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Introductory

SPECIES: Buchloe dactyloides | Buffalo Grass
ABBREVIATION : BUCDAC SYNONYMS : NO-ENTRY SCS PLANT CODE : BUDA COMMON NAMES : buffalo grass TAXONOMY : The currently accepted scientific name of buffalo grass is Buchloe dactyloides (Nutt.) Engelm. (Poaceae). There are no infrataxa. Buchloe is a monotypic genus [48,53,60,61]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Gail Winkler, September 1987 LAST REVISED BY AND DATE : Janet L. Howard, November 1995 AUTHORSHIP AND CITATION : Howard, Janet L. 1995; Winkler, Gail L. 1987. Buchloe dactyloides. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Buchloe dactyloides | Buffalo Grass
GENERAL DISTRIBUTION : Buffalo grass is distributed from central Montana east to Minnesota and south to eastern coastal Louisiana, Texas, New Mexico, eastern Arizona, and northern Mexico [48,53]. It is incidental in northern Idaho [95] and Virginia [79]. ECOSYSTEMS : FRES15 Oak-hickory FRES21 Ponderosa pine FRES32 Texas savanna FRES33 Southwestern shrubsteppe FRES35 Pinyon-juniper FRES38 Plains grasslands FRES39 Prairie FRES40 Desert grasslands STATES : AZ CO ID IA KS LA MN MO MT NE NM ND OK SD TX VA WY ADMINISTRATIVE UNITS : AGFO AMIS BADL CACA CHIC FOLA FOSC GUMO HOME LAME NEPE PIPE SCBL WICR WICA BLM PHYSIOGRAPHIC REGIONS : 7 Lower Basin and Range 10 Wyoming Basin 11 Southern Rocky Mountains 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K017 Black Hills pine forest K023 Juniper-pinyon woodland K031 Oak-juniper woodlands K044 Creosotebush-tarbush K053 Grama-galleta steppe K054 Grama-tobosa prairie K058 Grama-tobosa shrubsteppe K059 Trans-Pecos shrub savanna K060 Mesquite savanna K061 Mesquite-acacia savanna K064 Grama-needlegrass-wheatgrass K065 Grama-buffalograss K066 Wheatgrass-needlegrass K067 Wheatgrass-bluestem-needlegrass K068 Wheatgrass-grama-buffalograss K069 Bluestem-grama prairie K074 Bluestem prairie K075 Nebraska Sandhills prairie K084 Cross Timbers K085 Mesquite-buffalograss K087 Mesquite-oak savanna K096 Northeastern spruce-fir forest SAF COVER TYPES : 40 Post oak-blackjack oak 66 Ashe juniper-redberry (Pinchot) juniper 68 Mesquite 237 Interior ponderosa pine 239 Pinyon-juniper 242 Mesquite SRM (RANGELAND) COVER TYPES : 412 Juniper-pinyon woodland 504 Juniper-pinyon pine woodland 505 Grama-tobosa shrub 601 Bluestem prairie 604 Bluestem-grama prairie 605 Sandsage prairie 606 Wheatgrass-bluestem-needlegrass 607 Wheatgrass-needlegrass 608 Wheatgrass-grama-needlegrass 609 Wheatgrass-grama 611 Blue grama-buffalograss 615 Wheatgrass-saltgrass-grama 701 Alkali sacaton-tobosagrass 703 Black grama-sideoats grama 704 Blue grama-western wheatgrass 705 Blue grama-galleta 706 Blue grama-sideoats grama 709 Bluestem-grama 710 Bluestem prairie 711 Bluestem-sacahuista prairie 712 Galleta-alkali sacaton 713 Grama-muhly-threeawn 714 Grama-bluestem 715 Grama-buffalograss 717 Little bluestem-Indiangrass-Texas wintergrass 718 Mesquite-grama 720 Sand bluestem-little bluestem (dunes) 727 Mesquite-buffalograss 728 Mesquite-granjeno-acacia 729 Mesquite 732 Cross timbers-Texas (little bluestem-post oak) 733 Juniper-oak 734 Mesquite-oak 802 Missouri prairie HABITAT TYPES AND PLANT COMMUNITIES : Buffalo grass is codominant with blue grama (Bouteloua gracilis) over most of the shortgrass prairie [51]. It is a common component in mixed-grass prairie [3,30], semidesert grassland of New Mexico [17], and coastal prairie of Louisiana and Texas [33]. It is usually a minor element in undisturbed tallgrass prairie [26,46]. Buffalo grass also occurs in the understory of pinyon-juniper (Pinus-Juniperus spp.), mesquite (Prosopsis spp.), and eastern ponderosa pine (Pinus ponderosa) woodland [8], and in oak-hickory (Quercus-Carya spp.) savanna of the Cross Timbers region of Texas [39]. Plant community classifications naming buffalo grass as a community dominant are as follows: A framework for plant community classification and conservation in Texas [33] Remnant grassland vegetation and ecological affinities of the upper coastal prairie of Texas [34] A vegetation classification system for New Mexico, U.S.A. [37] Characteristics of major grassland types in western North Dakota [49] Distribution and ecology of loess hill prairies in Atchison and Holt counties in northwestern Missouri [58] A study of the vegetation of the sandhills of Nebraska [75] Plant communites of Texas [91] Analysis of grassland vegetation on selected key areas in southwestern North Dakota [109]

