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Introductory

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
ABBREVIATION : CHANOO SYNONYMS : Cupressus nookatensis D.Don. SCS PLANT CODE : CHNO COMMON NAMES : Alaska-cedar Alaska cedar Alaska yellow-cedar Alaska yellowcedar yellow-cedar Alaska cypress Nootka cypress Nootka false-cypress Sitka cypress yellow cypress mountain cypress cypress TAXONOMY : The currently accepted scientific name of Alaska-cedar is Chamaecyparis nootkatensis (D. Don) Spach. It is a member of the Cypress family (Cupressaceae) [32]. Alaska-cedar hybridizes with members of the genus Cupressus. The hybrids are as follows [23,24]: Cupressocyparis X notabilis (Cupressus glabra x Chamaecyparis nootkatensis) Cupressocyparis X ovensii (Cupressus lusitanica x Chamaecypais nootkatensis) Cupressocyparis X leylandii (Cupressus macrocarpa x Chamaecyparis nootkatensis) The hybrids have been extensively introduced in Great Britain [23]. LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Randy Scott Griffith, July 1992. LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Griffith, Randy Scott. 1992. Chamaecyparis nootkatensis. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
GENERAL DISTRIBUTION : Alaska-cedar is found in the Pacific Coast mountain ranges from south-central Alaska to southwestern Oregon with isolated groves in the Siskiyou Moutains of northern California [1,23,24]. The eastern edge of Alaska-cedar's range is defined by two disjunct populations: one in the Selkirk Mountains of southeastern British Columbia [33] and one in the Aldrich Mountains of central Oregon [1]. ECOSYSTEMS : FRES20 Douglas-fir FRES22 Western white pine FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce STATES : AK CA OR WA BC ADMINISTRATIVE UNITS : GLBA LACL MORA NOCA OLYM REDW SAJH WRST BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 4 Sierra Mountains KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K002 Cedar - hemlock - Douglas-fir forest K003 Silver fir - Douglas-fir forest K004 Fir - hemlock forest K012 Douglas-fir forest K015 Western spruce - fir forest SAF COVER TYPES : 205 Mountain hemlock 215 Western white pine 223 Sitka spruce 224 Western hemlock 225 Western hemlock - Sitka spruce 226 Coastal true fir - hemlock 227 Western redcedar - western hemlock 228 Western redcedar 229 Pacific Douglas-fir SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Alaska-cedar is listed as a dominant or codominant overstory species in the following publications: A preliminary classification system for vegetation of Alaska [55]. The forest communities of Mount Rainer National Park [17]. A preliminary classification of forest communities in the central portion of the western Cascades in Oregon [9]. Preliminary plant associationa of the southern Cascade Mountain Province [2]. Preliminary plant associations of the Siskiyou Mountain Province [3]. Vegetation and the environment in old growth forests of northern southeast Alaska: A plant association classification [44].

