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Wildlife, Animals, and Plants |
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INTRODUCTORY
ABBREVIATION:KOEMAC
|
| AZ | AR | CA | CO | CT | DE |
| ID | IL | IN | IA | KS | LA |
| ME | MA | MI | MN | MO | MT |
| NE | NV | NH | NM | NY | NC |
| ND | OH | OK | OR | PA | SC |
| SD | TX | UT | VT | WA | WI |
| WY | |||||
| AB | BC | MB | NB | NF | NS |
| ON | PE | PQ | SK | ||
| MEXICO |
| CO | MT | ND | UT | WY | |
| Cattle | Good | Good | Good | Good | Good |
| Domestic Sheep | Good | Good | Good | Good | Good |
| Horses | Good | Good | Good | Good | Good |
| Pronghorn | ---- | Fair | Poor | Fair | Good |
| Elk | ---- | Fair-Good | ---- | Good | Good |
| Mule deer | ---- | Fair | Poor | Fair | Good |
| White-tailed deer | ---- | ---- | Poor | ---- | Good |
| Small mammals | ---- | ---- | Poor | Good | Fair |
| Small non-game birds | ---- | ---- | Poor | Good | Fair |
| Upland game birds | ---- | ---- | Poor | Fair | Fair |
| Waterfowl | ---- | ---- | Fair | Poor | Fair |
Prairie Junegrass is rated poor in protein value and good in energy value [37]. Nutritional value in accordance with relative phenological stage is [81]:
| Hay, dry | Aerial part fresh, immature | Aerial part fresh, mature | Aerial part fresh overripe | |
| % Dry matter | 100.0 | 100.0 | 100.0 | 100.0 |
| % Ash | 7.8 | 7.8 | 8.1 | 7.7 |
| % Crude Fiber | 34.2 | 25.8 | 38.6 | 41.0 |
| % Ether extract | 2.9 | 2.3 | 2.5 | 1.8 |
| % N-Free extract | 46.0 | 40.3 | 45.0 | 45.7 |
| % Protein (N x 6.25) | 9.1 | 23.8 | 5.8 | 3.8 |
| % digestible protein - Cattle | 4.8 | 18.1 | 2.8 | 1.1 |
| % digestible protein - Horses | 5.2 | 17.7 | 2.5 | 0.8 |
| % digestible protein - Sheep | 4.7 | 19.2 | 2.4 | 0.5 |
| Energy | ||||
| Cattle DE1 Mcal/kg | 2.11 | 2.50 | 2.31 | 2.31 |
| Sheep DE1 Mcal/kg | 1.99 | 2.59 | 2.44 | 2.47 |
| Cattle ME2 Mcal/kg | 1.73 | 2.05 | 1.89 | 1.89 |
| Sheep ME2 Mcal/kg | 1.63 | 2.12 | 2.00 | 2.03 |
| Cattle TDN3 % | 47.8 | 56.7 | 52.5 | 52.3 |
| Sheep TDN3 % | 45.2 | 58.8 | 55.3 | 56.0 |
The short stature and scattered distribution of prairie Junegrass provide minimum coverage for larger birds and mammals. Relative coverage classifications for wildlife species are [37]:
| MT | ND | UT | WY | |
Small mammals |
Poor | Poor | Good | Fair |
| Small non-game birds | Poor | Fair | Good | Fair |
| Upland game birds | Poor | Fair | Fair | Fair |
| Waterfowl | Good | Poor | Poor | Fair |
Prairie Junegrass can recolonize areas that have been subjected to severe water stress. Recolonization by prairie Junegrass provides protective cover to help subsequent post-drought, successional plant species growth [121].
