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Introductory

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
ABBREVIATION : ELYLAN SYNONYMS : Agropyron dasystachyum Agropyron albicans Agropyron elmeri Agropyron griffthsii Agropyron psammophilus Agropyron riparium SCS PLANT CODE : ELLA3 ELLAA ELLAL ELLAP COMMON NAMES : thickspike wheatgrass TAXONOMY : The currently accepted scientific name of thickspike wheatgrass is Elymus lanceolatus (Scribner & J. G. Smith) Gould. Barkworth and Dewey [3] combined the former taxa A. dasystachyum, A. elmeri, and A. riparium under E. lanceolatus. There are three subspecies of thickspike wheatgrass [3,4]: E. l. ssp. albicans (Scribner & J. G Smith) Barkw. & D. R. Dewey E. l. ssp. lanceolatus E. l. ssp. psammophilus (Gillett & Senn) A. Love LIFE FORM : Graminoid FEDERAL LEGAL STATUS : NO-ENTRY OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Kathy Ahlenslager, April 1986 LAST REVISED BY AND DATE : Kathy Ahlenslager, December 1987 AUTHORSHIP AND CITATION : Ahlenslager, Kathleen E. 1986. Elymus lanceolatus. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
GENERAL DISTRIBUTION : Thickspike wheatgrass occurs from Alaska east and south across Canada, south throughout the western United States and Mexico, in the Great Plains south to Kansas, and in the eastern United States as far south as West Virginia. It is best adapted to the northern Intermountain West and the drier areas of the Pacific Northwest where it is the most widely distributed sod-forming grass [7,22,23,34,36,41]. ECOSYSTEMS : FRES21 Ponderosa pine FRES23 Fir - spruce FRES29 Sagebrush FRES30 Desert shrub FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES36 Mountain grasslands FRES38 Plains grasslands STATES : AK AZ CA CO HI IL IN IA KS KY MI MN MO MT NV NM ND OH OR SD UT WA WV WI WY AB BC MB ON PQ SK MEXICO ADMINISTRATIVE UNITS : BLCA BRCA CODA DINO FOBU GLAC GRTE GRBA LABE SAJH SCBL SLBE THRO WICA YELL ZION BLM PHYSIOGRAPHIC REGIONS : 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K011 Western ponderosa forest K012 Douglas-fir forest K015 Western spruce - fir forest K016 Eastern ponderosa forest K017 Black Hills pine forest K018 Pine - Douglas-fir forest K021 Southwestern spruce - fir forest K023 Juniper - pinyon woodland K037 Mountain mahogany - oak scrub K038 Great Basin sagebrush K040 Saltbrush - greasewood K051 Wheatgrass - bluegrass K055 Sagebrush steppe K056 Wheatgrass - needlegrass shrubsteppe K057 Galleta - three-awn shrubsteppe K063 Foothills prairie K064 Grama - needlegrass - wheatgrass K065 Grama - buffalograss K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem prairie K098 Northern floodplain forest SAF COVER TYPES : 220 Rocky Mountain juniper 237 Interior ponderosa pine SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Thickspike wheatgrass is a codominant or indicator species in many habitat types in sagebrush-grassland vegetation. It is found in communities with Indian ricegrass (Oryzopsis hymenoides), sand dropseed (Sporobolus cryptandrus), big sagebrush (Artemisia tridentata), prairie sandreed (Calamovilfa longifolia), bluebunch wheatgrass (Pseudoroegernia spicata), Idaho fescue (Festuca idahoensis), and needlegrass (Stipa spp.) [40]. Thickspike wheatgrass is named as a dominant in the following vegetation classifications: Climax vegetation of Montana based on soils and climate [31] Plant associations (habitat types) of Region 2., 3rd ed. [37]

