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Introductory

SPECIES: Carex filifolia | Threadleaf Sedge
ABBREVIATION : CARFIL SYNONYMS : NO-ENTRY SCS PLANT CODE : CAFI COMMON NAMES : threadleaf sedge TAXONOMY : The fully documented scientific name of threadleaf sedge is Carex filifolia Nutt. [11,12]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Crystal Walkup, June 1991 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Walkup, Crystal J. 1991. Carex filifolia. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Carex filifolia | Threadleaf Sedge
GENERAL DISTRIBUTION : Threadleaf sedge is distributed over much of the western prairies of North America from the Yukon Territory south to Washington (east of the Cascade Mountains), Oregon, northeastern Nevada, Utah, and New Mexico, and east through the Rocky Mountains and onto the plains of Manitoba, Nebraska, and Texas [11,12]. ECOSYSTEMS : FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES20 Douglas-fir FRES21 Ponderosa pine FRES23 Fir - spruce FRES29 Sagebrush FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES36 Mountain grasslands FRES37 Mountain meadows FRES38 Plains grasslands FRES39 Prairie STATES : AK AZ CA CO ID KS MN MT NE NV NM ND OK OR SD TX UT WA WY AB BC MB SK YT ADMINISTRATIVE UNITS : AGFO BADL BIHO BITH BICA CHCU CRMO DENA DETO FOBO GLAC SCBL THRO WICA YELL BLM PHYSIOGRAPHIC REGIONS : 4 Sierra Mountains 5 Columbia Plateau 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : KO11 Western ponderosa forest KO12 Douglas-fir forest KO15 Western spruce - fir forest KO16 Eastern ponderosa forest KO17 Black Hills pine forest KO18 Pine - Douglas-fir forest KO20 Spruce - fir - Douglas-fir forest KO21 Southwestern spruce - fir forest KO23 Juniper - pinyon woodland KO37 Mountain-mahogany - oak scrub KO38 Great Basin sagebrush KO51 Wheatgrass - bluegrass KO55 Sagebrush steppe KO56 Wheatgrass - needlegrass shrubsteppe KO63 Foothills prairie KO64 Grama - needlegrass - wheatgrass KO65 Grama - buffalograss KO66 Wheatgrass - needlegrass KO67 Wheatgrass - bluestem - needlegrass KO68 Wheatgrass - grama - buffalograss KO70 Sandsage - bluestem prairie KO74 Bluestem prairie KO75 Nebraska Sandhills prairie KO81 Oak savanna KO98 Northern floodplain forest SAF COVER TYPES : 42 Bur oak 201 White spruce 206 Engelmann spruce - subalpine fir 210 Interior Douglas-fir 216 Blue spruce 238 Western juniper 239 Pinyon - juniper 243 Sierra Nevada mixed conifer 251 White spruce - aspen 256 California mixed subalpine SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Threadleaf sedge forms a climactic climax with needle-and-thread grass (Stipa comata) in protected areas of Theodore Roosevelt National Park (TRNP) [9]. Blue grama (Bouteloua gracilis) dominates unprotected areas which have been heavily grazed. Some consider the presence of blue grama an indication of xeric soils [14], but the majority believe that blue grama replaces threadleaf sedge as a result of overgrazing [3,8,9,36]. An edaphic climax of wheatgrass (Agropyron spp.)/sedge (Carex spp.) also occurs in Theodore Roosevelt National Park. Following is a list of habitat type (hts) and community type (cts) publications in which threadleaf sedge is listed as a dominant or subdominant. Analysis of grassland vegetation on selected key areas in southwestern north Dakota [33]. Grassland and shrubland habitat types of western Montana [18]. Range plant communities of the Central Grasslands Research Station in south central North Dakota [15]. The vegetation of the Grand River/Cedar River, Sioux, and Ashland Districts of the Custer National Forest: a habitat type classification [9].