VALUE AND USE

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
WOOD PRODUCTS VALUE : Alaska-cedar commands a high price for stumpage due to its fine texture, straight grain, durability, freedom from splitting and checking, resistance to acid, and excellent milling qualities [1,24,33,35]. The wood is used in window frames, doors, boat building, utility poles, marine pilings, cabinets [24,56], carving, and greenhouse construction [33]. Most of the harvested wood is exported to Japan where, because of its similar bright yellow color, it is used as a substitute for the rare hinoki (Chamaecyparis obtusa) [6]. The wood has an unusual and distinct "potato-like" odor [48]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Alaska-cedar is of minor importance to livestock and wildlife as browse. When densities of black-tailed deer are high, Alaska-cedar is browsed [51]. The Alaskan brown bear girdles the upslope side of the tree in the spring to feed on the phloem, which is high in sucrose [27]. PALATABILITY : Alaska-cedar browse is unpalatable to blue grouse [36]. NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : Alaska-cedar as a component of old-growth forests can provide critical thermal and hiding cover for large ungulates [22] and small mammals [58]. VALUE FOR REHABILITATION OF DISTURBED SITES : Alaska-cedar seedlings can be planted in the subalpine environment where disturbance is recurrent, for it is the only conifer capable of surviving on sites with frequent avalanches [15]. OTHER USES AND VALUES : Native Americans used Alaska-cedar wood to produce bows [52], masks, bowls, and dishes. The roots were split and used for the framework of baskets and hats [48]. Alaska-cedar is grown as an ornamental in North America and Europe [41]. MANAGEMENT CONSIDERATIONS : In southeast Alaska, Alaska-cedar is suffering from dieback that started around the turn of the century [28,30,31]. Most of the mortality has occurred in bog and semibog sites [28]. The search for a pathogen has been exhaustive with little results. It now seems likely the cause is abiotic [28,30,31]. The most plausible hypothesis offered thus far is that of a warming trend that started in Alaska in the late 1800's which has decreased the snow pack [28]. Because Alaska-cedar has low frost resistance [40], the decreased snow pack renders the fine roots susceptible to frost damage. This is the first sign of Alaska-cedar decline [28]. Alaska-cedar is relatively free of damaging agents due to chemical compostion of the wood [24]. It is virtually rot-free, and the snags can persist for 100+ years [29]. Hennon [26] lists the 77 known fungi associated with Alaska-cedar. Clearcutting changes the species compostion of second-growth forests in the Western Hemlock Zone, increasing Alaska-cedar's percent composition [23]. The recommended silvicultural practice of cutting old-growth Alaska-cedar is clearcut with planting [60]. Plantation-grown Alaska-cedar has a growth rate comparable to that of Douglas-fir; this is much greater than natural regeneraton of Alaska-cedar within its range [34]. Equations have been developed for Alaska-cedar based on growth percent as an estimation of future productivity on different soil types [54]. Hamilton [21] explored the response of Alaska-cedar to single-tree selection method, and he determined that Alaska-cedar will respond favorably to the method.

