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Wildlife, Animals, and Plants |
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BOTANICAL AND ECOLOGICAL CHARACTERISTICS
GENERAL BOTANICAL CHARACTERISTICS:Timber oatgrass is a native, strongly caespitose, perennial bunchgrass [23,67]. The erect culms are densely tufted and generally reach 4 to 20 inches (10-50 cm) in height [62]. Leaves are mainly basal, flat or involute [69]. The old sheaths and blades are often persistent and wither at the base of the plant [23]. The inflorescence is a narrow panicle, often 1-sided, with short mostly erect branches generally bearing 4 to 9 spikelets [10,23]. One- to 2-flowered spikelets occasionally occur in the axils of the lower leaves [69]. With age, the culms often separate at the nodes where these seed-bearing spikelets are borne. Timber oatgrass has a shallow and fibrous root system [58]. RAUNKIAER LIFE FORM:Chamaephyte REGENERATION PROCESSES:Timber oatgrass reproduces by seed and tillering [57]. This grass also produces self-fertilized spikelets in the axils of the lower leaves [69]. These cleistogamous spikelets enable the plant to reproduce even if development of the flower stalks is retarded [50]. In the Intermountain region, timber oatgrass is apparently largely apomictic (setting seed without fertilization), as the anthers are mostly abortive [10]. Mean germination of seed collected in southeastern British Columbia was 22.2% under laboratory conditions [54].SITE CHARACTERISTICS:Timber oatgrass grows in a wide range of habitats including rock outcrops, sphagnum bogs, dry meadows, grassy balds, and on alluvial flats of river floodplains [3,29,30,53]. It occurs on dry to moist sites from the prairies and grasslands to rocky alpine ridges [22]. In parts of the northern Rocky Mountains timber oatgrass is locally common in subalpine meadows in the fir-spruce zone [40,47]. In British Columbia, it grows in alpine tundra zones [1] and in the Sierra Nevada of California, it occurs in mountain meadows [56]. In fescue grasslands, timber oatgrass occurs in small patches or as a "fairly constant scattering" across the stand [24]. At high elevations timber oatgrass is most abundant in subalpine and alpine parks and meadows but also occurs in openings in upper elevation coniferous types, including fir-spruce and lodgepole pine communities [58,65,69]. It is common in ponderosa pine forests of the Black Hills of South Dakota [59]. Across its wide range, timber oatgrass grows from sea level to more than 12,000 feet (3660 m) in elevation [30]. Generalized elevational ranges of timber oatgrass are as follows [12,61,58,69]: 7,500 to 12,800 feet (2286-3901 m) in CO 3,200 to 8,900 feet ( 975-2713 m) in MT 4,950 to 6,780 feet (1509-2067 m) in SD 8,000 to 12,000 feet (2440-3660 m) in UT 5,000 to 11,000 feet (1981-3353 m) in WYSites at high elevations are often rocky with permeable, well-drained, shallow to deep soils [22,27,34]. Often, soils are characterized by relatively high soil moisture derived from snowmelt above [13]. Soils are commonly derived from shale, limestone, and other sedimentary materials, or from granite or redeposited volcanics [25,27]. Timber oatgrass commonly grows on loam and silt-loam soils [30]. Growth is described as good on organic and andic soils [12]. SUCCESSIONAL STATUS:Timber oatgrass is an indicator of climax in a number of subalpine and alpine grasslands, fescue grasslands, and forest communities. According to Beetle [3] timber oatgrass is "indicative of long-standing stability in the vegetation." In Nevada, it is often associated with "stable" riparian communities [38]. Depending on the type and severity of disturbance, timber oatgrass is also a constituent in some early seral communities. In lodgepole pine forests of Wyoming, timber oatgrass is most common in immature stands as compared with mature stands [3]. Similarly in ponderosa pine stands of South Dakota, it is more abundant and more productive in clearcut and heavily thinned sapling and pole stands (measured 8 to 15 years after treatment) than in unthinned stands [64]. SEASONAL DEVELOPMENT:Timber oatgrasss begins growth early in the spring [57]. Flowering occurs from June to August, depending on the location [50]. The seed crop is disseminated during August and September [58]. At lower elevations in California, flowerstalks may appear in April, with the seed cast by June [50]. At higher elevations, these dates are correspondingly later. In Colorado, Montana, and Wyoming, flowering begins in July and ends in August [12].
In western Montana, seasonal development varies according to
aspect as follows [44]: Southwest Northeast
range mean duration range mean duration
date (days) date (days)
growth
starts 4/28-5/24 5/12 10.2 4/28-5/29 5/16 11.9
1st
bloom 6/10-7/14 6/25 7.8 6/8-7/8 6/27 10.0
blooming
over 6/29-7/20 7/9 6.2 7/1-7/21 7/13 6.2
dissemination
starts 7/22-8/10 7/30 6.3 7/26-8/21 8/3 6.1
plant
dried 8/18-10/15 9/25 17.4 9/10-11/2 10/6 13.6
Related categories for SPECIES: Danthonia intermedia | Timber Oatgrass |
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