout the United States. It occurs in all 50 states, almost all Canadian provinces, and Mexico [62,126]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES12 Longleaf - slash pine FRES13 Loblolly - shortleaf pine FRES14 Oak - pine FRES15 Oak - hickory FRES16 Oak - gum - cypress FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES19 Aspen - birch FRES20 Douglas-fir FRES21 Ponderosa pine FRES22 Western white pine FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce FRES25 Larch FRES26 Lodgepole pine FRES27 Redwood FRES28 Western hardwoods FRES29 Sagebrush FRES30 Desert shrub FRES32 Texas savanna FRES33 Southwestern shrubsteppe FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES36 Mountain grasslands FRES37 Mountain meadows FRES38 Plains grasslands FRES39 Prairie FRES40 Desert grasslands FRES41 Wet grasslands FRES42 Annual grasslands FRES44 Alpine STATES : AL AK AZ AR CA CO CT DE FL GA HI ID IL IN IA KS KY LA ME MD MA MI MN MS MO MT NE NV NH NJ NM NY NC ND OH OK OR PA RI SC SD TN TX UT VT VA WA WV WI WY AB BC MB NB NF NT NS ON PQ SK YT MEXICO ADMINISTRATIVE UNITS : ACAD AGFO ALPO ANTI APIS ARCH ASIS BADL BAND BISO BITH BICA BLCA BLRI BRCA BUFF CACH CACO CAHA CARE CACA CATO CEBR CHCH CHIR COLO COLM COSW CODA CUGA CUVA DEVA DEWA DETO EFMO FIIS DINO FLFO FODO FOBU GATE GWCA GWMP GLAC GRCA GRTE GRKO GRBA GRSA GRSM GUMO GUIS HALE HOBE INDU ISRO JECA JOFL LACL LAME LAMR LAVO MACA MEVE MORA MORU NERI NOCA OBRI OLYM PIRO PINN PIPE PORE REDW RICH ROCR ROMO SAJH SAMO SARA SCBL SHEN SHIL SLBE SUCR THRO TICA VAFO VOYA WACA WICA YELL YOSE ZION BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : Dandelion is found in nearly all Kuchler Plant Associations. SAF COVER TYPES : Dandelion is found in nearly all SAF cover types. SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Dandelion is an indicator species in ruderal vegetation types in North Dakota, South Dakota, and Washington [51,137].

VALUE AND USE

SPECIES: Taraxacum officinale | Dandelion
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Dandelion is a preferred food of domestic sheep grazing on mountain meadows [83] and is readily eaten by cattle on rough fescue (Festuca scabrella) prairies in Alberta [38]. Dandelion is commonly eaten in the spring by sharp-tailed grouse [89]. It is a minor component of bighorn sheep diets in the Upper Yellowstone Valley [63] and is an important food for pocket gophers on mountain grasslands of Colorado [132]. Dandelion is an important source of nectar and pollen for bees in Alaska [96]. Dandelion is consumed by deer and elk in the spring, summer, and fall in meadows of the Rocky Mountains [73]. In Yellowstone National Park, dandelion is an important food for grizzly bears in summer. Peak use in in June [82]. Leaves, stems, seeds, and flowers were found in grizzly and black bear scats in Glacier National Park [65]. In Alberta, black bears browse on earlier phenological stages of dandelion (spring and early summer) because of the higher nutrient quality. Dandelion is one of the dominant species found in spring bear scats [52]. During prenesting through incubation of greater prairie chicken broods (April-May) on the Sheyenne National Grasslands in North Dakota, dandelion flowers were one of the primary diet items. Individual fecal samples contained up to 96 percent dandelion flowers during April and May [106]. Dandelion is one of the favored foods of sage grouse in the spring, summer, and fall in Nevada. Of all meadow forbs consumed, dandelion contributed 82 percent to spring forb diets [40,67]. In British Columbia, deer consumed dandelion at significantly higher (P<0.05) rates on harvested lodgepole pine sites than on unharvested sites [28]. PALATABILITY : Dandelion is more palatable to wildlife and livestock in prebloom stages than in postbloom stages [81]. It is poor to fair in palatability on ponderosa pine sites throughout the West [85]. Palatability ratings for dandelion from selected western states are as follows [37]: UT CO WY MT ND Cattle good good fair fair good Sheep good good good good good Horses good good fair good good Elk good ---- good good ---- Mule deer good ---- good fair fair White-tailed deer ---- ---- good fair fair Pronghorn good ---- good good fair Upland game birds good ---- good good good Waterfowl fair ---- poor ---- good Small nongame birds fair ---- fair fair fair