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
GENERAL BOTANICAL CHARACTERISTICS : Alaska-cedar is a native, evergreen, long-lived (as long as 3,500 years [16]), monecious tree [1,24]. It is slow growing with a narrow crown; the twigs are four-angled [56]. The boles of mature trees have buttressed and fluted bases, and the bark is shreddy [33]. Alaska-cedar is a medium-sized tree, although at treeline it is reduced to a shrub. It can obtain heights of 100 to 125 feet (30-38 m) with a d.b.h. as great as 12 feet (3.7 m) [24]. The root system is shallow with complex layering [24]. The leaves are scalelike and roughly 0.125 inch (0.32 cm) in length [1,46]. The stroboli are borne on the tips of branchlets. The male strobili are yellow. The female strobili are green, spherical, and 0.5 inch (1 cm) in diameter [23]. RAUNKIAER LIFE FORM : Ligneous Chamaephyte Phanerophyte REGENERATION PROCESSES : Sexual: The frequency of good seed crops is irregular (4 or more years) [59], and germination rates are low [35]. A germination rate of around 12 percent can be obtained with a warm stratification (30 days at 68 to 86 degrees Fahrenheit [20-30 deg C]) followed by a moist stratification (30 days at 40 degrees Fahrenheit [4 deg C]). A tetrazolium stain has been recommended for a test of seed viability [23]. The seed are quite small with an average of 108,000 seeds per pound (240,000 seeds/kg) [23,24]. The seed can be stored dry at 32 degrees Fahrenheit (0 deg C) for 3 to 5 years [59]. Bower and others [5] recommend foliar application of gibberellin A3 to increase flowering and filled seed. From the parent tree the mean dissemination distance is about 400 feet (120 m) [24]. Germination is epigeal [24], and mineral soil or well decomposed organic matter are the preferred germination substrates [37]. Vegetative: Alaska-cedar reproduces asexually by layering. It layers readily under the deep, heavy coastal snowpacks [49]. Vegetative reproduction is the method of choice to meet the demands for containerized stock, due to the low germination rate and infrequent good seed crops [35]. Cuttings, treated with indolebutyric acid and potted in the greenhouse, were ready for planting in 1 year [24]. Clones have advantages over seedlings such as fewer multiple leaders and uniformity in size [35]. Karlsson [34] and Karlsson and Russell [35] provide in-depth information on age of the donor, clone survival, establishment, and planting guidelines. Preliminary results indicate that there is genetic variation between provenances for shoot growth; however, further testing is needed to establish transfer zones [6]. SITE CHARACTERISTICS : Alaska-cedar occurs in hypermaritime to submaritime, subalpine, boreal, and summer-wet, cool mesothermal climates [39]. It occurs from shoreline to treeline in the northern portion of its range but is restricted to higher elevations in the southern portion [24]. Elevation: Elevational ranges for Alaska-cedar in several western states are as follows [24,49]: Feet Meters Alaska 0 to 3,000 0 - 910 Washinton and Oregon 2,000 to 7,500 600 - 2300 California 4,950 to 7,260 1,500 - 2,200 Soil: Alaska-cedar has a strong affinity for deep, well-drained soils rich in calcium and magnesium, and derived from parent materials of andesite, diorite, gabbro, or basalt (Histosol and Spodosol soil orders) [24]. It also can be found on the poor, rocky soils of the alpine environment far above the limits of other conifers [1]. Associates: In addition to those previously listed under Distribution and Occurrence, Alaska-cedar's overstory associates include California red fir (Abies magnifica), subalpine fir (A. lasiocarpa), Pacific silver fir (A. anabilis), noble fir (A. procera), Brewer spruce (Picea breweriana), whitebark pine (Pinus albicaulis), shore pine (P. contorta), incense-cedar (Libocedrus decurrens), and Pacific yew (Taxus brevifolia) [24]. Understory associates include big huckleberry (Vaccinium membranaceum), Alaska blueberry (V. alaskaense), fool's huckleberry (Menziesia ferruginea), and copperbush (Cladathamnus pyroliflorus) [24]. SUCCESSIONAL STATUS : Oblgate Initial Community Species Facultative Seral Species Obligate Climax Species Depending on the site, Alaska-cedar can be a long-lived seral species or a climax species [14,16]. In the subalpine environment it is the first tree species to become established, later forming large krummholz stands from layering [15]. Alaska-cedar is classified as shade tolerant; it will respond to 10 percent of full light and reach photosynthetic saturation at 60 percent [20]. SEASONAL DEVELOPMENT : Flowering of Alaska-cedar occurs progressively earlier in the spring as elevation decreases, suggesting that bud developement is based on heat sums [5]. Alaska-cedar flowers from April to June depending on latitude and elevation [24]. The cones of trees in the southern portion of its range mature from September to October, and dispersal begins in October and lasts through spring. In the northern portion of its range and in alpine environments, maturation of the cones is also based on heat sums, with 2- and 3-year reproductive cycles, respectively, being the norm [10]. In the northern portion of its range pollination of cones initiated the previous summer occurs from mid-April to late May; cones mature the following year [24]. The mature cones can be identified by their yellow-brown color [23].

FIRE ECOLOGY

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
FIRE ECOLOGY OR ADAPTATIONS : Fire is not an important factor in Alaska-cedar's cool, wet habitats. Alaska-cedar's bark is thin and offers little protection from fire [1]. The fire regime of Alaska-cedar's habitats is one of long-interval (150 to 350+ years) severe crown or suface fires resulting in stand replacement [44]. POSTFIRE REGENERATION STRATEGY : Tree without adventitious-bud root crown Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
IMMEDIATE FIRE EFFECT ON PLANT : The immediate effect of a cool to hot fire on Alaska-cedar is damage to the cambium layer, usually resulting in the death of the tree [1]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : Fire resistance is rated as low for Alaska-cedar [49], although a few individuals will survive a cool fire [7,25]. PLANT RESPONSE TO FIRE : Alaska-cedar will invade a burned site via wind-dispersed seed from adjacent unburned forests [24]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : The burning of slash is controversial. Fyles and others [18] recommend the burning of slash to improve access for planters, increase plantable sites, reduce brush competition, and reduce fire hazard; however, there is little information about the effects of slash burning on Alaska-cedar [12,38]. Feller [13] gives information on the effects of slashburning on nutrient loss (see Fire Case Study). After fire in the subalpine environment Alaska-cedar is slow to regenerate in the krummholz zone [8].

