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Introductory

SPECIES: Vulpia myuros | Foxtail Fescue
ABBREVIATION : VULMYU SYNONYMS : Festuca myuros Vulpia megalura Festuca megalura SCS PLANT CODE : VUMY VUMYH COMMON NAMES : foxtail fescue rattail fescue six weeks fescue TAXONOMY : The fully documented scientific species name is Vulpia myuros (L.) C.C. Gmelin. The annual fescues, only weakly distinguished from the perennial fescues, were previously placed in the subgenus Vulpia within the genus Festuca. According to Cronquist and others [2], most American researchers now follow European workers in giving the annual fescues generic status. Two varieties are recognized: V. m. var. myuros and V. m. var. hirsuta Hack. [13]. Foxtail fescue was formerly known as Vulpia (or Festuca) megalura. This species was thought to be a native North American species closely related to Vulpia myuros, a European introduction. The distinguishing characteristic was considered to be the ciliate lemma of V. megalura [2]. However, ciliate lemmas are also found on some European specimens, and most researchers now place Vulpia megalura within the species V. myuros [2,14]. Foxtail fescue (V. m. var. myuros) hybridizes with red fescue (Festuca rubra) in Britain [13]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : NO-ENTRY OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : D. Tirmenstein, February 1987 LAST REVISED BY AND DATE : D. Tirmenstein, January 1988 AUTHORSHIP AND CITATION : Tirmenstein, D. A. 1987. Vulpia myuros. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Vulpia myuros | Foxtail Fescue
GENERAL DISTRIBUTION : Foxtail fescue, a European native, is widely distributed throughout North America from Alaska and British Columbia eastward to Montana and southward to Baja California [2,8]. It is occasional in the central United States but is common in the eastern United States [2]. This species is common throughout southern Europe, Asia, Australia, and the Americas [13]. Vulpia myuros var. myuros is widely distributed throughout Eurasia, Africa, Australia, and North and South America [13,14]. In Canada, the United States, and Mexico, it occurs most commonly in coastal areas. Vulpia myuros var. hirsuta occurs widely throughout Europe, North America, South America, Hawaii, Japan, and Australia. It is occasional in the Rocky Mountains and the Midwest [13]. ECOSYSTEMS : FRES29 Sagebrush FRES34 Chaparral - mountain shrub FRES42 Annual grasslands STATES : AL AK AZ AR CA CT DE GA HI ID LA MA MD ME MI MT NC NJ NV OH OK OR PA RI SC TN TX UT WA WI VA WV BC YT MEXICO ADMINISTRATIVE UNITS : CALO CHCH COLO HAVO NOCA OLYM REDW RICH SAGU SAMO WHIS BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K038 Great Basin sagebrush K048 California steppe K055 Sagebrush steppe SAF COVER TYPES : NO-ENTRY SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Foxtail fescue is most typically a constituent of early seral plant communities. It occurs sporadically through a range of habitat types and may becomes abundant on some disturbed sites [7]. This species has been described as an indicator of climax on some sites, and it occurs as a dominant in the annual grasslands of California [16]. Foxtail fescue, along with smooth brome (Bromus mollis), dominate a vegetation zone surrounding vernal pools in southern California [11].

VALUE AND USE

SPECIES: Vulpia myuros | Foxtail Fescue
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Foxtail fescue is a fair to good seasonal forage for livestock because of the relatively large volume of herbage produced at some sites [18]. It may be a slightly better forage for cattle and horses than for sheep. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : Nutritional value is low after the plant matures in late spring [18]. COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : Foxtail fescue generally increases in response to moderate to heavy grazing [18]. It generally replaces perennial bunchgrasses following overuse [7]. Foxtail fescue is considered a serious weed in some areas [14].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Vulpia myuros | Foxtail Fescue
GENERAL BOTANICAL CHARACTERISTICS : Foxtail fescue is a single or loosely tufted weedy annual [2]. Culms grow from 7.9 to 23.6 inches (20-60 cm) in height and are decumbent and geniculate at the lower nodes [2,8]. Blades may be long or short and either flat or involute [13]. Sheaths are glabrous, blades are narrow, and panicles are elongated [2]. Spikelets are four- or five-flowered. The first glume is generally less than one-half as long as the second glume [10]. The inflorescence is contracted on a rather dense panicle or spicate raceme [13]. RAUNKIAER LIFE FORM : Therophyte REGENERATION PROCESSES : Foxtail fescue reproduces by seed which drops to the ground when mature. Seed production tends to be abundant, and seed is usually viable [18]. Optimum germination occurs over a wide range of temperatures. Germination was inhibited by a constant temperature of 92 degrees Fahrenhiet (30 deg C) and by wide diurnal fluctuations [21]. SITE CHARACTERISTICS : Foxtail fescue is widely distributed on dry, disturbed sites [2]. Although foxtail fescue stands may fluctuate greatly from year to year, this species tends to be fairly uniformly distributed over extensive foothill ranges of California [18]. It also occurs as a codominant with soft brome (Bromus mollis) on an outer, seasonally moist zone of some California vernal pools [11]. It has been reported at 4,300 feet (1,312 m) in Utah [4] and occurs from sea level to 4,500 feet (1,373 m) in California [18]. It is favored on thin to heavily packed acid soils and on recently burned chaparral in California [18]. In North America, V. m. var. myuros occurs most commonly in coastal areas [13]. SUCCESSIONAL STATUS : Facultative Seral Species Annual fescues such as foxtail fescue generally occur sporadically but may become abundant where disturbance occurs [7]. Foxtail fescue is most typically a constituent of early seral plant communities. It dominantes later seres of the annual grasslands of California, however [16]. SEASONAL DEVELOPMENT : Foxtail fescue, an annual, begins growth in the spring and matures earlier than associated perennial grasses. At many California sites maturation is completed from May 1 to May 15 [18].