VALUE AND USE

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : NO-ENTRY PALATABILITY : The palatability of thickspike wheatgrass to cattle is good in the summer and fair in the winter. Palatability of this grass to elk is good yearlong; palatability to deer is fair in the spring [40]. The degree of use shown by livestock and wildlife species for thickspike wheatgrass in several western states is rated as follows [14]: CO MT ND UT WY Cattle Good Good Good Good Good Sheep Fair Fair Fair Fair Good Horses Fair Good Good Good Good Pronghorn ---- Poor Poor Fair Fair Elk ---- Fair Good ---- Good Mule deer ---- Poor Poor Good Fair White-tailed deer ---- Poor ---- ---- Poor Small mammals ---- Poor ---- Good Fair Small nongame birds ---- Poor ---- Fair Fair Upland game birds ---- ---- ---- Fair Fair Waterfowl ---- ---- ---- Fair Fair NUTRITIONAL VALUE : Compared to other grasses, thickspike wheatgrass is rated good in energy value and fair in protein value [14]. COVER VALUE : The degree to which thickspike wheatgrass provides environmental protection during one or more seasons for wildlife species is as follows [14]: MT ND UT WY Pronghorn Poor Fair Poor Poor Elk Poor ---- Poor Poor Mule deer Poor Fair Poor Poor White-tailed deer Poor Fair ---- Poor Small mammals Fair ---- Fair Good Small nongame birds Fair ---- Fair Good Upland game birds Fair ---- Fair Fair Waterfowl Good ---- Fair Fair VALUE FOR REHABILITATION OF DISTURBED SITES : Due to its drought tolerance and ability to form a dense sod, thickspike wheatgrass and cultivars of it are widely used for soil stabilization on disturbed range sites and dry areas subject to erosion, as well as on roadsides, airports, irrigation canal banks, recreation areas, construction sites, abandoned farm lands, and planting corners where circle move sprinklers are used [2,32,36]. The cultivar 'Critana' is adapted for use in Montana, Wyoming, and the western Dakotas in areas with mean annual precipitation of 10 to 16 inches (25-40 cm). It has been successfully used on mine spoils and in range seedings in Montana and Wyoming [35].. The cultivar 'Sodar' is widely used as a low-growing, low-maintenance cover in the Intermountain area. It is used for roadside seedings, recreation areas, disturbed areas, and other sites where a low-maintenance turf is desirable [35]. Vesicluar-arbuscular mycorrhizae are a common component of arid soils. They usually increase nutrient and water intake, as well as increase the dry mass of plants. The disturbance of land often leads to the reduction or elimination of mycorrhizal fungi propagules. Water relations in thickspike wheatgrass plants are not affected by inoculation of mycorrhizae or the presence of annuals [1]. OTHER USES AND VALUES : Although thickspike wheatgrass is often seeded alone, it is also seeded in mixtures for cover and forage. In mixtures with taller species, it is sometimes used as a spreading undercover plant [15,40]. Cultivars with good seed production and high seedling vigor include 'Sodar,' released in Idaho and Washington, 'Critana,' released in Montana, and 'Elbee,' released in Alberta [18,30,34]. Commercially available seeds of thickspike wheatgrass were used successfully to revegetate subalpine slopes of Montana ski areas [6]. MANAGEMENT CONSIDERATIONS : Weed control is usually needed on new seedings of thickspike. Stands should not be grazed until plants are firmly rooted. In the sandy soil where thickspike wheatgrass usually grows, trampling damage is often as serious as overgrazing. Livestock should be managed to prevent overuse and subsequent wind erosion problems. As soil becomes stabilized and plants get older, yields tend to decrease [32]. When seeded alone, a fertilization program helps to maintain high forage production [40].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
