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Wildlife, Animals, and Plants |
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INTRODUCTORY
ABBREVIATION:JUNOCC SYNONYMS:
Juniperus occidentalis ssp. australis (Vasek) A. Holmgren & N. Holmgren NRCS PLANT CODE:
JUOC COMMON NAMES:
western juniper TAXONOMY:
The scientific name of western juniper is Juniperus occidentalis Hook. (Cupressaceae) [44,45,53]. The following varieties of western juniper are recognized: LIFE FORM:Tree FEDERAL LEGAL STATUS:No special status OTHER STATUS:No entry AUTHORSHIP AND CITATION:Tirmenstein, D. (1999, April). Juniperus occidentalis. In: Remainder of Citation DISTRIBUTION AND OCCURRENCE
GENERAL DISTRIBUTION:
Western juniper occurs from southeastern Washington and Oregon southward to the upper slopes of the Sierra Nevada and San Bernardino Mountains of southern California [87]. It occurs along the western edge of the Great Basin in southwestern Idaho and northwestern Nevada [70]. ECOSYSTEMS:
FRES21 Ponderosa pine STATES:
CA ID NV OR WA BLM PHYSIOGRAPHIC REGIONS:
1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range KUCHLER PLANT ASSOCIATIONS:
K007 Red fir forest K008 Lodgepole-pine-subalpine forest K011 Western ponderosa forest K023 Juniper-pinyon woodland K024 Juniper steppe woodland K030 California oakwoods SAF COVER TYPES:
205 Mountain hemlock 207 Red fir 208 Whitebark pine 211 White fir 218 Interior Douglas-fir 237 Interior ponderosa pine 238 Western juniper 247 Jeffrey pine 250 Blue oak-gray pine 256 California mixed subalpine SRM (RANGELAND) COVER TYPES:
107 Western juniper/big sagebrush/bluebunch wheatgrass 109 Ponderosa pine shrubland 110 Ponderosa pine-grassland 201 Blue oak woodland 209 Montane shrubland 210 Bitterbrush 212 Blackbush 322 Curlleaf mountain-mahogany-bluebunch wheatgrass 412 Juniper-pinyon woodland 415 Curlleaf mountain-mahogany HABITAT TYPES AND PLANT COMMUNITIES:
Western juniper is an indicator of climax in a variety of sagebrush (Artemisia spp.) grassland, shrub-steppe, and dry coniferous habitat types. It occurs as a codominant with singleleaf pinyon (Pinus monophylla), big sagebrush (A. tridentata), gray low sagebrush (A. arbuscula ssp. arbuscula), stiff sagebrush (A. rigida), curlleaf mountain-mahogany (Cercocarpus ledifolius), antelope bitterbrush (Purshia tridentata), rubber rabbitbrush (Chrysothamnus nauseosus), green rabbitbrush (C. viscidiflorus), arrowleaf balsamroot (Balsamorhiza sagittata), Sandberg bluegrass (Poa secunda), Idaho fescue (Festuca idahoensis), bluebunch wheatgrass (Pseudoroegneria spicata), smilo grass (Piptatherum micranthum), and Thurber needlegrass (Achnatherum thurberiana). Western juniper also occurs with spiny hopsage (Grayia spinosa), gray horsebrush (Tetradymia canescens), junegrass (Koeleria macrantha), needle-and-thread grass (Hesperostipa comata), bottlebrush squirreltail (Elymus elymoides), and other grasses [27,37,87]. VALUE AND USE
WOOD PRODUCTS VALUE:
Western juniper has been used since historic times for firewood, charcoal, corrals, poles, and fence posts [22]. The wood is extremely durable and resistant to rot [77]. Juniper wood splits easily, burns clean and produces little ash [22,43]. Western juniper woodlands can produce 8 to 11 cords of firewood per acre. However, it is estimated that 7 hours of labor are required per cord to cut, limb, pile slash, and gather the wood [14]. Western juniper dulls saws since wind-blown sand particles readily adhere to its shaggy bark [43]. IMPORTANCE TO LIVESTOCK AND WILDLIFE:
Western juniper provides food and cover for a variety of bird and mammal species [94]. It is browsed to some extent by mule deer and elk [68]. In parts of California, mule deer feed on small amounts of western juniper during winter and spring [10,59]. In the high desert of Oregon, western juniper is an important winter pronghorn food [86]. Western juniper is used primarily as an emergency food source by most classes of livestock and many big game species [77]. In California, it may be an important critical deer food source during severe winters; at these times, it is consumed in large quantities [59]. PALATABILITY:
Western juniper is palatable browse for elk, mule deer, mountain cottontails, porcupines and black-tailed jackrabbits [68]. However, palatability of western juniper varies by individual tree [87] and Rosentreter and Jorgensen [82] describe overall palatability as "low." NUTRITIONAL VALUE:
Western juniper is fairly nutritious for mule deer and other large mammals, but is not highly digestible [60]. Crude protein (%) is as follows [10]: January 6.9 February 5.6 March 7.2 April 7.0 May 8.3 August 7.5 October 7.8 November 8.5 December 7.0 Nutritive values vary by season and plant part. Protein and ash content (%) is as follows [35]: crude protein ash green foliage 8.1 3.9 cured foliage 7.6 4.2 bark 3.2 7.1 COVER VALUE:
Western juniper provides perching and nesting sites for at least 27 species of birds, as well as cover and hibernation sites for small mammals [68]. In southeastern and south-central Oregon western juniper provides excellent hiding and thermal cover for mule deer [61,62]. In parts of Nevada, it also provides some cover for pronghorn [101]. Western juniper also provides shade for domestic livestock [22]. VALUE FOR REHABILITATION OF DISTURBED SITES:Western juniper can be propagated from cuttings or by layering [22,87]. Trees have been used as riprap for stabilizing streambanks [22]. OTHER USES AND VALUES:
Western juniper has been cultivated as an ornamental since 1840 [50]. The wood is used in toys, sporting goods, jewelry boxes, suitcase and closet liners, inlay products, clocks, decorative items, and pencils [43,77]. Pipe bowls are made from the roots of western juniper, and pet bedding from the shavings [43]. Juniper boughs have been used for Christmas wreaths and other decorations. Over 100 tons of boughs from central Oregon were sold in 1983 at 1 to 2 cents per pound [77]. MANAGEMENT CONSIDERATIONS:
In western juniper woodlands, wet to moderately wet soils grazed during the early spring are subject to damage through increased soil compaction. Grazing on dry or frozen soils causes fewer negative effects [13]. At least 3 years of rest from grazing is suggested after seeding in western juniper [82]. BOTANICAL AND ECOLOGICAL CHARACTERISTICS
Western juniper is a tree that typically grows 15 to 30 feet (4.6 to 9.1 m) [87]. It rarely exceeds 60 feet (18.3 m) in height. The largest recorded specimen is 86 feet tall (26 m), with a circumference of 480 inches (1,219 cm) and crown spread of 58 feet (18 m) [12]. Trees develop full crowns and heavy limbs at maturity [22].
