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Introductory

SPECIES: Fraxinus pennsylvanica | Green Ash
ABBREVIATION : FRAPEN SYNONYMS : Fraxinus pennsylvanica var. lanceolata Fraxinus lancolata SCS PLANT CODE : NO-ENTRY COMMON NAMES : green ash red ash Darlington ash white ash swamp ash water ash TAXONOMY : The fully documented scientific name of green ash is Fraxinus pennsylvanica Marshall. The taxonomy presented here follows that of the Great Plains Flora Association [26] which does not recognize any varieties or forms within North America. Review of the literature indicates some disagreement as to the existence of the variety F. pennsylvanica var. subintegerrima (Vohl) Fern. (F. p. var. lanceolata (Borkh.) Sarg.), a glabrous form of the species more commonly encountered in the northern Great Plains and western plantings. Fully documented names of varieties and synonyms of green ash encountered in the literature are presented below [15,26,30,44,53,56]. Fraxinus pennsylvanica var. campestris (Britt.) F.C. Gates-Gates Fraxinus Pennsylvanica var. lanceolata (Borkh.) Sarg.-Budd & Best Fraxinus pennsylvanica var. austinii Fern.-Boivin Fraxinus pennsylvanica var. subintegerrima (Vahl) Fern.-Fernald Fraxinus campestris Britt. Fraxinus lanceolata Borkh.-Rydberg Fraxinus viridis Mixhx.f. LIFE FORM : Tree FEDERAL LEGAL STATUS : NO-ENTRY OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : L. C. Rosario, August 1988 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : NO-ENTRY

DISTRIBUTION AND OCCURRENCE

SPECIES: Fraxinus pennsylvanica | Green Ash
GENERAL DISTRIBUTION : Green ash is the most widely distributed of all the American ashes. Its range extends from Cape Breton Island and Nova Scotia to southeastern Alberta and Montana, and southward to central Texas and northern Florida [52,57]. The disputed glabrous variety, Fraxinus pennsylvanica var. subintegerrima (lanceolata) is found mainly in the northern Great Plains region [30,53]. ECOSYSTEMS : FRES14 Oak - pine FRES15 Oak - hickory FRES16 Oak - gum - cypress FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES19 Aspen - birch FRES28 Western hardwoods STATES : MT ND SD WY NB KS OK TX MN IA MO AR LA WI IL KY TN MS AL GA FL SC NC VA WV OH MI MD DE NJ NY PA CT RI MA VT NH MA NB NS PQ ON MB SK AB ADMINISTRATIVE UNITS : ACAD APIS BADL BISO BITH BLRI BUFF COLO COSW CUVA DEWA DETO EFMO FODO GATE GWCA GWMP GRKO GRSM HOBE INDU JELA MACA MORR NATR NERI OZAR PIRO PIPE ROCR SARA SCBL SHEN SHIL SLBE THRO TICA VAFO VOYA WICR WICA BLM PHYSIOGRAPHIC REGIONS : 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K28 Alder - ash forest K81 Oak savanna K89 Blackbelt K98 Northern floodplain forest K99 Maple - basswood forest K100 Oak - hickory forest K101 Elm - ash forest K102 Beech - maple forest K103 Mixed mesophytic forest K106 Northern hardwoods K106 Northern hardwoods; seral stages K111 Oak - hickory - pine forest K112 Southern mixed forest K113 Southern floodplain forest SAF COVER TYPES : 16 Aspen 26 Sugar maple - basswood 42 Bur oak 52 White oak - black oak - northern red oak 62 Silver maple - American elm 63 Cottonwood 65 Pin oak - sweetgum 82 Loblolly pine - hardwood 87 Sweetgum yellow poplar 88 Willow oak - water oak - diamondleaf (laurel) oak 89 Live oak 91 Swamp chestnut oak - cherrybark oak 92 Sweetgum - willow oak 93 Sugarberry - American elm - green ash 94 Sycamore - sweetgum - American elm 95 Black willow 96 Overcup oak - water hickory 101 Baldcypress 102 Baldcypress - tupelo 103 Water tupelo - swamp tupelo 235 Cottonwood - willow 236 Bur oak SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Green ash has been used as a climax indicator in a number of forested habitat typing systems in the central and western part of its range in portions of Montana, Wyoming, North Dakota, and South Dakota. The understory dominants used as indicator species within the green ash series are common chokecherry (Prunus virginiana) and western snowberry (Symphoricarpos occidentalis). Green ash also forms co-climax stands with American elm (Ulmus americana) with common chokecherry as the understory dominant, and is a dominant overstory component in various deciduous forest and woodland plant association types, including: Rocky Mountain juniper (Juniperus scopulorum); bur oak (Quercus macrocarpa); eastern hophornbean (Ostrya virginiana); aspen (Populus tremuloides); and plains cottonwood (Populus sargentii) in North Dakota, South Dakota, Nebraska, Montana and Wyoming. Published classification schemes listing green ash as a climax indicator are presented below. Location Authority MT Hansen, Hoffman and Steinauer 1984 [28] Johnston 1987 [32] ND Girard 1985 [23] Hansen, Hoffman and Steinauer 1984 [28] Hoffman and Alexander 1987 [29] WY Hansen, Hoffman and Steinauer 1984 [28]

VALUE AND USE

SPECIES: Fraxinus pennsylvanica | Green Ash