GENERAL BOTANICAL CHARACTERISTICS : Thickspike wheatgrass is a long-lived perennial, cool-season, drought-tolerant, coarse-leaved grass [36,39]. It is a sod-forming species with long, branched rhizomes, by which plants spread rapidly. The erect culms are 16 to 32 inches (40-80 cm) tall. In rich soils in the more eastern areas of its range, the root depth of thickspike wheatgrass plants may reach 6.8 to 11.7 feet (2.1-3.6 m). In the more arid western portions roots may penetrate to only 4.8 feet (1.5 m) [11]. Although morphologically similar to western wheatgrass (Pascopyrum smithii), thickspike wheatgrass is more drought resistant, albeit less productive [29]. RAUNKIAER LIFE FORM : Geophyte REGENERATION PROCESSES : Seeds usually germinate in 21 days under ideal laboratory conditions. Germination is improved by prechilling (stratification). Seedling vigor is fair, which is better than that of western wheatgrass (Pascopyrum smithii) but not as good as that of desert wheatgrass (A. desertorum) and intermediate wheatgrass (A. intermedium) [40]. Thickspike wheatgrass regenerates vegetatively through rhizomes, as well as sexually via wind-dispersed seeds. Due to its matted root system, other species have difficulty establishing in thickspike stands. Thickspike wheatgrass increases in frequency of individual plants and percent cover in response to grazing [16]. SITE CHARACTERISTICS : Thickspike wheatgrass occurs in grasslands and sagebrush deserts on low dry plains, dry hillsides, exposed flats and ridges, benches, and well-drained meadows. Plants rarely form pure stands over any extensive area, although they can appear to dominate and can form almost pure stands over a small area [36]. Although thickspike wheatgrass is best adapted to sand, sandy loam, and loam soils, it is also found on heavy alkaline flats and weakly saline soils. This grass grows on well-drained areas where the water table is 3 feet (1 m) or more beneath the surface [40]. Thickspike wheatgrass plants require from 8 to 15 inches (20-38 cm) of annual precipitation [29]. When growing in areas with more than 20 inches (51 cm) annual precipitation, plants are not long-lived, because of leaf and stripe rusts [32]. Plants occur over wide elevational ranges. These vary as follows [14]: from 4,800 to 10,000 ft (1,463-3,049 m) in CO 5,000 to 6,500 ft (1,524-1,982 m) in MT 3,800 to 8,900 ft (1,159-2,713 m) in WY SUCCESSIONAL STATUS : Thickspike is present in several climax plant communities [27]. It has the ability to increase in abundance during secondary succession [36]. SEASONAL DEVELOPMENT : This cool-season grass produces its best growth in spring. Thickspike wheatgrass blooms from May to July. Flowering dates are as follows [14]: State Earliest date observed Latest date observed CO June August MT May July ND July August WY June August Thickspike wheatgrass plants mature about 3 weeks earlier than those of western wheatgrass (Pascopyrum smithii). The summer regrowth of plants is fair and the fall regrowth is good [40]. Listed below are the average dates that plants reached various developmental phases on the Snake River Plain, Idaho [9]: Growth starts March 1 Flower stalks appear May 21 Heads fully out June 16 Flowers in bloom June 27 Plant drying July 12 Seeds ripen July 27 Dissemination starts August 8 Dissemination over August 20 Plant dried August 11