Stand Height Diameter
inch cm inch cm
Open, dominant 3.5 9 0.3 0.8
Closed, subdominant 3.5 9 0.1 0.4
Closed, young 1.2 3 0.05 0.2
Highest rate 4.3 11 0.5 1.3
RAUNKIAER LIFE FORM:Phanerophyte REGENERATION PROCESSES:
Western juniper regenerates through seed. Green pistillate megastrobili form seed cones in abundance [87,92]. Female cones first develop 2 weeks after the male cones, and seed cones remain on the tree for 2 years until mature [72,84]. Seed cones generally ripen in mid-September. Seed cones are blue-green prior to ripening, and bluish-black and glaucous when ripe [50]. Cone-berries contain 1 to 4 (more rarely up to 12) brownish seeds [69]. Western juniper is monoecious or dioecious [92]. Many plants have the capacity to shift their sex from year to year, depending on conditions. Trees bordering roads or clearings typically produce much greater numbers of female cones than do trees within stands [28]. with big sagebrush - 47% with western juniper - 15% with bunchgrasses - 14% in the open - less than 1%Burkhardt and Tisdale [19] found that most seedlings established on the north side of existing trees where the young plants were protected from intense solar radiation. Seedlings are often found along fences, hedges, or under taller trees where large numbers of birds perch [69,87,103]. Western juniper does not reproduce vegetatively [22]. SITE CHARACTERISTICS:
Western juniper occurs in open stands on mountain slopes and high plateaus [87]. It grows as scattered individuals on rimrock or rock outcrops and in higher densities along streams, on scablands, and lower slopes where water has dispersed the seed [28]. It has remained dominant in shifting sand dune communities of south-central Washington where fire is unlikely [1]. SUCCESSIONAL STATUS:
Western juniper is a highly competitive invasive species [81]. It is found on all exposures and slopes but is relatively intolerant of shade [22,23]. This long-lived species has been expanding its range into adjacent shrub-steppes, grasslands, and savannas during the past 100 to 150 years [8]. It has doubled its range in central and eastern Oregon during the past 80 to 100 years [81]. Expansion into low sagebrush communities has been slower than into big sagebrush communities [2]. Expansion into shrub steppe communities in Nevada has reduced understory diversity and productivity [7]. Western juniper expansion has been attributed to livestock grazing and associated reductions in fine fuels, climatic changes (mild temperatures and above average precipitation in the late 1880s and early 1900s), and reduction in fire frequency due to fire suppression and cessation of Native American burning [8,29,74,75]. SEASONAL DEVELOPMENT:In Oregon, leaf elongation begins in June with 15 to 20% annual leaf turnover [72]. Western juniper cones develop from mid-April to mid-May and pollen is shed in May [50,72,87]. Seed cones ripen in mid-September after the second growing season [50]. Cones may persist on the trees for 2 to 3 years [69]. Most germination occurs during the spring [87]. In Oregon, most natural germination occurs during April [22]. FIRE ECOLOGYFIRE ECOLOGY OR ADAPTATIONS:
Young western junipers have thin bark and are readily killed by surface fires. "Older" trees with thicker bark are described as "moderately resistant" to fire [87]. In general, the taller the juniper, the greater the severity of the fire required to kill it [64]. Western juniper does not sprout [19]. Reestablishment is through seed that is dispersed fairly slowly by water and animals. POSTFIRE REGENERATION STRATEGY:
Tree without adventitious bud/root crown FIRE EFFECTS
IMMEDIATE FIRE EFFECT ON PLANT:Western juniper is a nonsprouter that is generally killed by severe fires [11]. Younger trees have thin bark and are readily killed by surface fires [87]. Older trees with thicker bark, little fuel near the stem, and higher foliage are moderately resistant to fire [38,87]. Fire may not carry in open stands of mature trees with sparse understory vegetation [15]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT:Western juniper foliage is not easily ignited and/or scorched when moist [17]. Western junipers under 4 to 6 feet (1.2-1.8m) tall are readily killed by fire [64,17]. A more severe fire is necessary to kill taller trees [64]. PLANT RESPONSE TO FIRE:Mature trees are somewhat resistant to fire if the crown is not scorched, so some larger trees may survive low-severity fires and serve as a seed source [67]. Western juniper first becomes dominant 30 to 50 years after fire [15]. Reoccupation of a site occurs fairly slowly through dispersed seed [19,25]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE:Recovery time depends on the size of burn, location of seed source, stand maturity, and presence of animal dispersers [19]. Postburn succession in western juniper communities depends on season of burn, postfire mortality, and on seed of associated species present in the preburn community. Postfire succession is also related to the effects of competition from herbaceous species and shrubs as well as drought. Large burns and long distances from seed sources slow recovery rates. According to Bunting and others [17] "because these factors vary widely early seral community composition is highly variable." Fire scars have been observed on western junipers found growing in fire-resistant low sagebrush communities lacking perennial grass cover [103]. FIRE MANAGEMENT CONSIDERATIONS:
Western juniper sites with less than 1,322 pounds per acre (600 kg/ha) of fine fuels are difficult to burn [16]. As the crown of an established western juniper expands over time, herbaceous production declines from the combined effects of shading, litter accumulation, and soil moisture [1]. Trees create their own fine fuel break, so these stands may be virtually "fireproof" except under the "most severe burning conditions" [1,103]. Many western juniper woodlands have advanced to a point where prescribed fire is no longer a viable management option. For example, in extremely dense stands, prescribed burning would be both hazardous and expensive. In some cases, weedy annuals prevent the establishment of more desirable perennial species after fire [16].