VALUE AND USE

SPECIES: Buchloe dactyloides | Buffalo Grass
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Buffalo grass is one of the most important forage grasses of the shortgrass and mixed-grass prairies. All classes of livestock graze it during all seasons [59]. The foliage is nutritious and palatable when green, and its nutritional quality does not decline greatly as it cures [52,68]. Buffalo grass is also important wildlife forage. Wildlife consumers of buffalo grass include white-tailed deer [22], bison [73], pronghorn, black-tailed jackrabbit [38], and prairie dogs [24]. Buffalo grass is especially important in bison [73] and black-tailed prairie dog diets [24]. Bison in northeastern Colorado consumed it year-round, with mean percent composition in their diet least in May (41%) and highest in August (84%) [73]. Black-tailed prairie dog also use it year-round, consuming all parts of the plant [13]. Annual black-tailed prairie dog consumption of buffalo grass averaged 23 percent in western South Dakota, peaking in June and July (34%) and reaching a low in December (8%) [90]. Mountain plover nest on blue grama-buffalo grass flats in Colorado [47]. PALATABILITY : Palatability of buffalo grass has been rated good for cattle, domestic sheep, and horses [36]. Reitz and Morris [81] rated it one of the most palatable grasses in Montana. Livestock utilization may vary by region and year, however. In a cattle utilization study in South Dakota mixed-grass prairie, cattle selected several other graminoids over buffalo grass even though buffalo grass was the most productive graminoid on the study site [96]. NUTRITIONAL VALUE : Nutritional value of fresh, mature buffalo grass in the United States was [70]: Percent dry matter 48.9 ash 6.1 crude fiber 13.0 digestible protein cattle 2.9 domestic sheep 2.9 domestic goats 2.7 horses 2.7 rabbits 2.9 The National Academy of Sciences [70] also provides nutritional analyses of fresh and cured buffalo grass in other phenological stages (immature, dough stage, ripe, overripe), and for fresh buffalo grass-western wheatgrass-bluestem (Grama spp.) mixes. Energy (calories/kg) and mineral analyses of buffalo grass are given. Harlan [50] published a nutritional analysis of buffalo grass seed. Dittberner and Olson [36] rated buffalo grass fair in energy and protein value. They rated its nutritional value for wildlife in several states as follows: UT CO MT ND elk good fair poor ---- mule deer good poor poor ---- white-tailed deer ---- ---- ---- poor pronghorn good ---- ---- poor upland game birds good good poor poor waterfowl fair ---- ---- ---- nongame birds good ---- poor ---- small mammals good good poor ---- COVER VALUE : Dittberner and Olson [36] rated the value of buffalo grass in providing cover for wildlife as follows: WY ND upland game birds good poor small nongame birds good poor small mammal cover good poor VALUE FOR REHABILITATION OF DISTURBED SITES : Buffalo grass sod is ranked superior in controlling erosion [100,106]. It has been ranked first among native grasses in controlling wind erosion [10]. It is recommended for rehabilitating surface-mined lands [92], and has been successfully established on bentonite [87] and coal mine [98] spoils. OTHER USES AND VALUES : Buffalo grass is planted for lawns [42,66,103] and used in hayfield mixtures. It is of limited use as a hay grass when planted alone, however, due to its short stature [89]. The sod houses of early Great Plains settlers were constructed mostly from buffalo grass [52]. MANAGEMENT CONSIDERATIONS : Buffalo grass is highly resistant to grazing [5,9,16]. It usually increases under heavy grazing, especially at the expense of tallgrass species [51,76]. Its response to grazing may vary by site, however. On the Central Plains Experimental Range in Colorado, Archer and Tieszen [9] found that buffalo grass importance value increased with moderate to heavy continuous grazing on ridges and midridges, but decreased with such grazing on swales. Importance value was low on all sites with light grazing. Buffalo grass responses to various other grazing regimes are described [66,74,93]. Buffalo grass is highly drought resistant, although somewhat less so than blue grama [89]. Buffalo grass seed is commerically available [43,81]. Guidelines for seeding buffalo grass onto rangeland, or establishing it from cut sod, are available [10,21,31,43] Marcum and Engelke [65] provide field test results on buffalo grass response to various pre- and postemergent herbicides.