WOOD PRODUCTS VALUE : Green ash wood, which is heavy, hard, strong and yellowish with wide, white sapwood, has moderately high specific gravity and a low wood moisture content which make it a valued species for solid wood products as well as for pulp and paper requiring hardwood fibers [46]. Crating, boxing, handle stock and rough lumber can be obtained from merchantable-size trees [14]. Some plants exhibit poor form, such as crooked growth and very open tops [53], but generally apical dominance is strong enough so that vigorous, uninjured, open grown trees have a single, straight stem until they are 15 feet (4.6m) or more in height [57]. In Georgia, green ash, useful for pulpwood, lumber and veneer, is probably the most valuable commercial species adapted to the wet flats of Piedmont bottomlands. Because of its pioneer nature and ability to grow rapidly in pure, even-age stands it is well-suited for plantation management systems. Properly established and managed plantations and natural stands should produce high yields of fiber and quality solid-wood products [21]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Green ash is the principle component of wooded draws in the northern Great Plains region, providing critical habitat for many wildlife species and protecting domestic livestock both in summer and in winter. Domestic livestock are attracted to these green ash-dominated communities in the summer for shade, water, late season green vegetation, and to escape from insects; during the winter they use these areas as thermal cover and for protection from wind. Wooded draws provide sites that enhance accumulation of snow which is a valuable source of water for stockponds [7]. Green ash woodlands are considered to be important habitats for a number of wildlife species. They provide important year-round deer habitat for both mule and whitetail deer, contributing both browse and shelter [25]. These areas constitute up to 50% of the critical habitat for deer throughout much of the Great Plains [7]. Other mammal species commonly found on native woodlands include: squirrels, coyotes, rabbits, racoons and porcupines and several species of small mammals. Woodland vegetation is essential to the breeding, nesting and fledging of a number of avian species [23]. Green ash-dominated wooded draws provide 70% of sharptail grouse habitat, and nongame birds occupy the draws in the summer [7]. A rich diversity of invertebrate species are also present in the woodlands, and serve as an important food source for many species [23]. PALATABILITY : Generally green ash is of low palatability to all classes of livestock, however, it is often overbrowsed by these animals in small woodlots or around farmsteads. Deer will browse green ash, and tend to overbrowse when other preferred species are not available [25,53]. Dittberner and Olson [18] rate palatability of this species as poor for cattle, sheep, and horses in North Dakota. Palatability for wildlife has been rated as follows [18]: CO MT ND Mule deer ---- poor good White-tailed deer ---- poor good Antelope ---- ---- fair Upland game birds ---- poor ---- Sm nongame birds fair poor ---- Sm mammals ---- poor ---- NUTRITIONAL VALUE : The nutritional value of green ash is relatively low for livestock, with a fair energy value, and a poor protein value [18]. COVER VALUE : Green ash is most valuable to livestock and wildlife in the extensive prairie/plains regions where quality cover is often lacking. The degree to which this species provides environmental protection during one or more seasons for wildlife species is as follows [18]: CO MT ND Mule deer ---- fair good White-tailed deer ---- good good Antelope ---- poor ---- Upland game birds ---- good fair Sm nongame birds good good fair Sm mammals ---- fair ---- VALUE FOR REHABILITATION OF DISTURBED SITES : Green ash has been used to plant spoil banks resulting from strip mine operations in Pennsylvania, West Virginia, and the Central states with reported high survival and growth rates of up to one foot (30.5cm) per year [57]. In the Kansas, Missouri, Oklahoma area, green ash planted on strip mined areas gave 97% survival the first year and 96% survival the second year on shale and clay with a pH of 7.5 [14]. In this same study, green ash had overall the highest percent survival of five native species planted, or 63%. In another study by Bjugstad [7], green ash showed the highest survival of seven hardwood species reestablished on bentonite and low-salt coal spoils in northeast Wyoming, about 44% at the start of the third growing season. Survival and growth of this species is good on a wide variety of sites [14,34,51,57], but green ash does best, and is better developed on moist sites [8,14,34]. Bjugstad [8] reports that restoration of green ash/common chokecherry (Prunus virginiana) habitat types on coal mine spoils is greatly increased by trickle irrigation over a two year initiation period. On barren spoils, green ash should be planted in equal proportions with black locust (Robinia pseudoacacia) or other hardwoods [8,14]. This species is tolerant of moderately heavy competition from shrubs and herbs [14] and can be underplanted in black locust stands if the ground cover is not too dense [34]. Green ash has also been used extensively in wind break plantings, rated by Van Deusen and Cummingham [51] as the best medium- to fast-growing windbreak species from the standpoint of its survival and adaptability. Read [41] recommends green ash, with its moderate drought resistance, for windbreak plantings on these sites in the central Great Plains: 1) River and creek lowlands (deep, moist, permeable soils). 