Small mammals good ---- fair fair fair NUTRITIONAL VALUE : Protein content of dandelion exceeds the minimum requirement needed for body maintenance for deer in ponderosa pine communities [94]. Dandelion meets the nutritional requirements of beef cattle in Alberta [16]. Protein and manganese content increase from early June to early July, when it is harvested on ranges in Alberta. By late September, protein content decreases significantly [16]. Chemical composition (in percent) of dandelion from an irrigated pasture during 1986 was as follows [16]: June 3 July 7 September 24 Average Acid detergent fiber 28.1 22.4 25.8 25.4 Crude protein 13.8 22.8 14.7 17.1 Ca 1.21 1.55 1.61 1.46 P 0.30 0.48 0.29 0.36 Mg 0.31 0.47 0.50 0.43 K 2.58 2.24 2.46 2.43 COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Dandelion has low short-term and long-term revegetation potential on disturbed sites. Erosion-control potential is low [37]. OTHER USES AND VALUES : The Gwich'in Athabaskan Indians of Fort Yukon, Alaska frequently eat the leaves of dandelion in salads or boil and eat them [54]. Roots of dandelion can be ground and used as a mild laxative or to treat heartburn. Tea and wine can be made from flowers [140]. MANAGEMENT CONSIDERATIONS : Dandelion is an invader species that commonly inhabits overgrazed rangelands [85]. Dandelion availability for deer decreases on cattle-grazed sites [7]. Dandelion meets the nutritional requirements of beef cattle and is readily grazed by them [16]. Producers may want to control dandelion in irrigated pastures to restrict seed movement to adjacent land where dandelion may be undesirable [16]. Dandelion is a threat in upper forest and alpine zones of western Montana because of its ability to invade little disturbed or undisturbed native vegetation through seed dispersal [133]. In Montana, dandelion seedlings compete with conifer seedlings on forest sites. Grass seeding on these sites will eventually decrease the dandelion population in 4 to 5 years [14]. Clearcuts and thinning of forests stimulates dandelion production. Sage grouse and deer populations benefit from increased production of dandelion [10]. Sage grouse habitat loss due to development and postdevelopment land use can be minimized by regulation of livestock on important adjacent nondeveloped lands [10]. Dandelion can be readily controlled with 2,4-D. It is most effective to spray in early spring before first bloom. Sites should not be mown for 3 to 5 days before spraying or 1 to 2 days after [92]. Strip spraying in Idaho in relatively high annual precipitation (13 inches [33 cm]) areas benefits sage grouse brood-rearing habitat due to quick recovery of dandelion and other forbs. Average cover of dandelion in sprayed areas was 17.2 percent, whereas average cover in nonsprayed areas was 11.2 percent [23]. A decrease in the population of dandelion occurs where pocket gophers are present. When gophers were removed, dandelion population increased by 50 percent in 2 years on mountain grasslands and meadows of Colorado, Utah, and Oregon [42].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Taraxacum officinale | Dandelion
GENERAL BOTANICAL CHARACTERISTICS : Dandelion is an introduced, cool-season, perennial forb [140]. It has a thick taproot up to 6 inches (15.2 cm) long [135]. Stems are very short and wholly underground, producing a rosette of leaves at the ground surface. Leaves are 2 to 16 inches (5-40 cm) long [134]. The flower heads are solitary at the end of naked, hollow stalks. Stalks can reach heights up to 2 feet (60 cm) [126,135]. One head contains from 100 to 300 flowers [126]. Seeds of dandelion are topped by a parachute of bristles that aid in dissemination [55]. Dandelion forms vesicular-arbuscular mycorrhizal associations [15,37]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Dandelion reproduces apomictically through parthenogenesis [62]. Plants develop from unfertilized gametes. Dandelion is an aggressive seed producer and reproduces mainly from seed [42]. Seeds travel a considerable distance because of the parachuting effect produced by the spreading pappus. In a tallgrass pairie in Iowa, achenes of dandelion were blown by the wind several hundred meters from the nearest source population [98]. Dandelion creates a long-lived seedbank [11,99]. In a seedbank of a ponderosa pine community