FIRE CASE STUDIES

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
CASE NAME : Fuel properties and slash-burning-induced nutrient losses REFERENCE : Feller, M. C. 1988 [13] SEASON/SEVERITY CLASSIFICATION : The plots were burned on 10 different days in July, August, and September 1984 to incorporate a range of fuel moisture conditions. STUDY LOCATION : The prescribed fire took place on a clearcut area of the University of British Columbia's Research forest which is located appoximately 24 miles (40 km) east of Vancouver, British Columbia. The cordinates are 49 degrees 17 minutes N latitude, 122 degrees 35 minutes W longitude. PREFIRE VEGETATIVE COMMUNITY : The preburn community was a productive coastal western hemlock forest composed of western hemlock (Tsuga heterophylla), western redcedar (Thuja plicata), Alaska-cedar (Chamaecyparis nootkatensis), and Douglas-fir (Pseudotsuga menziesii) that had been clearcut. TARGET SPECIES PHENOLOGICAL STATE : NO-ENTRY SITE DESCRIPTION : The study site was on a gentle, easterly slope at an elevation of 500 m (1,650 ft). The climate of the area is marine, warm-temperate, rainy. The mean annual precipitation is from 87 to 138 inches (220-350 cm), which is received mainly in the form of rain. The soil over most of the area was a Typic Haplorthod with a mor forest floor with a mean depth of 10 inches (26 cm). The area had been logged during a snow-free period using a high lead harvesting system. After clearcutting the slash was sorted by species* into five diameter classes: (1) < 1 cm (2) 1.1-3.0 cm (3) 3.1-5.0 cm (4) 5.1-7.0 cm (5) > 7 cm *Alaska-cedar and western redcedar were combined and shall be henceforth referred to as cedar. The area was then divided into 50 2.25-square-meter plots that were greater than 0.5 meter apart. These were slpit into 10 groups of 5 plots; within each group the plots were randomly assigned a species. Western hemlock slash had three fuel loadings (4.4, 9.9, 17.7 kg/m2) while cedar and Douglas-fir had one fuel loading (9.9 kg/m2). Each plot received all size classes of slash. Ten samples of slash were oven dried and used to determine prefire chemical and percent composition. FIRE DESCRIPTION : The fire was ignited on each plot using a strip of gasoline around the perimeter of the plot. The five plots within a group were ignited separatly, but within minutes of each other. Atmospheric conditions and fuel moisture at the time of ignition of each fire were as follows: Temp. Relative Wind speed Fine fuel C Humidity (km/hr) moisture (%) July 17 23.8 68 7 15 23 27.7 37 5 14 26 21.4 59 6 20 30 25.1 59 7 15 August 2 20.5 64 7 18 13 14.8 82 4 19 16 25.1 52 3 13 22 22.4 58 7 15 Sept. 14 22.2 49 5 32 26 15.3 64 4 20 Depth of burn into the forest floor (L, F, H layers combined) averaged 1.9 cm on the cedar plots, 1.6 cm on Douglas fir plots, and 1.8 on the western hemlock. The total mean slash consumption per species in kilograms per square meter was 4.2, 3.3, and 3.6 for cedar, Douglas-fir, and western hemlock, respectively. FIRE EFFECTS ON TARGET SPECIES : Cedar slash had the greatest depth of burn, which in turn ment greater losses of nutrients to the atmosphere. The mean nutrient loss (grams per square meter) for seven elements from the three types of slash were as follows: N P S K Na Mg Ca Cedar 26.3 1.3 2.7 3.7 0.1 2.0 19.5 Douglas-fir 19.5 1.2 2.2 2.3 0.1 1.1 11.0 Western hemlock 20.9 0.2 2.3 1.0 0.1 0.9 3.8 FIRE MANAGEMENT IMPLICATIONS : This study revealed that western hemlock/western redcedar/Alaska-cedar forests produce greater nutrient losses to the atmosphere when the slash composition has a greater proportion of Alaska-cedar and western redcedar. One can expect smaller nutrient losses when western hemlock makes up the majority of the slash. Nutrient losses can be limited if the the forest floor and larger fuels are moist when burned. This limits fuel consumption. Also nutrient loss can be reduced by more complete utilization during harvest, thus reducing the slash load.