2) Medium to deep upland soils (silty or clayey loams). 3) Medium to deep upland soils (sandy loams and loamy sands). 4) Very wet, poorly drained soils (saline or alkaline uplands and lowlands). In all cases green ash is a medium sized tree with a medium crown density and root habit, and a moderate growth on all but very wet, poorly drained soils, where growth is slow. Green ash reaches a maximum height of 45 feet (14m) on the river and creek lowlands and only 20 feet (6.2m) on the very wet soils. The Natural Resources Conservation Service has selected and propogated from seed collected from a farmstead windbreak in Wibaux County, Montana, the cultivar 'Cardan'. This cultivar has been planted extensively by the SCS in farmstead and field windbreaks in North Dakota, South Dakota, and western Minnesota. "Cardan" is well suited to the northern Great Plains, giving excellent survival and growth rates on a wide range of soils and climatic conditions typical of the Plains [58]. Green ash is also rated as having moderate potential for erosion control and for long term revegetation [18]. Samaras of green ash should be harvested in the fall when the color fades from yellow to brown [11] or when the seed within is white, crisp, firm, and fully elongated [53]. It has been suggested that moisture content is a more important determiner of readiness for collection than color of the samara. Harvest is recommended when the average moisture content falls to 49% [17], or over a 3-week period when the average moisture content falls below 100% [16]. Samaras may be picked by hand or with pruners, shaken from the limbs of standing trees onto sheets spread below, or swept up from the streets in urban areas [11]. Seeds should be spread in shallow layers for complete drying; dewinging is not necessary. Seeds may be stored in sealed containers for up to seven years at 41 F (5C) with about 8% moisture without losing viability. There are an average of 17,000 seeds/lb [53]. Care should be taken to select seed or planting stock of the same geographical origin as the planting site, as a large variation in drought and cold-tolerance is encountered in this widely distributed species [37]. Green ash seeds may be sown in the fall without any stratification, especially in the northern reaches of green ash's range, by planting them as soon as collected, before October 15, and mulching overwinter with burlap or straw. Spring planted seed requires pregermination treatment. This can be a warm-cold stratification of 60 days at 32 to 41 F (0-5C) [11], or a 4-day soak in 100 ppm giberellic acid or 2 days in 1 to 10 ppm 6-benzyladenine followed by a 30-day stratification at 37 F (3C). This last treatment will save 30 to 60 days of stratification time and 21 days of germination time [48]. Tinus [48] also states that germination of dewinged green ash stored seed is faster and greater than germination of nondewinged stored seed. Another stratification technique to improve regeneration if seedlings are needed rapidly is to depericarp water soaked seed and partially remove the endosperm [36]. Germination is about 75% with stratified seed, which germinates in 20 days [53]. Stratified seeds should be sown in the spring, drilling them in rows 6 to 12 inches (14-30cm) apart, with 25 to 30 seeds per linear foot, or be broadcast to a density of 10 to 15 seedlings per square foot. Normal outplanting for seedstock is 1-0 or sometimes 2-0 [11]. OTHER USES AND VALUES : Green ash, a cultivated ornamental throughout its range, has often been planted for shade and landscape beautification in urban parks, recreation areas, and residential areas [11,51]. Its leaves turn golden yellow in the fall [53]. Soil Conservation Service field plantings have resulted in commercial production and extensive plantings of green ash in the northern Great Plains for field shelterbelts and farmstead windbreaks [35,51]. MANAGEMENT CONSIDERATIONS : Green ash is highly susceptible to the effects of rubbing and trampling by livestock [25], and unfenced windbreaks or shelterbelts may be damaged by cattle. Heavy utilization of green ash draws by livestock greatly reduces the occurence of mature green ash trees, and subsequent recruitment of green ash from