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Wildlife, Animals, and Plants |
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INTRODUCTORY
ABBREVIATION:ACHHYM SYNONYMS:Oryzopsis hymenoides (Roemer & J.A. Schultes) Ricker ex Piper [33,38,50,76,117] NRCS PLANT CODE:ACHY COMMON NAMES:
Indian ricegrass TAXONOMY:The fully documented scientific name of Indian ricegrass is Achnatherum hymenoides (Roemer & J.A. Schultes.) Barkworth [55]. Indian ricegrass hybridizes with various needlegrasses (Stipa spp. sensu lato) including green needlegrass (Nassella viridula) [65]. Achnatherum contractum is a fertile derivative of an Indian ricegrass × smilo grass (Piptatherum micranthum) cross [10]. LIFE FORM:graminoid FEDERAL LEGAL STATUS:No special status OTHER STATUS:No entry AUTHORSHIP AND CITATION:Tirmenstein, D. (1999, March). Achnatherum hymenoides. In: Remainder of Citation Species Index FEIS Home DISTRIBUTION AND OCCURRENCE
GENERAL DISTRIBUTION:Indian ricegrass is widely distributed throughout the western United States. [18,113]. It occurs east of the Cascade Range from British Columbia south to southern California and northeastern Mexico and east to Alberta, the Dakotas, and Texas [56]. ECOSYSTEMS:
FRES21 Ponderosa pine STATES:
AR AZ CA CO ID KS MN MO MT NE NV BLM PHYSIOGRAPHIC REGIONS:
3 Southern Pacific Borders KUCHLER PLANT ASSOCIATIONS:
K011 Western ponderosa forest SAF COVER TYPES:
220 Rocky Mountain juniper SRM (RANGELAND) COVER TYPES:
101 Bluebunch wheatgrass HABITAT TYPES AND PLANT COMMUNITIES:
Indian ricegrass occurs throughout the sagebrush series and is present in several salt-desert shrub communities. It also grows in pinyon-juniper (Pinus-Juniperus spp.) and ponderosa pine (P. ponderosa) forest communities. It is also locally abundant in mixed-grass prairie of Canada and in sagebrush (Artemisia spp.) and desert shrub [21,58,127,129]. Common associates are needle-and-thread grass (Hesperostipa comata), sideoats grama (Bouteloua curtipendula), blue grama (B. grama), and wheatgrass (Triticeae) [58]. Indian ricegrass often grows interspersed in open stands of big sagebrush (A. tridentata) with species such as Thurber needlegrass (S. thurberiana), Sandberg bluegrass (Poa secunda), and bottlebrush squirreltail (Elymus elymoides) [127]. It occurs in black sagebrush (A. nova), winterfat (Krascheninnikovia lanata), and shadscale (Atriplex confertifolia) communities [58,60]. VALUE AND USE
IMPORTANCE TO LIVESTOCK AND WILDLIFE:
Indian ricegrass has good forage value for domestic sheep, cattle, and horses. It can be important cattle forage in winter, particularly in salt desert communities [24,130]. Indian ricegrass is often used most heavily in late winter, when succulent and nutritious new green leaves are produced [59,60]. It supplies a source of green feed before most other native grasses have produced much new growth [95]. Consequently, Indian ricegrass is often heavily grazed before animals leave winter ranges [51]. PALATABILITY:Indian ricegrass is described as highly palatable to all classes of livestock in both green and cured condition [18,58]. Indian ricegrass has been reported as "dependably palatable" in winter [56]. However, flowerstalks are unpalatable late in the spring and are not grazed at that time [89]. Seeds and seedstalks are relished by domestic sheep [60]. NUTRITIONAL VALUE:
Indian ricegrass does not provide enough digestible protein to meet minimum requirements for some ungulates [116]. It is deficient in phosphorous, carotene, and digestible protein but is a relatively good energy source [29]. Indian ricegrass seed has approximately 4058.0 calories per gram (16.1 ash-free calories per seed) [71].
Total weight (%)
dry matter = 92.6 structural carbohydrates = 32.2 ash = 4.5
free water = 7.4 lignin = 8.4 crude protein = 13.1
cell walls = 43.1 acid insoluble ash = 2.5 solvent extract = 2.8
soluble ash = 2.0 cell contents = 56.9 soluble carbohydrates = 39.0
non-nutritive matter = 10.9
A study involving seasonal variation of carbohydrate reserves in roots and crowns of desert range species in northern Utah indicated that the crowns of Indian ricegrass accumulate higher concentrations of total available carbohydrates than the roots. Fall depletion of both roots and crowns is common and is proportional to growth activity in the fall. Spring depletion of total available carbohydrates is less when precipitation is higher [31]. Additional nutritional studies are available [32,101,121].