VALUE AND USE

SPECIES: Carex filifolia | Threadleaf Sedge
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Threadleaf sedge provides excellent forage for several livestock and wildlife species, although production is very low [25]. It provides extremely valuable early spring growth and remains palatable throughout the growing season [21,25,34]. Threadleaf sedge is an important winter and spring forage for both elk and bison [37,38]. It is also one of the five main forages of the black-tailed prairie dog [26]. Domestic animals such as cattle, sheep, and horses also use this species [5]. PALATABILITY : Although palatability varies widely with location, it is generally fair to good for livestock and wildlife [5]. Young plants are preferred, but threadleaf sedge maintains high palatability throughout the growing season [21,25]. Palatability has been rated at 70 percent for cattle [21]. The relish and degree of use shown by livestock and wildlife species for threadleaf sedge in several western states has been rated as follows [7]: CO MT ND UT WY Cattle Fair Good Good Fair Fair Sheep Good Good Good Fair Good Horses Good Good Good Fair Fair Elk ---- Fair ---- Fair Good Mule deer ---- Poor Good Fair Fair White-tailed deer Fair Good ---- ---- ---- Pronghorn ---- Poor Good Fair Fair Upland game birds ---- ---- ---- Fair ---- Waterfowl ---- Fair ---- Poor ---- Small nongame birds ---- ---- ---- Fair ---- Small mammals ---- ---- ---- Fair Good NUTRITIONAL VALUE : The nutrient content of threadleaf sedge has been reported as follows [18]: (Protein is more digestible in fresh grass than in overripe grass). N-free Ether Crude Protein Extract Extract Fiber Ash Fresh 7.5 45.6 3.2 32.9 10.8 Overripe 4.1 43.4 3.2 37.5 11.8 COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : The ability of the root system of this species to hold soil in place makes it an ideal candidate for controlling wind- and water-induced soil erosion. It exhibits good growth on moderate slopes and fair growth on steep slopes. Erosion-control potential has been rated high for Colorado, and medium for Utah, Montana, Wyoming, and North Dakota. Long-term revegatation potential has been rated medium for Utah, Wyoming, Montana, and North Dakota [7]. Limited seed availability restricts the use of this plant. Native seed harvests would be costly due to the low seed production, and currently no commercial sources are available. OTHER USES AND VALUES : Food: American Indians used culm bases as food during famines [25]. Drought tolerance: Threadleaf sedge has value as a drought-tolerant species [31]. MANAGEMENT CONSIDERATIONS : Threadleaf sedge decreases under heavy grazing pressure. Reproduction is almost entirely from seed, so if the stand is grazed so closely that no seed production occurs, it will soon deteriorate [5]. Plants lose vigor when overgrazed and are eventually replaced by blue grama and buffalograss (Buchloe dactyloides), which are more resistant to close grazing [10,31].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Carex filifolia | Threadleaf Sedge
GENERAL BOTANICAL CHARACTERISTICS : Threadleaf sedge is a perennial, native, cool-season grasslike plant with a densely caespitose growth habit [25]. It reaches up to 12 inches (30 cm) in height. Its root system binds the soil more firmly than that of most grasses. Although very small, (1 mm or less in diameter) roots occur in enormous numbers. Root systems extend farther laterally (as far as 15 inches [38 cm] away from the plant) than vertically [4,28]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Sexual: Plants are monoecious; male flowers occur on the upper portion of the floret and female flowers on the lower portion. Fruits are triangular achenes [7]. Flowers are often damaged by frost, preventing seed production [21]. Vegetative: Threadleaf sedge can reproduce vegetatively by rhizomes, but this is uncommon. The majority of reproduction is by seed [7]. SITE CHARACTERISTICS : Threadleaf sedge is typically found on dry soils of open prairies and rolling hills. It can grow on moist sites but not in shallow water or swamps [25]. Threadleaf sedge generally occurs on the upper slopes of hills but can occur on all aspects [14]. It usually grows in full sunlight, although fair stands sometimes occur in the shade [5]. Soils: Threadleaf sedge is generally found on deep, loamy soils in meadows and shallow rocky soils on slopes [5]. Production is fair to good on all soil types except clay to dense clay. Optimum production occurs on sandy loams and loams with a depth of 10 to 20 inches (26-51 cm) [7]. Elevation: The elevational range is wide due to the wide distribution of the plant. In the Northwest and Canada it can be found within several hundred feet of sea level, but in the Southern Rocky Mountains it occurs at timberline (12,000 feet [3,658 m]) and has been reported at 13,000 feet (3,958 m) in Colorado [5,7]. SUCCESSIONAL STATUS : Threadleaf sedge decreases in response to heavy grazing and is eventually removed from the plant community. In Nebraska it was identified as a codominant with needlegrass (Stipa spp.) in a mixed-prairie climax community. Under grazing pressure, needlegrass decreased and was replaced by blue grama. With continued heavy grazing, buffalograss replaced threadleaf sedge [27]. Results of a study in southwestern North Dakota showed cover of threadleaf sedge was significantly higher within exclosures than without [2]. Blue grama appeared to replace threadleaf sedge. SEASONAL DEVELOPMENT : Seasonal development varies slightly according to elevation. Growth starts in late March to early April in North Dakota [21], early April in Nebraska [27], and April in Colorado [6]. Flowering occurs from May to June in North Dakota, late April in Nebraska, late April to May in Colorado, May in Wyoming [7], and April to July in Montana [7]. Seed dissemination starts the first of June in Nebraska and mid-May in Colorado. Seed dispersal is completed by the end of July in North Dakota. The plant is dried by early to mid-November in Colorado.