Prairie Junegrass's ability to suppress invasive annuals is variable. Reestablishing prairie Junegrass populations, through greenhouse propagation and transplant, within an area overrun by annuals has been successful [72]. Borman and others [22] observed a weak ability to exclude annual weeds in southwest Oregon. When grown with the invasive Japanese brome (Bromus japonicus) under densities of 0, 50, 100, 200 and 400 m-2, emergence and seedling survival of prairie Junegrass were not affected [96]. Prairie Junegrass has the ability to revegetate areas of high soil disturbance. Sod obtained from a native rough fescue (Festuca scabrella) climax grass community (containing prairie Junegrass) in Canada was used to revegetate a steep-sloped ravine edge disturbed during road construction and community facility development [91]. Prairie Junegrass is a component in seeding mixtures designed for restoration of disturbed Fescue grassland sites within Glacier National Park [69]. A study evaluating secondary successional patterns in a big sagebrush (Artemisia tridentata) community in the Piceane Basin of northwest Colorado observed a positive response in prairie Junegrass vegetation cover after induced soil disturbance. Increased vegetation cover over a six year period occurred after three different soil disturbance regimes [14]:| a) | Vegetation mechanically removed and soil scarified to 12 inches (30 cm) |
| b) | Top soil and subsoil (C horizon) removed to 3.3 feet (1 m) mixed and replaced |
| c) | 2 layers, 3.3 feet (1 m), soil removed and replaced in reverse order (second layer on top) |
| Stratification | Days to 1st Germination | Days to peak Germination | Approx. # of seedlings / oz. of seed |
| 10 weeks (dry) | 13 | NA | low |
| 10 weeks (moist) | 7 | 19 | low |
Obtaining prairie Junegrass seed from plants within similar environments (i.e. elevation, climate), may be significant for re-establishing vigorous populations. A correlation between genotype and phenotype of individuals from different habitats has been observed [95].
Grazing on prairie Junegrass populations has both positive and negative results [15,39,80,88,102,103]. Evaluations of grazing are difficult because growing conditions may prove more significant than grazing pressure. Important grazing considerations for prairie Junegrass include site characteristics (e.g. soil, elevation) [109], environmental conditions [69,105], growing season, and frequency [69]. A study comparing rotational grazing (1½ months spring and 1½ months fall every other year) and continuous grazing saw increases under both regimes; precipitation was above the long term average [103]. A study comparing rotational grazing and season long grazing on a native northern mixed-grass prairie of North Dakota was associated with positive responses in prairie Junegrass basal cover between the 1st and 4th year. Degenerative effects were seen in the 6th year [15]. Prairie Junegrass responds well to grazing and is more palatable in spring and fall [114,117]. In general, grazing conducted August through October is sustainable, if adequate moisture is available [24]. Maintaining a 3 inch (7.6 cm) stubble is recommended for seral communities of the Great Basin, Pacific Northwest and northern Great Plains [80]. The increased presence of invasive plant species in areas with prairie Junegrass has led to studies evaluating the use of herbicidal controls. Tordon and Transline were evaluated for control of spotted knapweed (Centaurea maculosa) on native grasslands. No negative effects were documented for prairie Junegrass and increases in native grass density were observed [93]. Tebuthiuron applied to control sagebrush in Idaho had no effect on prairie Junegrass canopy cover at application rates reaching 1.1 kg ha -1 [123]. Correlations between mycorrhizal associations and prairie Junegrass stage of development have been observed. Positive associations are seen within early seedling development [51]. In later developmental stages, a negative association was documented within several grassland communities. In the absence of vesicular arbuscular mycorrhizae (VAM) in low P soils, prairie Junegrass outcompeted big bluestem (Andropogon gerardii); the opposite was true in the presence of VAM and P [52]. Suppression of mycorrhizal symbiosis increased the biomass of prairie Junegrass within a experimental tall-grass prairie consisting of big bluestem, purple prairie clover (Dalea purpurea), speedwell (Veronica fasciculata) and tall blazing star (Liatris aspera) [124].
Prairie Junegrass is a loosely-tufted, shallow-rooted, native grass of small stature. This cool-season perennial bunchgrass has long, mostly basal leaves [11,43,56,110]. The panicle is narrow and spikelike, except during spring flowering, when open [117].
Quantitative botanical characteristics are extremely variable depending upon sample location. The spikelike panicle can range from 1 to 7 inches (2.5-17.8 cm) in length [117] and is usually two flowered and compressed [54]. During flowering the spikelike branches are open [78].
The long, narrow, flat leaves range from 1.5 to 5 inches (3.8-12.7 cm) long from their basal point of attachment [117]. Prairie Junegrass's leaves are drought resistant and persist under dry conditions [33].
Hemicryptophyte
Regeneration is accomplished by seed, which ripens late summer to fall [110] and by sprouting from the residual plant [23,120]. Prairie Junegrass seeds are relatively small (4,000 gram-1 reported by Eddleman [40] and 3,125 gram-1 reported by Nernberg [83]) and of low viability [78,83,110].