COVER VALUE:
The degree to which Indian ricegrass provides cover for wildlife species has been rated as follows [28]:
UT CO WY MT ND
Elk poor ---- poor ---- ----
Mule deer poor ---- poor ---- fair
White-tailed deer ---- ---- poor ---- fair
Pronghorn poor ---- poor ---- poor
Upland game bird good ---- poor poor ----
Waterfowl poor ---- poor ---- ----
Small nongame bird good fair fair poor ----
Small mammals good fair poor ---- ----
VALUE FOR REHABILITATION OF DISTURBED SITES:
Indian ricegrass is well-suited for surface erosion control and desert revegetation although it is not highly effective in controlling sand movement [25,61,81]. Certain native ecotypes exhibit desirable characteristics such as drought and salinity tolerance, low seed dormancy, and good nutritional qualities [84]. However, Indian ricegrass can be difficult to establish [70]. Indian ricegrass can be useful in the reclamation of many arid and semiarid areas in the western United States [84]. Typical sites include those in which vegetation has been removed due to surface mining, construction activity, brush control, heavy grazing, or fire [110]. Indian ricegrass can be used for revegetating degraded rangelands in areas of low precipitation and has naturally revegetated overgrazed ranges in parts of Utah [110,125]. OTHER USES AND VALUES:
As its name suggests, Indian ricegrass was traditionally eaten by some Native American peoples [72]. The Paiutes used seed as a reserve food source [24]. MANAGEMENT CONSIDERATIONS:
Heavy early spring grazing may sharply reduce the vigor of Indian ricegrass and decrease the stand [107]. Plants that do survive exhibit poor vigor with short sparse foliage and dead centers, and may produce no litter or flower stalks [9]. In eastern Idaho, productivity of Indian ricegrass was at least 10 times greater in undisturbed plots than in heavily grazed ones [88]. In southeastern Idaho where grazing was eliminated for 25 years, Indian ricegrass increased in cover 5 fold. Both the density and basal area decreased with increasing grazing intensity in Glen Canyon National Recreation Area, Utah. The seed crop may be reduced where grazing is heavy [15]. BOTANICAL AND ECOLOGICAL CHARACTERISTICS
GENERAL BOTANICAL CHARACTERISTICS:Indian ricegrass is a hardy, cool-season, densely tufted, native perennial bunchgrass that grows from 4 to 24 inches (10-61 cm) in height [18,23,83]. Plant spread is estimated at 8 to 12 inches (20-30 cm) [61]. When mature, seedstalks are open, loosely branched, or with lacy tops [60]. Spikes are erect and range from 0.8 to 2.8 inches (2-7 cm) in length [18]. Indian ricegrass has deep, fibrous, extensive roots and is one of the most drought tolerant of the native range grasses [61,83,123]. The common name is derived from seeds that resemble grains of rice [83,111]. RAUNKIAER LIFE FORM:Chamaephyte REGENERATION PROCESSES:
Plummer and others [93] report that Indian ricegrass naturally spreads aggressively from seed, but Valentine [115] describes seedling establishment as "poor" in comparison to that of other species. Plummer and Frischknecht [94] reported that numerous seedlings often develop in years with wet springs, while very few develop during dry springs. Therefore, years of good seedling establishment can be quite infrequent. A high degree of correlation between April precipitation and Indian ricegrass abundance has been reported [46]. SITE CHARACTERISTICS:
Indian ricegrass grows on dry foothills, rocky valley or upper slopes, plains, and ridges [58,61,99]. It is moderately tolerant of both alkali and salt, and is adapted to soils of low fertility [29, 88]. This species is particularly well adapted to sand and attains greatest abundance on loose sandy soils where it forms almost pure stands [29,60,58]. Indian ricegrass dominates sandy soils throughout the sagebrush-grass zone [124]. Soils supporting Indian ricegrass range in texture from coarse sand to heavy clay [94]. It is also found on shallow shale soils [61]. SUCCESSIONAL STATUS:Koniak [73] reports that Indian ricegrass reaches greatest cover in early mid- to mid-seral stands following fire in pinyon-juniper communities of the Great Basin. Cover decreases by late succession [73]. In southeastern Idaho, Indian ricegrass increased during a 25-year disturbance-free period [22]. After approximately 20 more years, cover began to decline. In mixed-grass prairies of Colorado, Indian ricegrass plants commonly appear within the first 5 years after disturbance such as plowing [30]. Indian ricegrass is a pioneer on blowouts and loose