the herbaceous, shrub, and sapling strata is considerably lower on heavily grazed sites when compared to the lightly grazed sites [13]. Mean height of mature green ash is greater on the ungrazed site as compared to the grazed site, and saplings decrease in density with an increase in grazing use [25]. Generally, overgrazing green ash wooded draws causes sparse, low vigor stands with lots of dead material on the ground, compacts the soil, inhibits reproduction, and damages mature trees by rubbing, trampling and browsing [43]. If a wooded draw occupies only a small proportion of a pasture it will probably be overused because cattle concentrate in search of shade, water, and green vegetation, regardless of stocking rate [43]. Continued heavy utilization of green ash draws by livestock may eventually result in communities composed of a mixture of small-statured shrubs with green ash assuming a declining role [13]. Many woody stands in the upland draws have already disappeared from the High Plains landscape, and many more are in a serious state of decline [10]. Because many wildlife species, including mammals, birds and invertebrates, are dependent upon the woody species to complete their life cycles, alteration or loss of the woodland habitats would have severe consequences on wildlife populations. In order to protect these limited and high value green ash communities in the northern Great Plains area, Girard [23] offers these potential solutions: 1) Interseeding of highly palatable grass species in unused areas of the range will result in decreased use of wooded draws. 2) Construction of shelters at strategic location results in improved cattle production and more effective grazing distribution than volunteer and/or forced use of wooded areas. 3) Use of insecticide-impregnated ear tags for control of insects prevents livestock from using the shade of wooded draws to avoid insect attacks or from rubbing against trees in order to dislodge insects. 4) Fencing of woodlands to restrict livestock use, although this is a costly undertaking, results in these benefits: provision of a continually reproducing windbreak for cattle; grazing in winter and/or early spring, or during emergencies; firewood; and browse for wildlife. However, in decadent woodlands, fencing may have to be used in combination with other rehabilitation techniques. 5) Placement of salt blocks and water away from woodlands helps to improve livestock distribution. Some potential improvement techniques which could be used either alone or in combination to regenerate declining green ash communities are: 1) Burning to promote vigorous sprouting of shrubs and trees. 2) Selective cutting of mature trees to stimulate sprouting, taking into consideration age of the trees, as the ability to sprout decreases with age and diameter. 3) Underplanting trees and shrubs in partially cut stands with protection from cattle grazing. Any of these improvement techniques could and should be integrated with the management suggestions offered above for the best response [10,23].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Fraxinus pennsylvanica | Green Ash
GENERAL BOTANICAL CHARACTERISTICS : Green ash is a native, deciduous tree with a large, straight trunk and high branches. This dioecious tree grows up to 66 feet (20m) high with diameters of 1 1/2 to 2 feet (46-61cm), although in the wooded draws of the northern Great Plains it usually reaches about only 25 to 30 feet (8-10m) in height [26,44,53,57]. The largest living green ash is in Missouri, with a height of 106 feet (32m) and 4.4 feet (1.34m) in diameter [53]. A flood tolerant tree, green ash has an extensive, moderately shallow root system, which contributes to a high degree of windfirmness [53,57]. The bark is dark grey to brown with shallow furrows, and the wood is heavy, hard, strong and yellowish with wide, white sapwood [44]. Apical dominance is strong enough so that vigorous, uninjured, open-grown trees have a single, straight stem until they are 15 ft (5m) or more tall. In slow growing, shaded specimens, reassertion of apical dominance when the terminal bud is removed is slow. Therefore, understory seedlings frequently have poor stem form [57]. Leaves are opposite and oddly-pinnate about 8 to 12 in (20-30cm) long with 5 to 9 (usually 7) oblong-lanceolate or elliptic, serrate or entire leaflets [53]. The inconspicuous, unisexual flowers are borne over the entire outer part of the live crown, usually beginning when trees are 3-4 in (8-10cm) in diameter and 20 ft (6m) high [57]. Staminate flowers are dense panicles which are green with reddish anthers; pistillate flowers are greenish yellow in short panicles [44]. The fruit is an elongated, winged, single-seeded samara borne in clusters [11], and large seed crops are produced every year. According to Wright [57], green ash is composed of three or more ecotypes. The population from the