in Washington, viable dandelion seedlings emerged from litter and soil samples in greenhouse germination trials. Seed density of spring samples was 160 seeds per square yard (133 seeds/m sq) and of autumn samples was 60 seeds per square yard (50 seeds/m sq) [99]. Seeds of dandelion were viable up to 5 years in soil samples from Montana [11]. Seed germination on a control plot in Wisconsin was inhibited by thick mulch. Light mulch that remained on a mowed plot also reduced germination [36]. Germination was highest on a burned plot [36]. Vegetative: Dandelion sprouts from the caudex after disturbance [114,126]. SITE CHARACTERISTICS : Dandelion tolerates a wide range of site and soil conditions, but it most commonly occurs in disturbed areas such as cut-over or burned forests, avalanche areas, overgrazed ranges, and marshy floodplains [54,133]. It also occurs sites on highway and railroad rights-of-way, waste places, old fields, pastures, and lawns [114,126]. Dandelion occurs on soils that vary from thin layers above permafrost in the subarctic to deep loams in the western United States [37,114]. Soil texture ranges from clays and clayey loams to sandy loams. Dandelion does poorly on dense clay soils, saline soils, and acidic soils [37]. Dandelion occurs on flat to rolling topography or moderate to steep slopes [27,37]. It is found from sea level to high alpine elevations [126]. Regional elevational distributions are as follows [27,37,99]: feet meters Utah 4,100-11,300 1,250-3,445 Colorado 4,500-13,500 1,372-4,115 Wyoming 4,100- 9,600 1,250-2,926 Montana 2,900- 9,200 884-2,804 Washington 2,574- 2,722 780-825 Oregon 7,095- 7,920 2,150-2,400 Alberta 4,323- 6,336 1,310-1,920 Common shrubs, grasses, and forbs associated with dandelion include common snowberry (Symphoricarpos albus), Wood's rose (Rosa woodsii), russet buffalo berry (Sheperdia canadensis), blueberry (Vaccinium spp.), chokecherry (Prunus virginiana), black sagebrush (Artemesia arbuscula nova), Wyoming big sagebrush (A. tridentata ssp. wyomingensis), Oregon-grape (Manonia repens), rough fescue (Festuca scabrella), Idaho fescue (F. idahoensis), slender wheatgrass (Elymus trachycaulus), prairie junegrass (Koeleria cristata), timber danthonia (Danthonia intermedia), Richardson's needlegrass (Stipa richardsonii), timothy (Phleum pratense), tufted hairgrass (Deschampsia caespitosa), Kentucky bluegrass (Poa pratensis), aster (Aster spp.), willowweed (Epilobium spp.), prairiesmoke avens (Geum triflorum), small-leaf angelica (Angelica pinnata), Colorado columbine (Aquilegia caerula), rhexia-leaved paintbrush (Castilleja leonardii), Oregon fleabane (Erigeron speciousus), wallflower (Erysimum elatum), one-flower helianthella (Helianthella uniflora), Utah peavine (Lathyrus utahensis), and Richardson geranium (Geranium richardsonii) [32,83,117,124,129]. SUCCESSIONAL STATUS : Obligate Initial Community Species Dandelion is an important colonizer following vegetation disturbances in temperate climates throughout North America [85,99]. Although the role of dandelion as an early seral species does not change, the length of time dandelion populations are present varies among ecosystems. Dandelion enters a disturbed community and rapidly becomes abundant. It may achieve a peak in dominance within 2 to 3 years [7,14]. Holland found dandelion to be a transitory colonist of marsh habitats in Massachusetts; it was found for 10 years after the disturbance and then disappeared [53]. Dandelion was one of the earliest colonizers after tree havesting in a maple-beech-birch ecosystem in Michigan [32]. On an abandoned farmland in Arizona, dandelion was one of the predominant species following winter precipitation [30]. Dandelion was a pioneer species on a brine-killed forest site after elimination of brine discharge on the site in the spring of 1982 [7]. On a Douglas-fir clearcut in Colorado, dandelion was a dominant species in the understory the second year after cutting but was not present in the initial community [7]. Dandelion is not a member of the climax plant community on rangelands since it cannot withstand competition for moisture, nutrients, and light with the climax vegetation. It invades these areas after the preferred species have been removed by overgrazing [85]. SEASONAL DEVELOPMENT : Dandelion is one of the earliest spring bloomers on western