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Buchloe dactyloides | Buffalo Grass
GENERAL BOTANICAL CHARACTERISTICS : Buffalo grass is a warm-season, native perennial shortgrass [20]. It is drought-, heat-, and cold-resistant [106]. Foliage is usually 2 to 5 inches (5-13 cm) high, though in the southern Great Plains foliage may reach 12 inches (30 cm) [55,100]. Buffalo grass is usually dioecious. Plants are occasionally monoecious, sometimes with perfect flowers [48,49,54,61]. Flowerstalks are 4 to 8 inches (10-20 cm) tall [100]. The male inflorescense is a panicle; the female inflorescense consists of short spikelets borne in burlike clusters, usually with two to four spikelets per bur [48,104]. Buffalo grass sends out numerous, branching stolons [54,106]; occasionally it also produces rhizomes [85]. Roots are also numerous and thoroughly occupy the soil [100]. The numerous stolons and roots form a dense sod [54,106]. Buffalo grass roots are finer than those of most plains grasses, being less than 1 mm in diameter. Weaver [100] found that in the Great Plains, buffalo grass roots in silty loam reached 5 feet below ground, with 70 percent of roots (by weight) occurring in the first 6 inches (15 cm) of soil. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Primarily dioecious, buffalo grass usually outcrosses by wind pollination [77]. The monoecious sex form is more common in peripheral, sparse populations, and monecious plants may be self-fertile [56]. Seed production on native grasslands in Kansas was greatest when above-average rainfall occurred in May and June; seed production was lowest when rainfall during that period was below average [27]. At dispersal the entire bur, with seeds still inside, abscises from the plant. Burs usually disperse close to the parent plant; they cling only briefly, if at all, to animals or clothing [76]. Fresh seed is usually dormant, requiring abrasion or overwintering to break dormancy. Degree of dormancy, however, has been found to vary between populations [78]. Light is required for germination [43]. Ahring and Todd [2] found that prechilling at 41 to 50 degrees Fahrenheit (5-10 deg C), drying seeds for 6 to 48 hours at 104 to 158 degrees Fahrenheit (40-70 deg C), or soaking seeds 1 to 72 hours in sodium hypochlorite greatly increased germination. Fulright and others [43] provide a summary of recommended buffalo grass seed stratification regimes. Reports of germinative capacity of fresh seed that has been stratified and scarified range from 45 to 70 percent. Seed has remained viable in the seedbank for at least 7 years [29]. Twenty-five-year-old seed recovered from the walls of an abandoned sod house in Kansas was 15 to 78 percent viable [64]. Germination rates are usually better when seed remains within the bur [76]. In a seedbank study on the Central Plains Experimental Range, Colorado, buffalo grass seedling density in fine-textured soil samples gathered over a 15-month sampling period averaged 112 plants per square meter [25]. Seedlings grow rapidly under favorable conditions. They have produced stolons by 2 months of age. By the end of their first summer, seedlings grow a thick root mass that extends 2 feet (0.6 m) below ground [100]. Vegetative reproduction through spreading stolons is rapid given favorable environmental conditions [29]. Buffalo