arid northwestern part of green ash's range is more drought resistant than that from the moister central Great Plains, and as compared to the Coastal Plain ecotype (Virginia, North and South Carolina), the Northern States ecotype (Maine to Minnesota) grows more slowly, has greener petioles, is more winter hardy, and the leaves are less subject to damage by fall frosts. Several insects feed on green ash trees: oyster scale; carpenter worm; two ash saw flies; and unspecified borers particularly affect shade trees and windbreak plantings. Fungus, athracnose, rusts, and root rot sometimes damage trees and wood. Rabbits, deer and cattle may damage unfenced plantings [57]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Green ash regenerates both through sexual and vegetative reproduction often regenerating profusely from either seed or vegetatively after disturbance [38]. Large seed crops are produced each year, and the winged samaras are wind-dispersed, most within a few hundred feet of the parent tree. Some dispersal by water occurs, but the importance of water as a long distance dispersal agent is not known [57]. Wind and water dispersed seeds drop during the fall and winter months and germinate the following spring on a variety of ground types including moist litter as well as mineral soil, but rarely in dense vegetation [3]. This species grows best in partial shade [38]. Plants will reproduce from wind blown seeds along river banks [53]. This tree responds quickly to damage by sprouting when the top is removed, especially when trees are in smaller diameter classes [3]. The ability to sprout decreases with age and diameter of the parent tree [25]. The plants will sprout readily from the root crown or from stumps following damage [3,25,38,53,57], and it has been suggested that success and propagation of this species in an island environment is more due to its ability to sprout and resprout than to the number of successful instances of seedling establishment [3]. Land managers can take advantage of this reproductive strategy in northern Great Plains declining green ash communities by using selective cutting to help regenerate the woodlands through resprouting [25]. Green ash of sapling or pole size sprouts readily, resulting in clumps of several stems, and cuttings made from young trees root easily under greenhouse conditions. However no practical way to root cuttings from older trees has been found. This species can also be bench or field grafted [57]. SITE CHARACTERISTICS : Green ash, the most widely distributed of all the American ashes, grows in a subhumid to humid climate with an average annual precipitation of 15 to 60 inches (38-155cm) and an average length frost free season from 120 to 280 days. This flood tolerant species is almost completely confined to bottomland sites, but grows well when planted on moist upland soils. It is most commonly found on alluvial soils along rivers and brooks and less frequently in swamps, and is common on land subject to flooding once or twice a year, remaining healthy when flooded up to 40% of the time during the growing season [57]. Tree species most commonly associated with green ash are box elder (Acer negundo), red maple (Acer rubrum), American elm (Ulmus americana), pecan (Carya illenoensis), sugarberry (Celtis laevigata), sweetgum (Liquidambar styraciflua), American sycamore (Platanus occidentalis), eastern cottonwood (Populus deltoides), plains cottonwood (P. sargentii), quaking aspen (P. tremuloides), black willow (Salix nigra), and willow oak (Quercus phellos) [57]. In the western part of its range, mainly in the northern Great Plains, where rainfall is insufficient to suppport upland tree growth, green ash is commonly found in upland coulees and draws, broad valleys, and on floodplains [9]. The tree canopy in these wooded draws is primarily green ash, associated with American elm, boxelder, and Rocky Mountain juniper (Juniperus scopulorum). Chokecherry (Prunus virginiana) dominates the sapling layer, and with snowberry (Symphoricarpos occidentalis), makes up the shrub component. Green ash occurs on a wide variety of soils although it survives best on deep, permeable, well-drained loams [52], preferring these to river sand [54]. This species has been planted on medium to coarse-textured upland sands and loams with good moisture relations, and is tolerant of moderately strong acid (pH 4.0) to moderately basic reacting soils [53]. The elevational ranges for green ash in several northern Great Plains states are as follows [18,32]: Colorado 3,500 to 5,700 feet (1,067-1,737m) Montana 3,400 to 4,500 feet (1,036-1,372m) Nebraska 2,600 to 4,500 feet (793-1,372m) North Dakota 2,240 to 3,840 feet (683-1,170m) South Dakota 3,000 to 4,200 feet (915-1,280m) Wyoming 4,100 to 4,400 feet (1,250-1,341m) SUCCESSIONAL STATUS : Green ash is rated as intolerant to moderately tolerant of shade. In all but the