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Introductory

SPECIES: Vulpia microstachys | Small Fescue
ABBREVIATION : VULMIC SYNONYMS : Festuca microstachys Nutt. Vulpia pacifica (Piper) Rydb. Vulpia arida (Elmer) Henrard Vulpia reflexa (Buckley) Rydb. SCS PLANT CODE : VUMI VUMIC VUMIC2 VUMIP COMMON NAMES : small fescue Nutall's fescue TAXONOMY : The fully documented scientific species name of small fescue is Vulpia microstachys (Nutt.) Benth. Many varieties are recognized [18,19]: Vulpia microstachys var. microstachys Vulpia microstachys var. pauciflora (Beal) Lonard & Gould Vulpia microstachys var. ciliata (Beal) Lonard & Gould Vulpia microstachys var. confusa (Piper) Lonard & Gould The annual fescues, described as only weakly distinguished from perennial fescues, were previously placed in the subgenus Vulpia within the genus Festuca. Most American researchers now follow European workers in giving the annual fescues generic status [4]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : D. Tirmenstein, February 1987 LAST REVISED BY AND DATE : D. Tirmenstein, January 1988 AUTHORSHIP AND CITATION : NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Vulpia microstachys | Small Fescue
GENERAL DISTRIBUTION : Small fescue occurs from southern British Columbia through Washington and parts of Montana southward to New Mexico and Baja California [4,9]. Distribution by variety is as follows [12,13]: V. m. var. ciliata - central Washington and Oregon south through California to Arizona and Mexico V. m. var. confusa - southern Washington to California V. m. var. pauciflora - British Columbia and Montana south to Arizona and Mexico V. m. var. microstachys - British Columbia, Washington, Oregon to western Nevada, southern California and Mexico ECOSYSTEMS : FRES29 Sagebrush FRES42 Annual grasslands STATES : AZ CA ID MT NV NM OR WA UT BC MEXICO ADMINISTRATIVE UNITS : CHIS DEVA JODA JOTR LAVO LAME MORA NOCA REDW SAGU SAMO WHIS ZION BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains KUCHLER PLANT ASSOCIATIONS : K038 Great Basin sagebrush K055 Sagebrush steppe K048 California steppe SAF COVER TYPES : NO-ENTRY SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : NO-ENTRY

VALUE AND USE

SPECIES: Vulpia microstachys | Small Fescue
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Most annual fescues have little forage value [10]. Small fescue is relatively unpalatable and is increases in response to grazing [5,16]. PALATABILITY : Small fescue is relatively unpalatable to most species of wildlife and livestock. NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : Small fescue provides only minimal cover for most species of wildlife. VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : Grazing results in a reduction of perennial grasses and an increase of small fescue [5,6].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Vulpia microstachys | Small Fescue
GENERAL BOTANICAL CHARACTERISTICS : Small fescue is a weedy, native, annual, with erect or geniculate culms growing from 5.9 to 13.8 inches in height (1.5 dm-3.5 dm) [4]. The often purplish-tinged spikelets or branches of inflorescences are three- to five-flowered, and glumes are glabrous [4,13]. RAUNKIAER LIFE FORM : Therophyte REGENERATION PROCESSES : Small fescue, an annual, reproduces by seed which drops to the ground at maturity. SITE CHARACTERISTICS : Small fescue occurs on dry to moderately moist, disturbed sites [4]. It codominates with annual hairgrass (Deschampsia danthonioides) and hairy brome (Bromus commutatus) on many Oregon bluegrass (Poa spp.) scabland sites [16]. Vulpia microstachys var. pauciflora grows well on sandy, disturbed sites, whereas V. m. var. microstachys occurs most commonly in loose soil on open slopes, and along ditches and roadways [13]. Vulpia microstachys var. ciliata grows on loose sandy soil of forest openings [13]. SUCCESSIONAL STATUS : Facultative Seral Species Although small fescue is present in several climax communities, it is most abundant on early seral sites. In threetip sagebrush, (Artemisia tripartita)-Idaho fescue (Festuca idahoensis) communities of Washington and Oregon, small fescue increases in response to disturbance along with cheatgrass (Bromus tectorum), and plantain (Plantago spp.), while perennial grasses decrease [6]. Following disturbance in Washington steppe communities, small fescue becomes increasingly abundant as perennial grasses decline [5]. SEASONAL DEVELOPMENT : Small fescue begins growth in the spring and matures earlier than associated perennial grasses.