grass stolons have grown as rapidly as 2.25 inches (5.72 cm) per day [69]. Buffalo grass also spreads by tillering [106]. Following drought in western Kansas, buffalo grass cover increased vegetatively from 2 percent in 1940 to 93 percent in 1943. Buffalo grass has been reported to spread vegetatively following depletion even when heavily grazed [101]. Although blue grama is more drought resistant, buffalo grass usually recovers more quickly after drought through vegetative reproduction [29]. SITE CHARACTERISTICS : Buffalo grass occurs on all soil textures [10,21]. It is most common on clay, then fine-textured loam. It is rare on sandy soils [10,81]. Soils supporting buffalo grass are usually alkaline [10,19,49], with a high water-holding capacity (33-66%) [49]. Buffalo grass grows mostly on dry uplands and/or disturbed sites in mixed- and tallgrass prairie [3,48]. It grows best in regions of the Great Plains where annual precipitation ranges between 12 and 25 inches (305 and 635 mm) [106]. It is widely adapted, however, to extremes in climate and elevation [21,106]. Elevation at which buffalo grass occurs ranges from 2,000 feet (600 m) in Montana to 6,300 feet (1,890 m) in Wyoming [36]. SUCCESSIONAL STATUS : Buffalo grass appears in early to mid-stage secondary succession [4]. Costello [28] described an old-field successional sequence in north-central Colorado where the inital stage was dominated by annuals, followed by perennial forbs. Buffalo grass appeared after the perennial forbs, 10 to 20 years after abandonment. Buffalo grass is common on disturbed sites such as prairie dog towns [1,117]. In a pattern similar to old-field succession, buffalo grass established after forbs and three-awns (Aristida spp.) but before mid-grasses such as silver bluestem (Bothriochloa saccharoides) and sideoats grama (Bouteloua curtipendula) on an abandoned black-tailed praire dog town in Oklahoma tallgrass prairie [71]. Buffalo grass often establishes dominance in mixed-grass prairie with drought or heavy grazing, and may invade tallgrass prairie under such conditions [59,89]. Buffalo grass may not require disturbance in order to maintain dominance on all sites, however. It dominated fine-textured clay soils in a remnant mixed-grass Kansas prairie that had not been grazed or burned for 68 years [57]. Van Auken and Bush [97] found that honey mesquite (Prosopis glandulosa var. glandulosa) was unable to invade high-density buffalo grasslands in Texas. Buffalo grass is somewhat shade tolerant, but cannot tolerate dense shade [10,106]. Clements [23] described buffalo grass-blue grama shortgrass prairie as "proclimax:" a community held indefinitely from reaching "climax condition" by the natural disturbances of grazing and fire. SEASONAL DEVELOPMENT : Buffalo grass growth begins in late spring and continues through summer [54,89]. In the Great Plains flowering occurs mostly from April to June, but male plants may flower until late summer or fall [48]. Period of heaviest flowering varies by location. Flowering is reported from July through August in eastern Colorado [35]. August flowering is also reported from Kansas, with male plants beginning flowering slightly before female plants [103]. Seed ripens from early summer to late fall, depending upon location. Early July seed ripening is reported from Kansas [27].