FIRE ECOLOGY

SPECIES: Carex filifolia | Threadleaf Sedge
FIRE ECOLOGY OR ADAPTATIONS : Threadleaf sedge is generally able to survive fire due to basal meristems protected by soil and/or damp litter [30]. The low, dense growth habit allows for good continuity of fuel, which enhances fire spread [24]. POSTFIRE REGENERATION STRATEGY : Caudex, growing points in soil

FIRE EFFECTS

SPECIES: Carex filifolia | Threadleaf Sedge
IMMEDIATE FIRE EFFECT ON PLANT : Threadleaf sedge has been alternately reported as severely damaged or minimally damaged by fire. Degree of damage depends in part on season and intensity of the fire, and postburn precipitation. Of several publications listing threadleaf sedge as severely damaged by fire, only one was a report of an actual prescribed burn [17,20,23,35]. The fire was described as a "hot, clean burn" [17]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Threadleaf sedge recovers vegetatively, with surviving portions of the plants sending up new growth. Establishment occurs readily at any time following fire, with no adverse effects from competition with surviving plants of the same or other species. Reproductive maturity is usually not reached until the second year following the burn [30]. Postfire seed production is often prolific, possibly due to increased nutrients and available soil moisture [1]. An upland site with below average productivity took 1 to 3 years longer to recover following fire than more productive sites did [30]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : Low postburn precipitation may delay full recovery until postfire year 2 or 3 or longer, depending on the severity of the burn [30]. In South Dakota, productivity was increased by burning in April and October when precipitation was above average but was reduced when postburn precipitation was low [30]. Threadleaf sedge which was burned by a low-severity fire and which received adequate precipitation after the burn recovered relatively quickly (2 to 4 years after the fire). However, severe burns may cause more lasting effects, requiring 12 to 15 years for the plant community to recover [1]. Season of burn may also affect postfire response. In Montana threadleaf sedge was relatively unaffected by spring burning but reduced by fall burning [22]. However, in western North Dakota there was no reduction 4 years after a late summer burn, 3 years after a fall burn, or 3 months after a spring burn [8]. Therefore, if postfire precipitation is adequate, it appears that light- to moderate-severity fires (particularly spring fires) often cause only minimal damage to threadleaf sedge. FIRE MANAGEMENT CONSIDERATIONS : Prescribed burning to reduce sagebrush often involves plant communities with abundant threadleaf sedge. After burning, careful grazing management should be implemented to ensure full recovery of the plants [20]. To maintain a good stand, plants should not be burned during periods of drought, and burn severity should be light to moderate [2]. If late summer grazing is desired, burning threadleaf sedge in the fall or spring may allow leaves to remain on the plants longer than they would on unburned plants [32]. This provides forage later in the season.

FIRE CASE STUDIES

SPECIES: Carex filifolia | Threadleaf Sedge
CASE NAME : Northern Great Plains/Yield and Cover/Prescribed Burning REFERENCE : White, R. S.; Currie, P. O. 1983 [32] SEASON/SEVERITY CLASSIFICATION : Fall - 2nd week in October/severity not reported Spring - April 3/severity not reported STUDY LOCATION : Livestock and Range Research Station - Miles City, Montana PREFIRE VEGETATIVE COMMUNITY : Three communities were involved in this study: western wheatgrass (Pascopyrum smithii), blue grama (Bouteloua gracilis, and threadleaf sedge. Communities were relatively uniform, with pure stands of each of the three species. This minimized errors in yield and cover measurements that frequently occur when grass separation is required. TARGET SPECIES PHENOLOGICAL STATE : Fall burn: Threadleaf sedge was dormant. Spring burn: Prior to onset of spring growth. SITE DESCRIPTION : Soils of the study sites ranged in texture from loam to silt loam. All had good drainage and high fertility. Topography was level. Precipitation for the site is approximately 13.4 inches (34 cm) annually, with 60 percent received between May and August. During the postburn growing season, precipitation was near normal for June, but below normal for July and August. FIRE DESCRIPTION : Fires were confined by panels constructed of angle-iron and sheet metal. Plot size was sufficiently large (3 x 3 m) to approximate burning under natural conditions. FIRE EFFECTS ON TARGET SPECIES : Yield: Spring burning of threadleaf sedge yielded about the same amount of herbage through June as the unburned control. In contrast, fall burning showed about the same peak season yields, but until mid-June, yields were significantly lower than both the control and spring-burned plots. This resulted in 25 percent less herbage production on the fall-burned treatments. Basal Cover: Burning treatments substantially reduced the litter component and increased the proportion of bare soil in the threadleaf sedge community. Spring burning resulted in the highest threadleaf sedge cover, but there were no significant differences among other treatments. FIRE MANAGEMENT IMPLICATIONS : If grazing is needed in early June, neither spring nor fall burning should be undertaken. Plants on burned treatments never outyielded those on the control at that time. Spring or fall burning may effectively stimulate forage production for the rest of the grazing season. Available moisture must be considered before burning. Periods of drought place stress on the plants, making them less likely to recover from the burn.

