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Introductory

SPECIES: Bothriochloa barbinodis | Cane Bluestem
ABBREVIATION : BOTBAR SYNONYMS : Andropogon barbinodis Lag. Gen. & Sp. Andropogon saccharoides var. barbinodis Hackel in A. & C. DC. SCS PLANT CODE : BOBA3 COMMON NAMES : cane bluestem cane beardgrass feather bluestem plains beardgrass pinhole bluestem TAXONOMY : The currently accepted scientific name for cane bluestem is Bothriochloa barbinodis (Lag.) Herter. Two varieties are currently recognized [13]: Bothriochloa barbinodis var. barbinodis - cane bluestem Bothriochloa barbinodis var. perforata (Trin. ex Fourn.) Gould - pinhole bluestem LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Ronald Uchytil/November 1988 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Uchytil, Ronald J. 1988. Bothriochloa barbinodis. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Bothriochloa barbinodis | Cane Bluestem
GENERAL DISTRIBUTION : Cane bluestem is restricted to the southwestern United States and Mexico. Its range extends from southern California, across southern Nevada to southern Colorado, south through Oklahoma, Texas, New Mexico, and Arizona into Mexico [6,25]. ECOSYSTEMS : FRES21 Ponderosa pine FRES28 Western hardwoods FRES30 Desert shrub FRES35 Pinyon - juniper FRES38 Plains grasslands FRES40 Desert grasslands STATES : AZ CA NV NM OK TX UT MEXICO ADMINISTRATIVE UNITS : AMIS BIBE CABR CACH CACA CHIR CORO FOBO GLCA GRCA GUMO JOTR LAME LAMR ORPI SAGU SAMO WUPA BLM PHYSIOGRAPHIC REGIONS : 7 Lower Basin and Range 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains KUCHLER PLANT ASSOCIATIONS : K023 Juniper - pinyon woodland K031 Oak - juniper woodlands K053 Grama - galleta steppe K057 Galleta - threeawn shrubsteppe K058 Grama - tobosa shrubsteppe K060 Mesquite savanna SAF COVER TYPES : 239 Pinyon - juniper 241 Western live oak SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Cane bluestem is most commonly found in semidesert grasslands, oak woodlands, chaparral, pinyon-juniper woodlands, and shortgrass plains, where it usually occurs as scattered plants or in small groups [19,27]. It is a common grass in many Arizona chaparral communities [6]. In oak (Quercus emoryi) woodlands of Arizona, it is often an understory codominant with side oats-grama (Bouteloua curtipendula), hairy grama (Bouteloua hirsuta), threeawn (Aristida spp.), and plains lovegrass (Eragrostis intermedia). Cane bluestem has not been listed as a climax indicator in published classification schemes.