FIRE ECOLOGY

SPECIES: Buchloe dactyloides | Buffalo Grass
FIRE ECOLOGY OR ADAPTATIONS : Fire was an essential component of presettlement prairie ecosystems [6,18,76,82,99,116]. Prior to European settlement, buffalo grass probably burned in all seasons [12]. Areas dominated by warm-season grasses such as buffalo grass can carry fire even in winter and early spring [12], and Native Americans apparently burned various portions of the prairie year-round [7]. Buffalo grass survives grassland fire by several mechanisms. Vegetative regeneration is probably most important. Basal meristems are generally protected from grassland fire by soil and/or damp litter. Some stolon apices are usually protected by damp litter [107]. Buffalo grass regenerates after fire by basal tillering and sprouting from unburned stolon buds. Recovery time greatly accelerates when plants become vigorous enough to spread by stolons. Buffalo grass also regenerates from seed following fire [108]. Because it is enclosed in a bur, buffalo grass seed is more protected from fire than seed of most grass species. Buffalo grass burs have been shown to greatly reduce fire and heat damage to enclosed seed [76]. Regeneration from seed is slower than vegetative regeneration, however, and is probably most important when severe fire has killed a large proportion of stolon and basal buds. POSTFIRE REGENERATION STRATEGY : Caudex, growing points in soil Ground residual colonizer (on-site, initial community)

FIRE EFFECTS

SPECIES: Buchloe dactyloides | Buffalo Grass
IMMEDIATE FIRE EFFECT ON PLANT : Buffalo grass is not completely consumed in most grassland fires. Typically the upper part of the plant burns, and damage to basal portions of the plant is uncommon [45,99]. With grassland fires, flames fanned by even light winds seldom stay in one spot long enough to produce high temperatures at the soil surface. Fire intensities lethal to native perennial grasses such as buffalo grass rarely, if ever, occur during prescribed grassland fire [45]. Even with wildfires, temperatures near perennating tissues at the soil surface are usually not lethal. Unburned stubble often remains after fire has passed, and shallowly placed buds and seeds are unharmed [32 and references therein]. Wildfire occurring during drought, however, may generate temperatures high enough to kill buffalo grass perennating buds [32,45]. Lethal temperatures may also occur at the soil surface if woody plant invasion into grassland has occurred. Soil surface temperatures tend to rise when woody plants burn, and elevated temperatures last for longer periods of time [32]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Burning generally either favors buffalo grass or has no long-term effect upon it [108,110]. In some studies, buffalo grass productivity is unchanged or increases in the first postfire year [14,15]. Wright [110] concluded from several studies that in western Texas, buffalo grass was neither harmed nor favored by fire. In southern Nebraska, April burning of loess hill mixed-grass prairie had no significant (p=.1) effect on buffalo grass cover the following June or Septemeber. However, buffalo grass cover on burn plots had increased significantly compared to control plots by the second September after fire [84]. Fire had no long-term effect on buffalo grass at the Kansas Agricultural Experimental Station. Launchbaugh [63] reported that after a March wildfire in shortgrass prairie there, buffalo grass cover at postfire year 1 was reduced by 48 percent on burned areas as compared to adjacent unburned areas; height at the end of the first growing season was 6.7 inches (17.0 cm) on burned sites and 11.9 inches (30.2 cm) on unburned sites. By postfire year 2, buffalo grass cover on burned sites was 39 percent less than on unburned sites, and by postfire year 3 there was no significant difference in buffalo grass cover between burned and unburned sites. Buffalo grass recovery time may vary depending upon phenological stage, season of burning, fire severity, and/or postfire weather