rangelands [134]. It flowers from March to late fall in most states and will flower throughout the year in warmer areas [126]. General first flowering dates are from April 28 to May 19, and sometimes earlier in some locations [116]. By mid-June, dandelion has reached its maximum bloom stage, and the seeds from earlier flowering dates are mostly disseminated. By mid-July, all seeds are disseminated [40]. Reported dates for anthesis in some states are as follows [16,37,100]: Utah April-July Colorado April-August Wyoming May-August Montana April-September North Dakota April-June Virginia February-June Georgia February-June Mississippi February-June Tennessee February-June Kentucky February-June Iowa April-June Alberta June-July

FIRE ECOLOGY

SPECIES: Taraxacum officinale | Dandelion
FIRE ECOLOGY OR ADAPTATIONS : Dandelion is a component of diverse ecosystems in boreal and temperate regions with variable fire regimes. Dandelion is primarily adapted to fire through its prolific production of wind-dispersed seed [123]. Site colonization after fires occurs in many forested areas because of dandelion's persistent, viable seedbank [1]. POSTFIRE REGENERATION STRATEGY : Ground residual colonizer (on-site, initial community) Initial-offsite colonizer (off-site, initial community) Caudex, growing points in soil

FIRE EFFECTS

SPECIES: Taraxacum officinale | Dandelion
IMMEDIATE FIRE EFFECT ON PLANT : NO-ENTRY DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Dandelion generally establishes during the first or second postfire year. It usually increases in frequency after fire [22,36,41]. One year after a spring burn (May 24, 1983) in Galena Gulch, Montana, dandelion showed a 50 percent increase in frequency, but by the second year showed only a 47.5 percent increase over the preburn level [22]. Dandelion increased in frequency following a fire in 1974 in a Scotch pine forest in Scotland, but by postfire year 4, frequency started to decrease. Maximum frequency occurred at 3 years after fire [119]. Dandelion frequency was greater in burned than in unburned oak communities in Utah [74]. Following a prescribed fire in a Douglas-fir stand in south-central Idaho, dandelion frequency increased significantly by postfire year 2. Prefire frequency was 8 percent; at postfire year 1 frequency was 4 percent; and at postfire year 2 frequency was 24 percent [78]. In the Hedges Mountain area of the Helena National Forest, Montana, a sagebrush/rough fescue habitat type was burned in spring (May) and fall (September). Preburn and postburn community types, as named by the dominant species, were compared. Following the spring burn, bluegrass and dandelion were the dominant species during both postfire years 1 and 2. Following the fall burn, the dominant species during postfire year 1 were bluegrass, mountain brome (Bromus marginatus), and dandelion. By postfire year 2, dandelion was no longer a dominant; the site was dominated by bluegrass, Wood's rose, and common snowberry [109]. A fire on June 28, 1977 in Montana in a rough fescue community minimally disrupted reproduction and carbohydrate production of dandelion. Its frequency increased slightly on burned sites by the summer of 1978 [6]. In the timbered breaks along the Missouri River in central Montana, dandelion was favored by big game animals every postfire year except year 28. At postfire year 17 dandelion was found at high frequencies. First peak in frequency occurred at postfire year 4 [41]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Late spring burning in the tallgrass prairies of Kansas reduced dandelion cover compared with burning at earlier dates. In shortgrass prairies of western Kansas, dandelion was less affected by dormant season (fall and winter) burns than by spring burns [20]. Burning to decrease cover of dandelion on rangelands should be done in the spring after growth initiation. Annual burning in March or November in Nebraska resulted in the highest total cover of dandelion. Burning in April decreased cover [46]. Following logging, bulldozing, and slash burning, dandelion will establish in the open spots [14]. Dandelion competes with tree seedlings on burned sites. Grasses aerially seeded on burns may compete with and displace dandelion. After 4 to 5 years of grass seeding on sites in Montana dandelion populations eventually decreased [14].