VALUE AND USE

SPECIES: Bothriochloa barbinodis | Cane Bluestem
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Cane bluestem is both palatable and productive [19,20,28], and consumed by all classes of livestock. Cattle and horses utilize this grass more than sheep do, and it may be an important local food source for pronghorn [3,17,19,25]. The cane bluestem selection, PMT-333, which originates from Texas, has yielded up to 6,513 pounds per acre (7,300 kgs/ha) when irrigated [20]. PALATABILITY : Cane bluestem is most palatable to livestock during the spring and summer, before the stems become fully mature and fibrous [3,14,17]. Later in the season as plants mature, the leaves are consumed by cattle and horses but are considered too coarse for sheep [16,19]. Throughout its range, cane bluestem's palatability is generally considered to be good for cattle, horses, and pronghorn; and fair for sheep [3,16,28]. NUTRITIONAL VALUE : Cane bluestem has fair forage value. Nutritional analysis shows that protein, ash, and phosphorus levels all drop when the plant is mature. Seasonal trends in the nutritional composition of cane bluestem in the Edwards Plateau region of Texas are presented below [18]: % Composition date protein ash cell wall Phos DOM old and new growth 4/13 8 11 68 .12 33 leaves 5/24 9 10 65 .14 55 leaves and stems 6/28 9 9 66 .15 57 leaves and stems 8/30 6 8 68 .08 47 leaves and stems 10/3 6 5 74 .07 40 leaves and stems 10/25 5 7 70 .08 52 leaves and stems 12/27 3 9 71 .03 48 These results are consistent with Koshi and others [20], who reported that phosphorus and protein levels were inversely related to the age of plants at time of harvest. They observed that stands clipped three times per season yielded 5,443 pounds per acre (6,100 kg/ha) of forage containing 8.3 percent protein and 0.16 percent phosphorus; stands clipped twice produced 6,602 pounds per acre (7,400 kg/acre) of forage containing 7.0 percent protein and 0.17 percent phosphorus; and stands clipped once produced 6,959 pounds per acre (7,800 kg/ha) of forage containing 4.4 percent protein and 0.12 percent phosphorus. In this study protein levels were also found to be inversely related to the amount of water applied. COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Cane bluestem is recommended for reseeding arid southwestern rangelands. Seed, however, is available in very limited quantities. No cultivars have been released, and for reseeding purposes, seed may need to be collected from native stands. Cane bluestem is best suited for planting on silty or clayey soils in areas with 12 to 20 inches mean annual precipitation [24]. Cane bluestem averages 750,000 pure seeds per pound (1,653,000/kg). Recommended seeding rates are 17.2 pure seeds/square foot at 1 pound per acre (185 pure seeds/sq m at 1.12 kg/ha) [24]. OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : Cane bluestem is an indicator of range in good condition. This decreaser is one of the first grasses to disappear from ranges that are overgrazed [4,5,19]. Although cane bluestem is listed as a decreaser by numerous authors, Bernardon and others [2] observed under greenhouse conditions that clipping up to 60 percent of herbage at any stage during the first year of seedling development was not detrimental to subsequent root and herbage production. Cane bluestem is recommended for reseeding arid southwestern rangelands, but seed is available in very limited quantities. No cultivars have been released, and for reseeding, seed may need to be collected from native stands. Cane bluestem is best suited for planting in areas receiving 12 to 20 inches mean annual precipitation, on silty or clayey soils [24]. There are approximately 750,000 pure seeds per pound (1,653,000/kg). Recommended seeding rates are 17.2 pure seeds/square foot at one pound per acre (185 pure seeds/square meter at 1.12 kg/ha) [24].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Bothriochloa barbinodis | Cane Bluestem
GENERAL BOTANICAL CHARACTERISTICS : Cane bluestem is an extremely drought-resistant, native, warm-season, robust perennial bunchgrass, of medium height, with straw-colored pithy stems [16,19,25]. Culms are erect to spreading, 2 to 5 feet (0.6-1.5 m) tall and densly tufted [27]. Leaves are blue-green and cure to a dull red or yellow; they occur basally and on the flower stalks [7,25]. The inflorescence consists of a panicle, 2 to 5 inches (4.5-13 cm) long, with the appearance of a fan-shaped silvery-hairy cluster [25,27]. Pairs of spikelets occur at each joint. One is sessile and seed producing, with a bent awn about 0.75 inch (1.9 cm) long; the other is infertile with a 0.1- to 0.15-inch-long (3-4 mm) pedicel [7]. The presence (var. perforata) or absence (var. barbinodis) of a glandular pit on the first outer glume of the sessile spikelet is a consistent botanical characteristic used to distinguish the two varieties [14]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Cane bluestem produces an abundance of seed that is dispersed by wind [28]. The seeds apparently germinate readily, and the seedling are hardy [28]. Locally, during drought conditions, cane bluestem may be one of the only grasses producing seed. SITE CHARACTERISTICS : Cane bluestem occurs primarily in dry, sandy, gravelly or rocky sites [17,27]. It is best adapted to sandy-loam to calcerious-loam soils, with a pH range of 7.2 to 8.0 [8]. It is most common in semidesert grasslands, oak woodlands, and chaparral [19]. In open rangelands, it occurs primarily in areas of high water concentration or in areas that are occasionally flooded, which allows it to grow where annual precipitation is only 5 to 7 inches [16,19,25]. Cane bluestem usually occurs scattered; it seldom forms dense or pure stands [19,25]. Elevational ranges for cane bluestem in several western states are presented below [8,11,16,27]: from 1,000 to 5,800 feet (305-1,768 m) in AZ 5,000 to 5,600 feet (1,524-1,707 m) in CO 3,000 to 7,000 feet (914-2,134 m) in NM 3,020 to 6,000 feet (920-1,830 m) in UT SUCCESSIONAL STATUS : NO-ENTRY SEASONAL DEVELOPMENT : Growth of cane bluestem seems to be dependent upon available moisture. Most growth occurs between mid-June and late August when moisture is adequate. In Texas, irrigated plants produced 25 percent of seasonal growth by mid-June, 62 percent between mid-June and mid-July, and 13 percent after mid-July. Plants grown under natural rainfall, with water available later in the season, grew more later in the season. These plants produced 20 percent, 30 percent, and 50 percent of their seasonal growth during the same time periods [20]. Plants usually flower between May and October in Texas, but flowering will occur throughout the year under favorable growing conditions [14].