REFERENCES

SPECIES: Chamaecyparis nootkatensis | Alaska-Cedar
REFERENCES : 1. Arno, Stephen F.; Hammerly, Ramona P. 1977. Northwest trees. Seattle, WA: The Mountaineers. 222 p. [4208] 2. Atzet, Thomas; McCrimmon, Lisa A. 1990. Preliminary plant associations of the southern Oregon Cascade Mountain Province. Grants Pass, OR: U.S. Department of Agriculture, Forest Service, Siskiyou National Forest. 330 p. [12977] 3. Atzet, Thomas; Wheeler, David L. 1984. Preliminary plant associations of the Siskiyou Mountain Province. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 278 p. [9351] 4. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 5. Bower, R. C.; Ross, S. D.; Dunsworth, B. G. 1989. Effect of GA3 treatment timing in relation to natural day length on flowering and sex expression in Chamaecyparis nootkatensis. Canadian Journal of Forest Research. 19: 1422-1428. [10044] 6. Cherry, M. L.; Lester, D. T. 1992. Genetic variation in Chamaecyparis nootkatensis from coastal British Columbia. Western Journal of Applied Forestry. 7(1): 25-29. [18313] 7. Dale, Virginia H.; Hemstrom, Miles A.; Franklin, Jerry F. 1984. The effect of disturbance frequency on forest succession in the Pacific Northwest. In: New forests for a changing world: Proceedings of the 1983 convention of The Society of American Foresters; 1983 October 16-20; Portland, OR. Bethesda, MD: Society of American Foresters: 300-304. [4781] 8. Douglas, George W.; Ballard, T. M. 1971. Effects of fire on alpine plant communities in the North Cascades, Washington. Ecology. 52(6): 1058-1064. [6738] 9. Dyrness, C. T.; Franklin, J. F.; Moir, W. H. 1974. A preliminary classification of forest communities in the central portion of the western Cascades in Oregon. Bulletin No. 4. Seattle, WA: University of Washington, Ecosystem Analysis Studies, Coniferous Forest Biome. 123 p. [8480] 10. El-Kassaby, Y. A.; Maze, J.; MacLeod, D. A.; [and others]. 1991. Reproductive-cycle plasticity in yellow-cedar (Chamaecyparis nootkatensis). Canadian Journal of Forest Research. 21: 1360-1364. [16222] 11. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 12. Feller, M. C. 1982. The ecological effects of slashburning with particular reference to British Columbia: a literature review. Victoria, BC: Ministry of Forests. 60 p. [10470] 13. Feller, M. C. 1988. Relationships between fuel properties and slashburning induced nutrient losses. Forest Science. 34(4): 998-1015. [3752] 14. Franklin, Jerry Forest. 1966. Vegetation and soils in the subalpine forests of the southern Washington Cascade Range. Pullman, WA: Washington State University. 132 p. Thesis. [10392] 15. Franklin, Jerry F.; Dyrness, C. T. 1973. Natural vegetation of Oregon and Washington. Gen. Tech. Rep. PNW-8. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 417 p. [961] 16. Franklin, Jerry F.; Hemstrom, Miles A. 1981. Aspects of succession in the coniferous forests of the Pacific Northwest. In: Forest succession: concepts and application. New York: Springer-Verlag: 212-229. [7931] 17. Franklin, Jerry F.; Moir, William H.; Hemstrom, Miles A.; [and others]. 1988. The forest communities of Mount Rainier National Park. Scientific Monograph Series No 19. Washington, DC: U.S. Department of the Interior, National Park Service. 194 p. [12392] 18. Fyles, J. W.; Fyles, I. H.; Beese, W. J.; Feller, M. C. 1991. Forest floor characteristics and soil nitrogen availability on slash- burned sites in coastal British Columbia. Canadian Journal of Forest Research. 21: 1516-1522. [18312] 19. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 20. Grossnickle, Steve C.; Russell, John H. 1991. Gas exchange processes of yellow-cedar (Chamaecyparis nootkatensis) in response to environmental variables. Canadian Journal of Botany. 