FIRE ECOLOGY

SPECIES: Vulpia microstachys | Small Fescue
FIRE ECOLOGY OR ADAPTATIONS : There is little specific information on the adaptations of small fescue to fire; other annual fescues generally increase in abundance after fire [2]. Small fescue reoccupies a site through seed. In many areas, annual fescues mature early and drop their seeds before most wildfires occur [17]. Seeds are apparently undamaged by fire when buried in the soil, and late season fires probably have very little effect on small fescue. Recovery is rapid where viable annual fescue seeds remain buried in the soil, with an abundance of seedlings growing to maturity during the first year after the burn [3]. Small fescue generally increases in response to heavy grazing and other types of disturbance [5,6], and it is likely that fire creates an environment favorable to the germination and growth of this species. POSTFIRE REGENERATION STRATEGY : Ground residual colonizer (on-site, initial community)

FIRE EFFECTS

SPECIES: Vulpia microstachys | Small Fescue
IMMEDIATE FIRE EFFECT ON PLANT : Little is known about the specific effects of fire on small fescue. The dry foliage of annual grasses is typically consumed and the plant killed by fire. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Reestablishment of a site is through seed. Seeds buried in the soil can probably survive most fires. Where seeds do survive fire, recovery is rapid, with numerous seedlings germinating during the year after the burn. The relationship between fire intensity and severity, and seed surviviorship has apparently not been examined. Potential differences in plant response according to season of burn are also unknown. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Vulpia microstachys | Small Fescue
REFERENCES : 1. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 2. Cave, George H.; Patten, Duncan T. 1984. Short-term vegetation responses to fire in the upper Sonoran Desert. Journal of Range Management. 37(6): 491-496. [610] 3. Cline, J. F.; Uresk, D. W.; Rickard, W. H. 1977. Plants and soil of a sagebrush community on the Hanford Reservation. Northwest Science. 51(1): 60-70. [653] 4. Cronquist, Arthur; Holmgren, Arthur H.; Holmgren, Noel H.; [and others]. 1977. Intermountain flora: Vascular plants of the Intermountain West, U.S.A. Vol. 6. The Monocotyledons. New York: Columbia University Press. 584 p. [719] 5. Daubenmire, R. 1970. Steppe vegetation of Washington. Technical Bulletin 62. Pullman, WA: Washington State University, College of Agriculture, Washington Agricultural Experiment Station. 131 p. [733] 6. Franklin, Jerry F.; Dyrness, C. T. 1973. Natural vegetation of Oregon and Washington. Gen. Tech. Rep. PNW-8. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 417 p. [961] 7. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 8. Hironaka, M.; Fosberg, M. A.; Winward, A. H. 1983. Sagebrush-grass habitat types of southern Idaho. Bulletin Number 35. Moscow, ID: University of Idaho, Forest, Wildlife and Range Experiment Station. 44 p. [1152] 9. Hitchcock, A. S. 1951. Manual of the grasses of the United States. Misc. Publ. No. 200. Washington, DC: U.S. Department of Agriculture, Agricultural Research Administration. 1051 p. [2nd edition revised by Agnes Chase in two volumes. New York: Dover Publications, Inc.]. [1165] 10. Hitchcock, C. Leo; Cronquist, Arthur; Ownbey, Marion. 1969. Vascular plants of the Pacific Northwest. Part 1: Vascular cryptograms, gymnosperms, and monocotyledons. Seattle, WA: University of Washington Press. 914 p. [1169] 11. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 12. Lonard, Robert Irvin. 1970. A biosystematic study of the genus Vulpia (Gramineae). College Station, TX: Texas A&M University. 154 p. Dissertation. [3827] 13. Lonard, Robert I.; Gould, Frank W. 1974. The North American species of Vulpia (Gramineae). Madrono. 22(5): 217-280. [3826] 14. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 15. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 16. Volland, Leonard A. 1985. Plant associations of the central Oregon Pumice Zone. Rt-ECOL-104-1985. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 138 p. [7341] 17. Young, James A.; Evans, Raymond A.; Major, J. 1972. Alien plants in the Great Basin. Journal of Range Management. 25: 194-201. [2674] 18. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. [21992] 19. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 20. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090]

Index

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