FIRE ECOLOGY

SPECIES: Bothriochloa barbinodis | Cane Bluestem
FIRE ECOLOGY OR ADAPTATIONS : Information concerning the effects of fire on cane bluestem is lacking. This plant is a large, robust, coarse, perinnial bunchgrass with leaves occurring basally and along the stalk [19]. Wright and Bailey [29] and Young [30] report that large bunchgrasses with densely clustered stems are more susceptible to fire than small bunchgrasses with coarse stems because fire can burn down into the clump for sometimes 2 to 3 hours after the fire has passed. Small bunchgrasses with smaller amounts of leafy materials are consumed by a fire quickly, with little heat transferred downward into the clump. General botanical descriptions seem to place cane bluestem somewhere intermediate between these two types. POSTFIRE REGENERATION STRATEGY : Tussock graminoid Initial-offsite colonizer (off-site, initial community)

FIRE EFFECTS

SPECIES: Bothriochloa barbinodis | Cane Bluestem
IMMEDIATE FIRE EFFECT ON PLANT : NO-ENTRY DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : NO-ENTRY DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Bothriochloa barbinodis | Cane Bluestem
REFERENCES : 1. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 2. Bernardon, Abel E.; Huss, Donald L.; McCully, Wayne G. 1967. Effects of herbage removal on seedling development in cane bluestem. Journal of Range Management. 20: 69-72. [4768] 3. Buechner, Helmut K. 1950. Life history, ecology, and range use of the pronghorn antelope in Trans-Pecos Texas. American Midland Naturalist. 43(2): 257-354. [4084] 4. Canfield, R. H. 1948. Perennial grass composition as an indicator of condition of Southwestern mixed grass ranges. Ecology. 29: 190-204. [5308] 5. Canfield, R. H. 1957. Reproduction and life span of some perennial grasses of southern Arizona. Journal of Range Management. 10(5): 199-203. [3938] 6. Carmichael, R. S.; Knipe, O. D.; Pase, C. P.; Brady, W. W. 1978. Arizona chaparral: plant associations and ecology. Res. Pap. RM-202. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 16 p. [3038] 7. Cronquist, Arthur; Holmgren, Arthur H.; Holmgren, Noel H.; [and others]. 1977. Intermountain flora: Vascular plants of the Intermountain West, U.S.A. Vol. 6. The Monocotyledons. New York: Columbia University Press. 584 p. [719] 8. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 9. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 10. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 11. Gay, Charles W., Jr.; Dwyer, Don D. 1965. New Mexico range plants. Circular 374. Las Cruces, NM: New Mexico State University, Cooperative Extension Service. 85 p. [4039] 12. Gould, F. W. 1967. The grass genus Andropogon in the United States. Brittonia. 19: 70-76. [5526] 13. Gould, Frank W. 1975. The grasses of Texas. College Station, TX: Texas A&M University Press. 650 p. [5668] 14. Gould, Frank W. 1978. Common Texas grasses. College Station, TX: Texas A&M University Press. 267 p. [5035] 15. Gould, Frank W.; Shaw, Robert B. 1983. Grass systematics. 2d ed. College Station, TX: Texas A&M University Press. 397 p. [5667] 16. Humphrey, Robert R. 1960. Arizona range grasses: Description--forage value--management. Tucson, AZ: University of Arizona, Agricultural Experiment Station. 104 p. [5004] 17. Humphrey, Robert R. 1970. Arizona range grasses: Their description, forage value and management. Tucson, AZ: The University of Arizona Press. 159 p. [5567] 18. Huston, J. E.; Rector, B. S.; Merrill, L. B.; Engdahl, B. S. 1981. Nutritional value of range plants in the Edwards Plateau region of Texas. Report B-1375. College Station, TX: Texas A&M University System, Texas Agricultural Experiment Station. 16 p. [4565] 19. Judd, B. Ira. 1962. Principal forage plants of southwestern ranges. Stn. Pap. No. 69. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 93 p. [1302] 20. Koshi, P. T.; Eck, H. V.; Stubbendieck, J.; McCully, W. G. 1977. Cane bluestem: forage yield, forage quality, and water-use efficiency. Journal of Range Management. 30(3): 190-193. [5544] 21. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 22. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 23. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 24. Thornburg, Ashley A. 1982. Plant materials for use on surface-mined lands. SCS-TP-157. Washington, DC: U.S. Department of Agriculture, Soil Conservation Service. 88 p. [3769] 25. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. [2387] 26. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 27. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 28. Weaver, J. E.; Albertson, F. W. 1956. Grasslands of the Great Plains. Lincoln, NE: Johnsen Publishing Company. 395 p. [2463] 29. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. [2620] 30. Young, Richard P. 1983. Fire as a vegetation management tool in rangelands of the Intermountain Region. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 18-31. [2681] 31. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090]

Index

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