REFERENCES

SPECIES: Taraxacum officinale | Dandelion
REFERENCES : 1. Ahlgren, Clifford E. 1979. Buried seed in the forest floor of the Boundary Waters Canoe Area. Minnesota Forestry Research Note No. 271. St. Paul, MN: University of Minnesota, College of Forestry. 4 p. [3459] 2. Alaback, Paul B.; Herman, F. R. 1988. Long-term response of understory vegetation to stand density in Picea-Tsuga forests. Canadian Journal of Forest Research. 18: 1522-1530. [6227] 3. Allen, Eugene O. 1968. Range use, foods, condition, and productivity of white-tailed deer in Montana. Journal of Wildlife Management. 32(1): 130-141. [16331] 4. Almack, Jon. 1986. Grizzly bear habitat use, food habits, and movements in the Selkirk Mountains, northern Idaho. In: Contreras, Glen P.; Evans, Keith E., compilers. Proceedings--grizzly bear habitat symposium; 1985 April 30 - May 2; Missoula, MT. Gen. Tech. Rep. INT-207. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 150-157. [10815] 5. Anderson, Murray L.; Bailey, Arthur W. 1979. Effect of fire on a Symphoricarpos occidentalis shrub community in central Alberta. Canadian Journal of Botany. 57: 2820-2823. [2867] 6. Antos, Joseph A.; McCune, Bruce; Bara, Cliff. 1983. The effect of fire on an ungrazed western Montana grassland. American Midland Naturalist. 110(2): 354-364. [337] 7. Auchmoody, L. R.; Walters, R. S. 1988. Revegetation of a brine-killed forest site. Soil Science Society of America Journal. 52: 277-280. [11374] 8. Austin, D. D.; Urness, Philip J. 1982. Vegetal responses and big game values after thinning regenerating lodgepole pine. Great Basin Naturalist. 42(4): 512-516. [8354] 9. Austin, Dennis D.; Urness, Philip J. 1986. Effects of cattle grazing on mule deer diet and area selection. Journal of Range Management. 39(1): 18-21; 1986. [364] 10. Autenrieth, Robert; Molini, William; Braun, Clait, eds. 1982. Sage grouse management practices. Tech. Bull No. 1. Twin Falls, ID: Western States Sage Grouse Committee. 42 p. [7531] 11. Bard, Gily E. 1952. Secondary succession on the Piedmont of New Jersey. Ecological Monographs. 22(3): 195-215. [4777] 12. Barmore, William J., Jr.; Taylor, Dale; Hayden, Peter. 1976. Ecological effects and biotic succession following the 1974 Waterfalls Canyon Fire in Grand Teton National Park. Research Progress Report 1974-1975. Unpublished report on file at: U.S. Department of Agriculture, Forest Service, Intermountain Fire Sciences Laboratory, Missoula, MT. 99 p. [16109] 13. Basile, Joseph V.; Jensen, Chester E. 1971. Grazing potential on lodgepole pine clearcuts in Montana. Res. Pap. INT-98. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 11 p. [8280] 14. Bedunah, Don; Pfingsten, William; Kennett, Gregory; Willard, E. Earl. 1988. Relationship of stand canopy density to forage production. In: Schmidt, Wyman C., compiler. Proceedings--future forests of the Mountain West: a stand culture symposium; 1986 September 29 - October 3; Missoula, MT. Gen. Tech. Rep. INT-243. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 99-107. [5070] 15. Berch, Shannon M.; Gamiet, Sharmin; Deom, Elisabeth. 1988. Mycorrhizal status of some plants of southwestern British Columbia. Canadian Journal of Botany. 66: 1924-1928. [8841] 16. Bergen, Peter; Moyer, James R.; Kozub, Gerald C. 1990. Dandelion (Taraxacum officinale) use by cattle grazing on irrigated pasture. Weed Technology. 4(2): 258-263. [14775] 17. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 18. Bowes, G. G. 1991. Long-term control of aspen poplar and western snowberry with dicamba and 2,4-D. Canadian Journal of Plant Science. 