REFERENCES

SPECIES: Carex filifolia | Threadleaf Sedge
REFERENCES : 1. Blaisdell, James P. 1953. Ecological effects of planned burning of sagebrush-grass range on the upper Snake River Plains. Tech. Bull. 1975. Washington, DC: U.S. Department of Agriculture. 39 p. [462] 2. Brand, Michael D. 1980. Secondary succession in the mixed grass prairie of southwestern North Dakota. Fargo, ND: North Dakota State University. 77 p. Dissertation. [14147] 3. Coupland, Robert T. 1961. A reconsideration of grassland classification in the northern Great Plains of North America. Journal of Ecology. 49: 135-167. [12588] 4. Coupland, Robert T.; Johnson, R. E. 1965. Rooting characteristics of native grassland species of Saskatchewan. Journal of Ecology. 53: 475-507. [702] 5. Dayton, William A. 1931. Important western browse plants. Misc. Publ. 101. Washington, DC: U.S. Department of Agriculture. 214 p. [768] 6. Dickinson, C. E.; Dodd, Jerrold L. 1976. Phenological pattern in the shortgrass prairie. American Midland Naturalist. 96(2): 367-378. [799] 7. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 8. Dix, Ralph L. 1960. The effects of burning on the mulch structure and species composition of grasslands in western North Dakota. Ecology. 41(1): 49-56. [808] 9. Hansen, Paul L.; Hoffman, George R. 1988. The vegetation of the Grand River/Cedar River, Sioux, and Ashland Districts of the Custer National Forest: a habitat type classification. Gen. Tech. Rep. RM-157. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 68 p. [771] 10. Hansen, Paul L.; Hoffman, George R.; Bjugstad, Ardell J. 1984. The vegetation of Theodore Roosevelt National Park, North Dakota: a habitat type classification. Gen. Tech. Rep. RM-113. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 35 p. [1077] 11. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 12. Hitchcock, C. Leo; Cronquist, Arthur; Ownbey, Marion. 1969. Vascular plants of the Pacific Northwest. Part 1: Vascular cryptograms, gymnosperms, and monocotyledons. Seattle, WA: University of Washington Press. 914 p. [1169] 13. Weldon, Lyle W. 1956. A study of Artemisia tridentata Nutt. seedlings in relation to chemical control. Laramie, WY: Univeristy of Wyoming. 36 p. M.S. thesis. [55] 14. Lieffers, V. J.; Larkin-Lieffers, P. A. 1987. Slope, aspect, and slope position as factors controlling grassland communities in the coulees of the Oldman River, Alberta. Canadian Journal of Botany. 65: 1371-1378. [3957] 15. Lura, Charles L.; Barker, William T.; Nyren, Paul E. 1988. Range plant communities of the Central Grasslands Research Station in south central North Dakota. Prairie Naturalist. 20(4): 177-192. [7224] 16. McColley, Phillip D.; Hodgkinson, Harmon S. 1970. Effect of soil depth on plant production. Journal of Range Management. 23(3): 189-192. [1591] 17. Mueggler, Walter F.; Blaisdell, James P. 1958. Effects on associated species of burning, rotobeating, spraying, and railing sagebrush. Journal of Range Management. 11: 61-66. [1712] 18. Mueggler, W. F.; Stewart, W. L. 1980. Grassland and shrubland habitat types of western Montana. Gen. Tech. Rep. INT-66. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 154 p. [1717] 19. National Academy of Sciences. 1971. Atlas of nutritional data on United States and Canadian feeds. Washington, DC: National Academy of Sciences. 772 p. [1731] 20. Pechanec, Joseph F.; Stewart, George; Blaisdell, James P. 1954. Sagebrush burning good and bad. Farmers' Bulletin No. 1948. Washington, DC: U.S. Department of Agriculture. 34 p. [1859] 21. Sarvis, J. T. 1941. Grazing investigations on the Northern Great Plains. Bull. 307. Fargo, ND: North Dakota Experiment Station. 