Temperature, soil moisture and seasonality provide cues for germination of prairie Junegrass [40,104,112]. A comparison of germination and emergence of prairie Junegrass at 45 (7°C), 55 (13°C), 64 (18°C) and 80 (27°C) degrees Fahrenheit, showed the greatest % emergence at 64 degrees Fahrenheit (18°C) [104]. Blake [18] observed good germination within silt loam at 1/2 and 1/3 saturation. Prairie Junegrass also possesses the ability to germinate under periods of water stress. Germination responses were observed at water potential between 0 to 12 bars [40].
Prairie Junegrass is fairly cosmopolitan among prairie and grassland habitats of North America. A small percentage of prairie Junegrass cover is found within the majority of upland and high-prairie systems [38,90,122] in scattered stands [117]. Prairie Junegrass is normally found at elevations between 5,000 to 8,000 feet (1524-2438 m) [87]. Preferred sites are cool, semi-arid (xeric) [8,10,26], infertile grasslands and rock outcrops [27]. Annual precipitation requirements range from 16 to 21 inches (41-53 cm) [87] with 70% obtained during the growing season (August-September) [26].
The wide distribution of prairie Junegrass across several habitat types is consistent with its occurrence on many soil types [58,87]. A study evaluating ecological factors associated with production of grassland communities in western North Dakota found prairie Junegrass a member of several plant communities of different soils and vegetation [90]. Percent clay doesn't appear to have a great effect upon the presence of prairie Junegrass [58]. Steep slopes and well-drained soils with moderate to high water holding capacity are preferred [97]. Within sagebrush communities of Grand Teton National Park, a greater occurrence of prairie Junegrass was found on coarse textured soils [98]. An evaluation within the White Mountains of California found prairie Junegrass occurring on carbonate, noncarbonate, basalt, sandstone, and adamellite substrates, with a moderate affinity for noncarbonates [72]. A positive association with exchangeable Na was also found, along with a negative association to sandstone substrates and sites with high exchangeable K [72].Prairie Junegrass acts as a seral-recolonizer of previously water- stressed areas [122] and played a leading role in recolonizing bare soil of mid-continental grasslands after the droughts of 1933 to 1940 [121].
Prairie Junegrass is part of seral western juniper sites [35]. Prairie Junegrass also occurs in several climax communities. See the 'Habitat Types and Plant Communities' section of this species summary.
Prairie Junegrass has a wide distribution, so only a general description of seasonal development is given. Avoidance of growth during dry, summer months is ubiquitous [11]. Development is associated with rapid, early, spring growth [11,29] with maximum leaf growth taking place the first half of May [11].
| % Mature Foliage Height | |
| Mid-April | 65% |
| End-of-May | 90% |
| Beginning of Flowering | End of Flowering | |
| Colorado | April |
June |
| Idaho | May - June |
June - July |
| Montana | May |
July |
| North Dakota | June |
July |
| Saskatchewan | June |
June - July |
Prairie Junegrass is reported as showing little or no damage [129] to moderate damage [126] from fire. Perennial grasses possess growing points insulated near or below the soil surface [129]. The small stature of prairie Junegrass and coarse textured foliage aid in protection of these meristimatic tissue areas [129]. Possessing coarsely textured foliage and a small clump size also limits the potential for fire damage [129]. Coarse grasses like prairie Junegrass burn quickly, transferring little heat below the soil surface [120]. As a member of eastern Oregon grasslands, prairie Junegrass is considered a superior fire-resistant perennial bunch grass [126].