northwestern extension of its range (northern Great Plains) it establishes early in succession on alluvial soils either as a pioneer or following eastern cottonwood (Populus deltoides var. deltiodes), quaking aspen (P. tremuloides), or willow (Salix spp.). Green ash is less able to maintain a position in the crown canopy than its more rapidly growing associates such as red maple (Acer rubrum) and American elm (Ulmus americana); for this reason the proportion of ash usually decreases with increasing age in mixed elm-ash-maple stands [57]. However the northern Great Plains is beyond the range of red maple, and dutch elm disease has limited the expansion of American elm, therefore it apears that green ash is the climax species in these northern Great Plains green ash communities. Evidence also exists that it is replacing eastern cottonwood as the tree canopy dominant in floodplain communities where flooding no longer occurs [23]. SEASONAL DEVELOPMENT : Green ash, a dioecious tree, flowers before the leaf buds start to enlarge in March to April in Florida, and in late April to early May in Pennsylvania. Male flower buds require 1 to 2 weeks to pass from the unenlarged winter condition to completion of pollen shedding which takes about three days. Pollen is wind disseminated, most falling within 200 to 300 feet (60-90m) of the source. Female trees begin flower bud enlargement a few days later, and the stigmas are receptive as soon as they emerge from the bud for about a week. Within one month of pollination samaras developing from fertilized flowers reach mature size, although growth and ripening of the embryos is not completed until late September or early October. Seeds begin to fall as soon as they ripen until winter or early spring, and leaves fall at about the same time as when the seeds ripen [57].

FIRE ECOLOGY

SPECIES: Fraxinus pennsylvanica | Green Ash
FIRE ECOLOGY OR ADAPTATIONS : Generalized fire effects information indicates that green ash is adapted to disturbance by fire [3,24,43,53]. If the fire is hot enough to girdle even mature trees, which have little protection from burning because of their relatively thin bark, this species will sprout prolifically from the root crown when the main stem is damaged [43]. To a lesser degree postfire regeneration most likely involves the germination of on-site canopy stored seed and/or off-site wind or water dispersed seed as well [3,38,53]. POSTFIRE REGENERATION STRATEGY : Tree with adventitious-bud root crown/soboliferous species root sucker Initial-offsite colonizer (off-site, initial community)

FIRE EFFECTS

SPECIES: Fraxinus pennsylvanica | Green Ash
IMMEDIATE FIRE EFFECT ON PLANT : Limited fire effects information concerning green ash in the northern Great Plains portion of its range indicates that this species demonstrates good tolerance to fire when burned in a dormant state [25,53]. However, because of its thin bark, this species is subject to girdling by fire, depending on the intensity of the burn [20]. This species is known to sprout from the root crown following a burn if the top portion of the plant is killed [3,25]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Green ash is tolerant of burning and is stimulated to sprout from the root crown following damage from fire within the first year of the burn; however its ability to resprout decreases with age and with diameter [25]. Although it is thought that vegetative reproduction is the primary mode of regeneration after fire due to green ash's ability to sprout prolifically after damage, it is also expected that this tree, a prolific seeder [53], will regenerate from seeds. Seedling establishment may be either off-site or on-site, depending on the season and intensity of the fire: a mature tree with seeds intact could probably only survive a low intensity ground fire. Response and recovery time according to fire intensity, severity, and season of the burn has not been well documented for this species. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : In the northern Great Plains, fire has been suggested as a possible tool for treatment of degenerating woody draws [23,43] due to overuse by livestock (See Value and Use, Management Slot). Green ash communities showed an increase in biomass, density, and cover when burned by a wildfire in the Little Missouri grassland [25], and prescribed fire may be useful for opening up shrub thickets or for triggering sprouting in remnant trees and shrubs [23,43]. However, in the bottomland types of the North Central States where production of wood for sawtimber, veneer, paper and fuel is the primary management objective, it is recommended that fire not be used as a management tool, as bottomland species are susceptible to fire damage. Once larger trees are wounded, the entrance of heartwood decay producing fungi is facilitated, eventually leading to substantial cull and volume loss [38].