FIRE ECOLOGY

SPECIES: Vulpia myuros | Foxtail Fescue
FIRE ECOLOGY OR ADAPTATIONS : Reestablishment of a burned site is through seed, which is produced in abundance even in relatively poor years [18]. Foxtail fescue cures early and drops seed prior to the onset of the main fire season in many areas [22]. Seed buried in the soil or litter is capable of surviving most fires. Fire generally creates conditions that are favorable for the germination and growth of foxtail fescue. POSTFIRE REGENERATION STRATEGY : Ground residual colonizer (on-site, initial community)

FIRE EFFECTS

SPECIES: Vulpia myuros | Foxtail Fescue
IMMEDIATE FIRE EFFECT ON PLANT : Little is known about the specific effects of fire on foxtail fescue. The dry foliage is typically consumed by fire; seeds usually remain undamaged in the soil or litter. Late season fires presumably have very little effect on this species. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Foxtail fescue generally remains unchanged or increases in response to fire [7]. This species typically becomes abundant on dry, disturbed sites such as those created by fire [2,7]. It is a common constituent of many recently burned chaparral communities in California [18]. Limited research indicates that season of burn may significantly influence the fire response of this species. Wildfires often occur after the seeds of annual grasses such as foxtail fescue have dropped to the ground [22]. Seed already buried in the litter or soil is usually undamaged by fire. Late season fires probably have little effect on foxtail fescue, while creating an environment favorable for seedling germination and growth. When undamaged seed is present on a site, recovery is generally rapid and presumably occurs by the following growing season. Little information exists on the specific response to various fire intensities and severities. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Foxtail fescue has replaced original bunchgrass vegetation as a result of overgrazing and fire in foothills big sagebrush (Artemisia tridentata ssp. vaseyana var. xericensis)- bluebunch wheatgrass (Pseudoroegneria spicata) habitat types in southern Idaho [7].

REFERENCES

SPECIES: Vulpia myuros | Foxtail Fescue
REFERENCES : 1. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 2. Cronquist, Arthur; Holmgren, Arthur H.; Holmgren, Noel H.; [and others]. 1977. Intermountain flora: Vascular plants of the Intermountain West, U.S.A. Vol. 6. The Monocotyledons. New York: Columbia University Press. 584 p. [719] 3. Daubenmire, R. 1970. Steppe vegetation of Washington. Technical Bulletin 62. Pullman, WA: Washington State University, College of Agriculture, Washington Agricultural Experiment Station. 131 p. [733] 4. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 5. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 6. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 7. Hironaka, M.; Fosberg, M. A.; Winward, A. H. 1983. Sagebrush-grass habitat types of southern Idaho. Bulletin Number 35. Moscow, ID: University of Idaho, Forest, Wildlife and Range Experiment Station. 44 p. [1152] 8. Hitchcock, A. S. 1951. Manual of the grasses of the United States. Misc. Publ. No. 200. Washington, DC: U.S. Department of Agriculture, Agricultural Research Administration. 1051 p. [2nd edition revised by Agnes Chase in two volumes. New York: Dover Publications, Inc.]. [1165] 9. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 10. Hitchcock, C. Leo; Cronquist, Arthur; Ownbey, Marion. 1969. Vascular plants of the Pacific Northwest. Part 1: Vascular cryptograms, gymnosperms, and monocotyledons. Seattle, WA: University of Washington Press. 914 p. [1169] 11. Kopecko, Kathleen J. P.; Lathrop, Earl W. 1975. Vegetation zonation in a vernal marsh on the Santa Rosa Plateau of Riverside County, California. Aliso. 8(3): 281-288. [1373] 12. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 13. Lonard, Robert I.; Gould, Frank W. 1974. The North American species of Vulpia (Gramineae). Madrono. 22(5): 217-280. [3826] 14. Lonard, Robert Irvin. 1970. A biosystematic study of the genus Vulpia (Gramineae). College Station, TX: Texas A&M University. 154 p. Dissertation. [3827] 15. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 16. Paysen, Timothy E.; Derby, Jeanine A.; Black, Hugh, Jr.; [and others]. 1980. A vegetation classification system applied to southern California. Gen. Tech. Rep. PSW-45. Berkeley, CA: U.S. Department of Agriculture, Forest Service, Pacific Southwest Forest and Range Experiment Station. 33 p. [1849] 17. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 18. Sampson, Arthur W.; Chase, Agnes; Hedrick, Donald W. 1951. California grasslands and range forage grasses. Bull. 724. Berkeley, CA: University of California College of Agriculture, California Agricultural Experiment Station. 125 p. [2052] 19. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 20. Volland, Leonard A. 1985. Plant associations of the central Oregon Pumice Zone. Rt-ECOL-104-1985. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 138 p. [7341] 21. Young, James A.; Evans, Raymond A.; Kay, Burgess L. 1973. Temperature requirements for seed germination in an annual-type rangeland community. Agronomy Journal. 65: 656-659. [12717] 22. Young, James A.; Evans, Raymond A.; Major, J. 1972. Alien plants in the Great Basin. Journal of Range Management. 25: 194-201. [2674]

Index

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