sands of sand sage (Artemisia filifolia) communities of northeastern Colorado [97]. It is typically found on disturbed (but not undisturbed) sites in blackbrush (Coleogyne ramosissima) communities in parts of Nevada [47]. Indian ricegrass is present in several climax communities including climax bunchgrass communities of Oregon [1]. It forms part of a distinct sub-climax grassland community in parts of northern Utah [106]. SEASONAL DEVELOPMENT:Indian ricegrass generally flowers early prior to summer drought [42]. In Nevada flower buds may form as early as February or March [2]. However, flowering is indeterminate and can continue through the spring and summer if growing conditions remain favorable [120]. Ackerman and others [2] reported that in the Nevada desert, Indian ricegrass flowered after spring rains, but not following summer rains. This suggests that lower temperatures may be required for flowering. Time of flowering varies as follows [28]:
earliest flowering latest flowering
CO May July
MT May June
ND June June
WY May July
Average dates at which Indian ricegrass reached various developmental stages in Upper Snake Plains of southern Idaho are as follows [17]:
Date Range (days) Standard deviation
growth starts Apr 08
flowerstalks appear May 22 27 7.9
heads fully out Jun 17 26 8.6
flowers in bloom Jun 20 20 6.5
seed ripe Jul 03 21 7.3
dissemination starts Jul 06
dissemination over Jul 29
plant drying Jul 03 48 7.8
plant dried Aug 06
The following phenological development has been reported in Nevada [42]:
Mean dates breaking dormancy Mean days of active development March 31 72 April 2 74 April 7 74Indian ricegrass can produce green shoots in fall and generally stays dry until late winter or early spring when new growth begins. Lower portions of the stem may remain somewhat green during most of the winter [29]. Panicles of different maturities may be present on a particular plant at any given time during the growing season [120]. FIRE ECOLOGYFIRE ECOLOGY OR ADAPTATIONS:
Indian ricegrass reestablishes on burned sites through seed dispersed from adjacent unburned areas. Fire intervals in sagebrush-grass communities have been estimated at 7 to 70 years [131]. The range of fire intervals reported for some species that dominate communities in which Indian ricegrass occurs are listed below. To learn more about the fire regimes in those communities refer to the FEIS summary for that species, under "Fire Ecology or Adaptations." POSTFIRE REGENERATION STRATEGY:
Tussock graminoid FIRE EFFECTS
IMMEDIATE FIRE EFFECT ON PLANT:Indian ricegrass has not been the subject of intensive research. Pechanec and others [90] report that it is slightly damaged by fire. However, in Utah, Ralphs and others [96] note that is "moderately damaged" by fire. According to West [118], plants are generally killed by fire. DISCUSSION AND QUALIFICATION OF FIRE EFFECT:No entry PLANT RESPONSE TO FIRE:
Indian ricegrass regenerates from seed following fire [118]. In western and central Utah, Indian ricegrass may represent important cover on burned areas and often exhibits good growth characteristics. Low culm density reduces charring of crowns below soil, thereby protecting growing points. Spring burning in Utah did little damage to Indian ricegrass, and growth began within 3 weeks after burning [124]. In Nevada, summer wildfires reduced the basal area of Indian ricegrass, but little mortality was observed [122]. Rapid postfire responses such as these suggest that Indian ricegrass may also sprout from tillers. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE:
In singleleaf pinyon (Pinus monophylla)-Utah juniper (Juniperus osteosperma) communities of the Great Basin, Everett [41] reported that Indian ricegrass could increase in density soon after fire. Postfire recovery time of Indian ricegrass is listed as 2 to 4 years [124]. However, West and Hassan [119] report that only a small portion of preburn cover had been gained one year after a July burn in Utah. Specific values are as follows [119]:
After an annual early spring burn of grassland-shrublands in Canada, canopy cover and frequency of Indian ricegrass increased significantly (p<0.005) [7]:
FIRE MANAGEMENT CONSIDERATIONS:
Annual burning may produce a more arid environment than usually found in fescue (Festuca spp.) prairie, favoring mixed-prairie grasses such as Indian ricegrass.
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Related categories for SPECIES: Achnatherum hymenoides | Indian Ricegrass |
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