110 p. In cooperation with: U.S. Department of Agriculture, Northern Great Plains Field Station. [10853] 22. Senft, Dennis. 1983. Fire freshens rangeland. Agricultural Research. 32(3): 10-11. [2106] 23. Smith, Michael A.; Busby, Fee. 1981. Prescribed burning: effective control of sagebrush in Wyoming. RJ-165. Laramie, WY: University of Wyoming, Agriculture Experiment Station. 12 p. [2175] 24. Smith, Michael A.; Dodd, Jerrold L.; Rodgers, J. Daniel. 1985. Prescribed burning on Wyoming rangeland. Bull. 810. Laramie, WY: University of Wyoming Agricultural Extension Service. 25 p. [2176] 25. Stubbendieck, J.; Hatch, Stephan L.; Hirsch, Kathie J. 1986. North American range plants. 3rd ed. Lincoln, NE: University of Nebraska Press. 465 p. [2270] 26. Summers, Carol A.; Linder, Raymond L. 1978. Food habits of the black-tailed prairie dog in western South Dakota. Journal of Range Management. 31(2): 134-136. [2294] 27. Tolstead, W. L. 1942. Vegetation of the northern part of Cherry County, Nebraska. Ecological Monographs. 12: 255-292. [4470] 28. Weaver, J. E. 1958. Summary and interpretation of underground development in natural grassland communities. Ecological Monographs. 28(1): 55-78. [297] 29. Isaac, Leo A. 1963. Fire--a tool not a blanket rule in Douglas-fir ecology. In: Proceedings, 2nd annual Tall Timbers fire ecology conference; 1963 March 14-15; Tallahassee, FL. Tallahassee, FL: Tall Timbers Research Station: 1-17. [10700] 30. Whisenant, Steven G.; Uresk, Daniel W. 1989. Burning upland, mixed prairie in Badlands National Park. Prairie Naturalist. 21(4): 221-227. [11151] 31. White, Larry M.; Newbauer, John J., III; Wight, J. Ross. 1978. Vegetational differences on native range during 38 years in eastern Montana. In: Hyder, Donald N., ed. Proceedings of the 1st International Rangeland Congress; 1978 August 14-18; Denver, CO. Denver, CO: Society for Range Management: 260-262. [2535] 32. White, Richard S.; Currie, Pat O. 1983. Prescribed burning in the Northern Great Plains: yield and cover responses of 3 forage species in the mixed grass prairie. Journal of Range Management. 36(2): 179-183. [2541] 33. Whitman, Warren C. 1979. Analysis of grassland vegetation on selected key areas in southwestern North Dakota. Final Report on Contract No. 7-01-2. Bismark, ND: Regional Environmental Assessment Program. 199 p. [12529] 34. Williams, Thomas A. 1898. A report upon the grasses and forage plants and forage conditions of the eastern Rocky Mountain region. Bulletin No. 12. Washington, DC: U.S. Department of Agriculture, Division of Agrostology. 78 p. [4278] 35. Wright, Henry A.; Neuenschwander, Leon F.; Britton, Carlton M. 1979. The role and use of fire in sagebrush-grass and pinyon-juniper plant communities: A state-of-the-art review. Gen. Tech. Rep. INT-58. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Statio. 48 p. [2625] 36. Wright, John C.; Wright, Elnora A. 1948. Grassland types of south central Montana. Ecology. 29(4): 449-460. [2627] 37. Wydeven, Adrian P.; Dahlgren, Robert B. 1983. Food habits of elk in the northern Great Plains. Journal of Wildlife Management. 47(4): 916-923; 1983. [2630] 38. Wydeven, Adrian P.; Dahlgren, Robert B. 1985. Ungulate habitat relationships in Wind Cave National Park. Journal of Wildlife Management. 49(3): 805-813. [57] 39. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 40. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 41. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 42. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 43. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 44. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 45. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573]

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