FIRE ECOLOGY

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
FIRE ECOLOGY OR ADAPTATIONS : Thickspike wheatgrass is a fire-tolerant species. Plants remain green throughout much of the summer. Since they are low growing, they are less of a fire hazard than taller species. After planned sagebrush-grassland fires, plants soon spread and dominate the ground cover [40]. Fire probably stimulates these rhizomatous plants to initiate new shoots at primordial regions of the root system [20]. POSTFIRE REGENERATION STRATEGY : Rhizomatous herb, rhizome in soil

FIRE EFFECTS

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
IMMEDIATE FIRE EFFECT ON PLANT : Fires usually consume dry vegetation to ground level. Burning thickspike wheatgrass, with its coarse stems and lesser amounts of leafy material, usually results in rapid combustion and little downward transfer of heat to belowground plant parts [44]. The rhizomatous growth form of this species minimizes the effect of fire on it. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : Burns conducted in the spring after new growth is initiated can severely injure this species [44]. PLANT RESPONSE TO FIRE : Growth habit and season of burn are the principle variables regulating the response of grasses to fire [38]. Rhizomatous species are frequently favored by fire, as fire probably stimulates the initiation of new shoots at primordial regions of the root system. Thickspike wheatgrass increases in abundance following fire [42]. Production on burned plots remains above that on controls for about 30 years [21]. Postburn recovery time is rapid (2 to 5 years) in the sagebrush and pinyon-juniper zones of the Intermountain region [44]. Thickspike wheatgrass recovers more rapidly on ungrazed pastures than on grazed experimental plots [10,26,33]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : On a sagebrush-grass range on the Upper Snake River Plains, Idaho, thickspike increased markedly on burned areas by the end of the first growing season after fall burns. The degree of increase rose with the intensity of the burn. This relative increase continued so that by the end of the third year, production doubled or tripled that on the unburned range. Thickspike usually recovers fully within 3 years of a fire. After 15 years relative production declined, but thickspike wheatgrass still produced significantly more on the burned range than on the unburned one [8]. On a western wheatgrass and thickspike wheatgrass range in southeastern Alberta, 1 year after an August wildfire, production of the two grasses was reduced 19 percent [43]. An effect of fire is to increase water stress on plants on burned sites. An accidental fire in Saskatchewan burned a strip of thickspike wheatgrass-junegrass prairie several hundred feet wide. Sampling was done along burned and unburned sides of the fire line. Plant and soil water stress increased near the end of the May-August period following the fire on both burned and unburned sites. As the growing season progressed, the potentials in thickspike wheatgrass became lower on burned plots. Observed reductions in productivity were probably the result of increased plant water stress [45]. In an experiment using thickspike wheatgrass leaves, water potential and osmotic potential were lower in plants from burned areas [43]. FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Elymus lanceolatus | Thickspike Wheatgrass
REFERENCES : 1. Allen, E. B.; Allen M. F. 1986. Water relations of xeric grasses in the field: interactions of mycorrhizae and competition. New Phytologist. 104: 559-571. [2875] 2. Asay, K. H. 1983. Promising new grasses for range seedings. In: Monsen, Stephen B.; Shaw, Nancy, compilers, Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings of symposia; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 110-115. [356] 3. Barkworth, Mary E.; Dewey, Douglas R. 1985. Genomically based genera in the perennial Triticeae of North America: identification and membership. American Journal of Botany. 72(5): 767-776. [393] 4. Barkworth, Mary E.; Dewey, Douglas R.; Atkins, Riley J. 1983. New generic concepts in the Triticeae of the Intermountain Region: key and comments. Great Basin Naturalist. 43(4): 561-572. [394] 5. Baum, Bernard R.; Estes, James R.; Gupta, Pushpendra K. 1987. Assessment of the genomic system of classification in the Triticeae. American Journal of Botany. 74(9): 1388-1395. [198] 6. Behan, Mark J. 1983. The suitability of commercially available grass species for revegetation of Montana ski area. Journal of Range Management. 36(5): 565-567. [425] 7. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 8. Blaisdell, James P. 1953. Ecological effects of planned burning of sagebrush-grass range on the upper Snake River Plains. Tech. Bull. 1975. Washington, DC: U.S. Department of Agriculture. 39 p. [462] 9. Blaisdell, James P. 1953. Ecological effects of planned burning of sagebrush-grass range on the upper Snake River Plains. Tech. Bull. 1975. Washington, DC: U.S. Department of Agriculture. 39 p. [462] 10. Clarke, S. E.; Tisdale, E. W.; Skoglund, N. A. 1943. The effects of climate and grazing practices on short-grass prairie vegetation in southern Alberta and southwestern Saskatchewan. Technical Bulletin No. 46. Ottawa, Canada: Canadian Dominion, Department of Agriculture. 53 p. [635] 11. Coupland, Robert T.; Johnson, R. E. 1965. Rooting characteristics of native grassland species of Saskatchewan. Journal of Ecology. 53: 475-507. [702] 12. Dewey, Douglas R. 1983. Historical and current taxonomic perspectives of Agropyron, Elymus, and related genera. Crop Science. 23: 637-642. [793] 13. Dewey, Douglas R. 1983. New nomenclatural combinations in the North American perennial Triticeae (Gramineae). Brittonia. 35(1): 30-33. [794] 14. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 15. Dubbs, A. L.; Harada, R. T.; Stroh, J. R. 1974. Evaluation of thickspike wheatgrass for dryland pasture and range. Bulletin 677. Bozeman, MT: Montana State University, Montana Agricultural Experiment Station. 13 p. [3545] 16. Eckert, Richard E., Jr.; Spencer, John S. 1986. Vegetation response on allotments grazed under rest-rotation management. Journal of Range Management. 39(2): 166-174. [843] 17. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 18. Ferguson, Robert B.; Frischknecht, Neil C. 1985. Reclamation on Utah's Emery and Alton coal fields: techniques and plant materials. Res. Pap. INT-335. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 78 p. [917] 19. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 20. Gartner, F. R.; Butterfield, R. I.; Thompson, W. W.; Roath, L. R. 1978. Prescribed burning of range ecosystems in South Dakota. In: Hyder, D. N., ed. Proceedings, 1st international rangeland congress; Denver, CO. Denver, CO: Society for Range Management: 687-690. [999] 21. Harniss, Roy O.; Murray, Robert B. 1973. 30 years of vegetal change following burning of sagebrush-grass range. Journal of Range Management. 26(5): 322-325. [1086] 22. Hitchcock, A. S. 1951. Manual of the grasses of the United States. Misc. Publ. No. 200. Washington, DC: U.S. Department of Agriculture, Agricultural Research Administration. 1051 p. [2nd edition revised by Agnes Chase in two volumes. New York: Dover Publications, Inc.]. [1165] 23. Hitchcock, C. Leo; Cronquist, Arthur; Ownbey, Marion. 1969. Vascular plants of the Pacific Northwest. Part 1: Vascular cryptograms, gymnosperms, and monocotyledons. Seattle, WA: University of Washington Press. 914 p. [1169] 24. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 25. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 26. Mueggler, Walter F.; Blaisdell, James P. 1958. Effects on associated species of burning, rotobeating, spraying, and railing sagebrush. Journal of Range Management. 11: 61-66. [1712] 27. Quinnild, Clayton L.; Cosby, Hugh E. 1958. Relicts of climax vegetation on two mesas in western North Dakota. Ecology. 39(1): 29-32. [1925] 28. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 29. Reitz, Louis P.; Morris, H. E. 1939. Important grasses and other common plants on Montana ranges: description, distribution and relative value. Bull. 375. Bozeman, MT: Montana State College, Agricultural Experiment Station. 35 p. [1954] 30. Rogler, George A. 1973. The wheatgrasses. In: Heath, M. E.; Metcalfe, D. S.; Barnes, R. E., eds. Forages: the science of grassland agriculture. 3d ed. Ames, IA: The Iowa State University Press; 1973: 221-230. [2022] 31. Ross, Robert L.; Hunter, Harold E. 1976. Climax vegetation of Montana based on soils and climate. Bozeman, MT: U.S. Department of Agriculture, Soil Conservation Service. 64 p. [2028] 32. Schwendiman, John L. 1975. Grasses and legumes--new and improved for range seedings. In: Range multiple use management. Pullman, WA: Washington State University: 159-172. [2098] 33. Smith, Michael A.; Dodd, Jerrold L.; Rodgers, J. Daniel. 1985. Prescribed burning on Wyoming rangeland. Bull. 810. Laramie, WY: University of Wyoming Agricultural Extension Service. 25 p. [2176] 34. Sours, John M. 1983. Characteristics and uses of important grasses for arid western rangelands. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings of a symposia; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 90-94. [2201] 35. Thornburg, Ashley A. 1982. Plant materials for use on surface-mined lands. SCS-TP-157. Washington, DC: U.S. Department of Agriculture, Soil Conservation Service. 88 p. [3769] 36. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. [2387] 37. U.S. Department of Agriculture, Forest Service, Rocky Mountain Region. 1983. Plant associations (habitat types) of Region 2.,3rd ed. Lakewood, CO. 224 p. [2385] 38. Volland, Leonard A.; Dell, John D. 1981. Fire effects on Pacific Northwest forest and range vegetation. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region, Range Management and Aviation and Fire Management. 23 p. [2434] 39. Wambolt, Carl. 1981. Montana range plants: Common and scientific names. Bulletin 355. Bozeman, MT: Montana State University, Cooperative Extension Service. 27 p. [2450] 40. Wasser, Clinton H. 1982. Ecology and culture of selected species useful in revegetating disturbed lands in the West. FWS/OBS-82/56. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service, Office of Biological Services, Western Energy and Land Use Team. 347 p. Available from NTIS, Springfield, VA 22161; PB-83-167023. [2458] 41. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 42. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. [2620] 43. Wright, Henry A.; Neuenschwander, Leon F.; Britton, Carlton M. 1979. The role and use of fire in sagebrush-grass and pinyon-juniper plant communities: A state-of-the-art review. Gen. Tech. Rep. INT-58. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Statio. 48 p. [2625] 44. Young, Richard P. 1983. Fire as a vegetation management tool in rangelands of the Intermountain Region. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 18-31. [2681] 45. Redmann, R. E. 1978. Plant and soil water potentials following fire in a northern mixed grassland. Journal of Range Management. 31(6): 443-445. [1946] 46. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 47. U.S. Department of Agriculture, Soil Conservation Service. 1994. Plants of the U.S.--alphabetical listing. Washington, DC: U.S. Department of Agriculture, Soil Conservation Service. 954 p. [23104] 48. U.S. Department of the Interior, National Biological Survey. [n.d.]. NP Flora [Data base]. Davis, CA: U.S. Department of the Interior, National Biological Survey. [23119]

Index

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