foliage litter cones woody fuel
average ash content (%) 4.26 5.31 3.42 1.35-2.80
average heat content
with ash (mJ/kg) 23.64 22.53 23.68 20.04-20.27
average heat content
without ash (mJ/kg) 24.70 23.79 24.51 20.31-20.66
Van Wagtendonk and others [97] recommend adjusting fire spread models for use with Sierra Nevada conifers. If standard fuel models are used, they suggest adjusting fire front with the following "correction factors:"heat per Unit Area fireline intensity flame length (reaction intensity) 0.98 0.95 0.98Western juniper-low sagebrush communities that have been invaded by medusahead (Taeniatherum caput-medusae) are often extremely "fire hazardous" [103]. FIRE CASE STUDIES
CASE NAME:
Basin big sagebrush-bunchgrass prescribed burn, Dayville, OR REFERENCE:Sapsis, D. S. 1990 [83] SEASON/SEVERITY CLASSIFICATION:
Fall (9/25/87) /severity not specified STUDY LOCATION:The study was located approximately 5 miles (10 km) west of Dayville in east-central Oregon. The site was located in John Day Fossil Bed National Monument in T 11 S R 26 E , sections 31 and 32. PREFIRE VEGETATIVE COMMUNITY:Preburn vegetation was a basin big sagebrush (Artemisia tridentata ssp. tridentata)-Idaho fescue (Festuca idahoensis)-bluebunch wheatgrass ((Pseudoroegneria spicata) community. Dominant understory species included Idaho fescue, bluebunch wheatgrass, yarrow (Achillea millefolium) and threadstalk milkvetch (Astragalus fillipes). Western juniper (Juniperus occidentalis) did not occur as a dominant or codominant in the prefire vegetative community. Western juniper presumably grew as scattered individuals (many of which were seedlings). Prefire cover of western juniper was 369 individuals per ha. TARGET SPECIES PHENOLOGICAL STATE:Not specified SITE DESCRIPTION:
Aspect - north FIRE DESCRIPTION:Both burns were ignited with drip torches using a strip-head firing pattern. Pretreatment fuel loads ranged from 5-12 Mg/ha. Fuel loads in fall treatment units averaged 10.5 Mg/ha and in spring treatment units, fuel loads averaged 6.2 Mg/ha. Large amounts of herbaceous fuels (> 3 Mg/ha) were present. Burning Conditions:
Fire Behavior:
*significant difference between fall and spring burning (p<0.05) FIRE EFFECTS ON TARGET SPECIES:
Both spring and fall treatments killed all western juniper:
Western juniper frequency (%):
FIRE MANAGEMENT IMPLICATIONS:Prescribed fire was effective in limiting the abundance of western juniper. Prescribed fire may represent an effective and economical management tool for rangelands. Juniperus occidentalis: References1. Agee, James K. 1994. Fire and weather disturbances in terrestrial ecosystems of the eastern Cascades. Gen. Tech. Rep. PNW-GTR-320. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 52 p. (Everett, Richard L., assessment team leader; Eastside forest ecosystem health assessment; Hessburg, Paul F., science team leader and tech. ed., Volume III: assessment). [23656] 2. Agee, James K. 1996. Fire in the Blue Mountains: a history, ecology, and research agenda. In: Jaindl, R. G.; Quigley, T. M., eds. Search for a solution: sustaining the land, people and economy of the Blue Mountains. Washington, DC: American Forests: 119-145. [28827] 3. Anderson, E. William. 1956. Some soil-plant relationships in eastern Oregon. 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