conditions. In a bluestem (Andropogon gerardi and Schizacharium scoparium) pasture in Kansas, buffalo grass declined under 10 years of early (20 March) and late spring (1 May) annual burning compared to annual mid-spring (10 April) and no burning. Buffalo grass basal cover (%) after 10 years was [5]: Spring Burning --------------------------------- unburned early mid- late 2.21a 1.08b 2.65a 1.37a --------------------------------- Percentages followed by the same letter do not differ significantly (p<0.05). Spring (April) prescribed burning in mixed-grass praiaire in Badlands National Park, South Dakota, favored buffalo grass. Buffalo grass began vegetative expansion and produced seed during the first growing season after fire [107]. Compared to the control (no burn), buffalo grass standing crop increased for 2 to 3 postfire years, then returned to approximate prefire levels with onset of a drought [108]. Buffalo grass increased significantly (p=0.05) after various treatments involving prescribed burning on the South Texas Plains-Texas Gulf Prairie interface. Burning was effected to reduce woody plant invasion. Treatments were shredding, chopping, or scalping followed by prescribed burning 2 years later, and a control (prescribed burning only). All prescribed burning was done in September 1965. Percentage composition of buffalo grass in July 1966 was [14]: ------------------------------------------------------------------------- Control Shredded Chopped Scalped Average ------- -------- ------- ------- ------- U B U B U B U B U B 13 15 11 12 6 17 6 17 9 15 ------------------------------------------------------------------------- U=unburned; B=burned Brush cover was significantly (p=0.05) reduced from prefire levels at postfire year 1, although less than 15 percent of woody plants were actually killed by the fire [14]. In another southern Texas study on the Rob and Bessie Welder Wildlife Refuge, plots were subjected to a fall (September) fire, a winter (December) fire, or a fall fire with a winter reburn the following year. Burning was conducted in 1965 and 1966. Buffalo grass production (lb/acre) in August 1967 was [15]: Control Fall Winter Fall & Winter 355 330 315 401 April prescribed burning in cultivated buffalo grass in Kansas reduced subsequent summer seed yield. Unburned portions of the field produced 303 pounds of buffalo grass seed per acre compared to 79 pounds per acre on the burned portion [27]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Fall burning in a shortgrass-mixed-grass transition zone of the Flint Hills of Kansas reduced prairie threeawn (Aristida oligantha), an annual grass with little to no forage value for livestock, and increased relative abundance of the dominant perennial grasses, buffalo grass and blue grama. Percentages of total herbage production in fall, 1972, with no burning, fall burning, and spring burning, were [72]: prairie threeawn Perennial grasses Western ragweed ---------------- ------------------ --------------- unburned 73.7 21.2 5.0 spring (1) 84.0 14.0 2.0 fall (2) 13.7 75.8 7.0 ----------------------------------------------------------------------- 1=burned 4 April 1972; 2=burned 8 November 1971 April prescribed fire in mixed-grass prairie of southern Nebraska also reduced nonnative, cool-season annuals and increased the native, warm-season dominants, buffalo grass and blue grama [84]. There was no significant relationship between fireline intensity and postfire response of buffalo grass after spring burning in western Texas grassland. High-intensity (approximately 5,570 kW/m) headfire did no more damage to buffalo grass than low-intensity (approximately 70 kW/m) headfire [83]. Buffalo grass mortality may be higher with backfires than headfires. Being more slow-moving, backfires tend to generate more heat at ground level [45]. Wright has provided prescriptions for burning buffalo grass in the central and southern Great Plains [111], the Edwards Plateau [115] and Rio Grande Plains [113,114] regions of Texas, and in chained mesquite-tobosa communities [112].