71(4): 1121-1131. [19502] 19. Bowns, James E.; Bagley, Calvin F. 1986. Vegetation responses to long-term sheep grazing on mountain ranges. Journal of Range Management. 39(5): 431-434. [15584] 20. Bragg, Thomas B. 1991. Implications for long-term prairie management from seasonal burning of loess hill and tallgrass prairie. In: Nodvin, Stephen C.; Waldrop, Thomas A., eds. Fire and the environment: ecological and cultural perspectives: Proceedings of an international symposium; 1990 March 20-24; Knoxville, TN. Gen. Tech. Rep. SE-69. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station: 34-44. [16631] 21. Brown, David E. 1982. Great Basin montane scrubland. In: Brown, David E., ed. Biotic communities of the American Southwest--United States and Mexico. Desert Plants. 4(1-4): 83-84. [8890] 22. Bushey, Charles L. 1985. Summary of results from the Galena Gulch 1982 spring burns (Units 1b). Missoula, MT: Systems for Environmental Management. 9 p. [567] 23. Call, Mayo W. 1979. Habitat requirements and management recommendations for sage grouse. Denver, CO: U.S. Department of the Interior, Bureau of Land Management, Denver Service Center. 37 p. [591] 24. Canon, S. K.; Urness, P. J.; DeByle, N. V. 1987. Habitat selection, foraging behavior, and dietary nutrition of elk in burned aspen forest. Journal of Range Management. 40(5): 443-438. [3453] 25. Carleton, T. J.; Maycock, P. F. 1981. Understorey - canopy affinities in boreal forest vegetation. Canadian Journal of Botany. 59: 1709-1716. [14576] 26. Cole, C. Andrew. 1991. The seedbank of a young surface mine wetland. Wetlands Ecology and Management. 1(3): 173-184. [19468] 27. Cole, David N.; Hall, Troy E. 1992. Trends in campsite condition: Eagle Cap Wilderness, Bob Marshall Wilderness, and Grand Canyon National Park. Res. Pap. INT-453. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 40 p. [17764] 28. Collins, William B.; Urness, Philip J. 1983. Feeding behavior and habitat selection of mule deer and elk on northern Utah summer range. Journal of Wildlife Management. 47(3): 646-663. [6915] 29. Cooper, David J. 1990. Ecology of wetlands in Big Meadows, Rocky Mountain National Park, Colorado. Biological Report 90(15). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 45 p. [16106] 30. Cox, J. R.; Madrigal, R. M. 1988. Establishing perennial grasses on abandoned farmland in southeastern Arizona. Applied Agricultural Research. 3(1): 36-43. [11177] 31. Crouch, Glenn L. 1985. Effects of clearcutting a subalpine forest in central Colorado on wildlife habitat. Res. Pap. RM-258. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 12 p. [8225] 32. Crow, T. R.; Mroz, G. D.; Gale, M. R. 1991. Regrowth and nutrient accumulations following whole-tree harvesting of a maple-oak forest. Canadian Journal of Forest Research. 21: 1305-1315. [16600] 33. Currie, P. O.; Reichert, D. W.; Malechek, J. C.; Wallmo, O. C. 1977. Forage selection comparisons for mule deer and cattle under managed ponderosa pine. Journal of Range Management. 30(5): 352-356. [4697] 34. Davis, James N.; Harper, Kimball T. 1990. Weedy annuals and establishment of seeded species on a chained juniper-pinyon woodland in central Utah. In: McArthur, E. Durant; Romney, Evan M.; Smith, Stanley D.; Tueller, Paul T., compilers. Proceedings--symposium on cheatgrass invasion, shrub die-off, and other aspects of shrub biology and management; 1989 April 5-7; Las Vegas, NV. Gen. Tech. Rep. INT-276. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 72-79. [12872] 35. Deschamp, Joseph A.; Urness, Philip J.; Austin, Dennis D. 1979. Summer diets of mule deer from lodgepole pine habitats. 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