Fire survival strategy for prairie Junegrass is based upon seed germination and residual plant survival [23]. The extent of damage or benefit imposed by fire is highly variable. Response can vary according to fire severity, physiological state of plant, soil moisture, and season of burn [23,101]. Fire regimes for plant communities in which prairie Junegrass occurs are summarized below. For further information regarding fire regimes and fire ecology of communities where prairie Junegrass is found, see 'Fire Ecology and Adaptations' section of the FEIS species summary for the plant community or ecosystem.| Community or Ecosystem | Dominant Species | Fire Return Interval Range in Years (mean) |
| Pacific ponderosa pine* | Pinus ponderosa var. ponderosa | 1-47 [25] |
| interior ponderosa pine* | P. ponderosa var. scopulorum | 2-200 [25] |
| Colorado pinyon | P. edulis | 10-49 [25] |
| Mexican pinyon | P. cembroides | 20-70 [115,116 |
| Rocky Mountain Douglas-fir* | Pseudotsuga menziesii var. glauca | 25-100 [25] |
| coastal Douglas-fir* | Pseudotsuga menziesii var. menziesii | 40-240 [25,79,94] |
| quaking aspen (west of the Great Plains) | Populus tremuloides | 7-100 [48,75] |
| oak-hickory | Quercus-Carya spp. | 50-100 [1] |
| Texas savanna | Prosopis glandulosa var. glandulosa | < 10 [25] |
| California montane chaparral | Ceanothus and/or Arctostaphylos spp. | 50-100 [25] |
| basin big sagebrush | Artemisia tridentata var. tridentata | 12-43 [99] |
| mountain big sagebrush | Artemisia tridentata var. vaseyana | 5-15 [127] |
| Wyoming big sagebrush | Artemisia tridentata var. wyomingensis | 10-70 (40) [118,127] |
| mountain grasslands | Pseudoroegneria spicata | 3-40 (10) [7] |
| plains grasslands | Bouteloua gracilis and/or Buchloe dactyloides | 20-40 [25] |
| prairie | Andropogon gerardii var. gerardii | 1-6 [66] |
Tussock graminoid
Caudex, growing points in soil
Secondary colonizer - on-site seed
Prairie Junegrass is usually top-killed or killed by fire. Fast-moving, low-intensity fires will consume above ground vegetation without damaging the plant's crown [101]. In general, late-spring burns are more damaging to prairie Junegrass than early-spring, late-summer, fall, or winter burns [113].
The rootcrown of prairie Junegrass may sustain immediate damage depending upon amount of heat transferred through the soil [36]. Meristimatic tissue of most grasses is found at the ground surface in the rootcrown [101]. A study [36] in the southwest United States evaluated patterns of plant growth in relation to soil heating from wildfire in chaparral systems. Prairie Junegrass was found to have a strong negative correlation between heat load and resprouts.
Positive postfire vegetational responses are common for prairie Junegrass. Increased seedhead presence and height-of-inflorescence have been documented [41]. Annual burning of a native grassland in the aspen parkland of central Alberta caused a 40% increase in seedhead presence compared to unburned areas [5].
Prairie Junegrass's response to fire is related to season of burn, fire intensity, and postfire water availability [49]. Several studies evaluating the effect of fire on following season vigor report positive correlations [2,3,4,6,8]. Time required to acquire the approximate preburn frequency or coverage, is rapid, averaging 2 to 5 years [120].
Prairie Junegrass shows positive responses to fire. Fall and early-spring burns produce better responses than summer burns [4,62]. A summation on the effect of burn seasonality in eastern Oregon is given below [126]:
| Burn Season | Change in Basal Area | Mortality |
| Mid-May | -32% | 20% |
| Mid-June (Post-seed-out) | -18% | 0% |
| Mid-October | approx. -18% | 0% |
| Autumn 1977 | Spring 1978 | Summer 1978 | ||||
| Unburned | Burned | Unburned | Burned | Unburned | Burned | |
1.6 |
1.5 |
1.4 |
2.4 |
1.5 |
3.0 |
| Burn date | 1928 | 1929 |
1930 |
1931 |
1932 |
1933 |
Mean |
| Fall | 1,210 |
1,864 |
2,114 |
2,283 |
1,783 |
1,080 |
1,722 |
| Early-spring | 1,296 |
1,402 |
1,465 |
1,340 |
1,002 |
482 |
1,165 |
| Medium-spring | 574 |
555 |
756 |
1,031 |
767 |
523 |
701 |
| Late-spring | 1,058 |
1,052 |
1,571 |
1,818 |
1,068 |
447 |
1169 |
| Check plot | 619 |
1,084 |
1,052 |
992 |
627 |
181 |
759 |
| Fire Season | Years postfire | Unburned | Light-burn | Moderate-burn | Heavy-burn |
| August | 12 | 9.4 | 14.2 | 13.3 | 10.7 |
| September | 15 | 32.8 | 43.6 | 53.4 | 36.8 |
Burning, in general, has a positive influence on prairie Junegrass populations when conducted in early-spring and fall and followed by mean or above average annual precipitation [113]. Water availability after a burn is important for maintaining healthy postfire populations of prairie Junegrass. A fall burn and above average precipitation increased postfire prairie Junegrass density in a southwestern ponderosa pine (Pinus ponderosa) forest [86]. The opposite was seen at a fall burn in North Dakota, but burned areas possessed a lower moisture content than unburned areas [11].
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