REFERENCES

SPECIES: Fraxinus pennsylvanica | Green Ash
REFERENCES : 1. Ahlgren, C. E. 1957. Phenological observations of nineteen native tree species in northeastern Minnesota. Ecology. 38(4): 622-628. [74] 2. Andersen, C. P.; Sucoff, E. I.; Dixon, R. K. 1987. The influence of low soil temperature on the growth of vesicular- arbuscular mycorrhizal Fraxinus pennsylvanica. Canadian Journal of Forest Research. 17(8): 951-956. [2904] 3. Barnes, W. J. 1985. Population dynamics of woody plants on a river island. Canadian Journal of Botany. 63: 647-655. [2855] 4. Barton, Lela V. 1945. Viability of seeds of Fraxinus after storage. Contributions from Boyce Thompson Institute. 13: 427-432. [199] 5. Bellah, R. Glenn; Hulbert, Lloyd C. 1974. Forest succession on the Republican River floodplain in Clay County, Kansas. Southwestern Naturalist. 19(2): 155-166. [241] 6. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 7. Bjugstad, Ardell J. 1977. Reestablishment of woody plants on mine spoils and management of mine water impoundments: an overview of Forest Service research on the n. High Plains. In: Wright, R. A., ed. The reclamation of disturbed lands. Albuquerque, NM: University of New Mexico Press: 3-12. [4238] 8. Bjugstad, Ardell J. 1986. Wooded draws of the northern high plains: characteristics, value and restoration (North and South Dakota). Restoration & Management Notes. 4(2): 74-75. [4226] 9. Bjugstad, Ardell J.; Girard, Michele. 1984. Wooded draws in rangelands of the northern Great Plains. In: Henderson, F. R., ed. Guidelines for increasing wildlife on farms and ranches: With ideas for supplemental income sources for rural families. Manhattan, KS: Kansas State University, Cooperative Extension Service; Great Plains Agricultural Council, Wildlife Resources Committee: 27B-36B. [4239] 10. Boldt, Charles E.; Uresk, Daniel W.; Severson, Kieth E. 1979. Riparian woodlands in jeopardy on Northern High Plains. In: Johnson, R. Roy; McCormick, J. Frank, technical coordinators. Strategies for protection & management of floodplain wetlands & other riparian ecosystems: Proc. of the symposium; 1978 December 11-13; Callaway Gardens, GA. Gen. Tech. Rep. WO-12. Washington, DC: U.S. Department of Agriculture, Forest Service: 184-189. [4359] 11. Bonner, F. T. 1974. Platanus L. sycamore. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United States. Agriculture Handbook No. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 641-644. [7730] 12. Brutvan, B.; Klukas, R. (revised by R. Klukas). 1982. Checklist of plants of Wind Cave National Park.. [Place of publication unknown]: [Publisher unknown]. 32 p. On file with: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratoy, Missoula, MT. [374] 13. 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Related categories for Species: Fraxinus pennsylvanica | Green Ash

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