69: 2684-2691. [18343] 21. Hamilton, Ronald C. 1991. Single-tree selection method: An uneven-aged silviculture system. In: Genetics/silviculture workshop proceedings; 1990 August 27-31; Wenatchee, WA. Washington, DC: U.S. Department of Agriculture, Forest Service, Timber Management Staff: 46-84. [16562] 22. Hanley, Thomas A.; Robbins, Charles T.; Spalinger, Donald E. 1989. Forest habitats and the nutritional ecology of Sitka black-tailed deer: a research synthesis with implications for forest management. Gen. Tech. Rep. PNW-GTR-230. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 52 p. [7509] 23. Harris, A. S. 1974. Chamaecyparis Spach white-cedar. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 316-320. [7586] 24. Harris, A. S. 1990. Chamaecyparis nootkatensis (D. Don) Spach Alaska-cedar. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 97-102. [13373] 25. Hemstrom, Miles A.; Franklin, Jerry F. 1982. Fire and other disturbances of the forests in Mount Rainier National Park. Quaternary Research. 18: 32-51. [6747] 26. Hennon, P. E. 1990. Fungi on Chamaecyparis nootkatensis. Mycologia. 82(1): 59-66. [13291] 27. Hennon, P. E.; Hansen, E. M.; Shaw, C. G., III. 1990. Causes of basal scars on Chamaecyparis nootkatensis in southeast Alaska. Northwest Science. 64(1): 45-54. [11028] 28. Hennon, P. E.; Hansen, E. M.; Shaw, C. G., III. 1990. Dynamics of decline and mortality of Chamaecyparis nootkatensis in southeast Alaska. Canadian Journal of Botany. 68: 651-662. [10727] 29. Hennon, P. E.; Loopstra, E. M. 1991. Persistence of western hemlock and western red cedar trees 38 years after girdling at Cat Island in southeast Alaska. Research Note PNW-RN-507. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 5 p. [18341] 30. Hennon, P. E.; Shaw, C. G., III; Hansen, E. M. 1990. Dating decline and mortality of Chamaecyparis nootkatensis in southeast Alaska. Forest Science. 36(3): 502-515. [13011] 31. Hennon, P. E.; Shaw, C. G., III; Hansen, E. M. 1990. 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Dennis; Bendell, James F. 1982. Foods selected by blue grouse (Dendragapus obscurus fuliginosus). Canadian Journal of Zoology. 60(12): 3268-3281. [10169] 37. Klinka, K.; Feller, M. C.; Green, R. N.; [and others]. 1990. Ecological principles: applications. In: Lavender, D. P.; Parish, R.; Johnson, C. M.; [and others], eds. Regenerating British Columbia's forests. Vancouver, BC: University of British Columbia Press: 55-72. [10710] 38. Klinka, K.; Green, R. N.; Courtin, P. J.; Nuszdorfer, F. C. 1984. Site diagnosis, tree species selection, and slashburning guidelines for the Vancouver Forest Region, British Columbia. Land Management Report No. 25. Victoria, BC: Ministry of Forests, Information Services Branch. 180 p. [15448] 39. Klinka, K.; Krajina, V. J.; Ceska, A.; Scagel, A. M. 1989. Indicator plants of coastal British Columbia. Vancouver, BC: University of British Columbia Press. 288 p. [10703] 40. Krajina, V. J.; Klinka, K.; Worrall, J. 1982. Distribution and ecological characteristics of trees and shrubs of British Columbia. Vancouver, BC: University of British Columbia, Department of Botany and Faculty of Forestry. 131 p. [6728] 41. Kruckeberg, A. R. 1982. Gardening with native plants of the Pacific Northwest. Seattle: University of Washington Press. 252 p. [9980] 42. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 43. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 44. Martin, Jon Randall. 1989. 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Related categories for Species: Chamaecyparis nootkatensis | Alaska-Cedar

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