FIRE CASE STUDIES

SPECIES: Buchloe dactyloides | Buffalo Grass
CASE NAME : Postfire forage production in Custer State Park, SD REFERENCE : Easterly, T. G.; Jenkins, K. J. 1991 [40] SEASON/SEVERITY CLASSIFICATION : spring/variable STUDY LOCATION : The study area was on the north edge of French Creek Natural Area in Custer State Park, South Dakota. PREFIRE VEGETATIVE COMMUNITY : The study area was a mosaic of mixed-grass prairie and ponderosa pine (Pinus ponderosa) forest. Prairie was dominated by buffalo grass (Buchloe dactyloides), bluegrasses (Poa spp.), gramas (Bouteloua spp.), and western wheatgrass (Pascopyrum smithii). The forest understory was dominated by sedges (Carex spp.) and bluegrasses. TARGET SPECIES PHENOLOGICAL STATE : NO-ENTRY SITE DESCRIPTION : General: Site description was not given in detail. Aspect and slope were variable. Mean annual temperature at Custer, South Dakota (20 km northwest), was 42 degrees Fahrenheit (6 deg C). Mean annual precipitation was 18.4 inches (460 mm). Annual precipitation at Custer State Park was 14.9 inches (372 mm) in 1987 and 13.5 inches (337 mm) in 1988. Burn days: Peak temperature was 75 deg Fahrenheit (23 deg C) on 27 April 1987 and 82 degrees Fahrenheit (28 deg C) on 28 April 1987. Relative humidity ranged from 17 to 34 percent. Winds were from the south to southeast at 13 to 42 miles per hour (8-25 km/hr). Fine fuel moisture was approximately 5 percent. FIRE DESCRIPTION : Primary goals of the study were to reduce encroachment of ponderosa pine onto the prairie and increase forage available to bighorn sheep. The 235-acre (94-ha) burn was conducted on 27 and 28 April 1987. Strip-firing techniques were used to meet different burn objectives in different areas. In meadows and sites dominated by cool-season species, fire intensity was low and flame length was less than 12 inches (30 cm). Green-up of cool-season herbaceous species in the fall before the fire resulted in an incomplete burn pattern with large unburned patches. Sites dominated by warm-season species including buffalo grass burned hotter than cool-season sites, with flame lengths of 18 to 30 inches (45-75 cm). Fire intensities in forested areas varied widely depending upon fuels, aspect, time of day, and width of firing strip. Some direct mortality of pines occurred due to fire in tree crowns. FIRE EFFECTS ON TARGET SPECIES : Percent cover of buffalo grass in July was: 1987 1988 __________________ __________________ Burned Unburned Burned Unburned 4 2 5 3 FIRE MANAGEMENT IMPLICATIONS : Spring prescribed burning increased buffalo grass cover within the 2-year study period, and reduced ponderosa pine cover. Burning in mixed-grass prairie increased production of forbs and warm-season grasses including buffalo grass. Utilization of herbaceous species by elk, deer, and bison was higher in burned than unburned areas. Of all the Park ungulates, bison used the burns most intensively, with their use peaking in summer. Prescribed burning did not, however, increase the amount of forage available to bighorn sheep in late summer and presumably, winter. Mortality of ponderosa pine over the study area was approximately 50 percent. Pine expansion into mixed-grass prairie was curtailed.

REFERENCES

SPECIES: Buchloe dactyloides | Buffalo Grass
REFERENCES : 1. Agnew, William; Uresk, Daniel W.; Hansen, Richard M. 1986. Flora and fauna associated with prairie dog colonies and adjacent ungrazed mixed-grass prairie in western South Dakota. Journal of Range Management. 39(2): 135-139. [295] 2. Ahring, Robert M.; Todd, Glenn W. 1977. The bur enclosure of the caryopses of buffalograss as a factor affecting germination. Agronomy Journal. 69: 15-17. [2874] 3. Albertson, F. W. 1937. Ecology of mixed prairie in west central Kansas. Ecological Monographs. 7: 483-547. [5057] 4. Andelt, William F.; Kie, John G.; Knowlton, Frederick F.; Cardwell, Dean. 1987. Variation in coyote diets associated with season and successional changes in vegetation. Journal of Wildlife Management. 51(2): 273-277. [19860] 5. Anderson, Kling L.; Smith, Ed F.; Owensby, Clenton E. 1970. Burning bluestem range. Journal of Range Management. 23: 81-92. [323] 6. Anderson, R. C. 1970. Prairies in the Prairie State. Transactions of the Illinois State Academy of Sciences. 63(2): 214-221. [25838] 7. Anderson, Roger C. 1990. The historic role of fire in the North American grassland. In: Collins, Scott L.; Wallace, Linda L., eds. Fire in North American tallgrass prairies. Norman, OK: University of Oklahoma Press: 8-18. [14192] 8. Ansley, R. J.; Jacoby, P. W.; Cuomo, G. J. 1990. Water relations of honey mesquite following severing of lateral roots: influence of location and amount of subsurface water. Journal of Range Management. 43(5): 436-442. [14111] 9. Archer, Steven R.; Tieszen, Larry L. 1986. Plant response to defoliation: Hierarchical considerations. BioScience. 41(4): 236-247. [18790] 10. Beetle, Alan A. 1950. Buffalograss--native of the shortgrass plains. Bull. 293. Laramie, WY: University of Wyoming, Agricultural Experiment Station. 31 p. [414] 11. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. 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Related categories for Species: Buchloe dactyloides | Buffalo Grass

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