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Introductory

SPECIES: Rhododendron maximum | Rosebay
ABBREVIATION : RHOMAX SYNONYMS : NO-ENTRY SCS PLANT CODE : RHMA4 COMMON NAMES : rosebay rosebay rhododendron great-laurel great-rhododendron white-rhododendron TAXONOMY : The currently accepted scientific name for rosebay is Rhododendron maximum L. [14]. There are no recognized subspecies, varieties, or forms. LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Milo Coladonato, November 1991 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Coladoanto, Milo 1991. Rhododendron maximum. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Rhododendron maximum | Rosebay
GENERAL DISTRIBUTION : Rosebay grows mostly in the mountainous regions of eastern United States from western Maine to Georgia. Outlying populations can be found in the mountains of eastern Kentucky, eastern Tennessee, and Ohio [2,29]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES12 Longleaf - slash pine FRES13 Loblolly - shortleaf pine FRES14 Oak - pine FRES15 Oak - hickory STATES : AL CT GA KY MA MD ME NC NH NJ NY OH PA SC TN VA VT WV ADMINISTRATIVE UNITS : ALPO BISO BLRI CATO CUGA DEWA GRSM JOFL NERI OBRI SHEN BLM PHYSIOGRAPHIC REGIONS : NO-ENTRY KUCHLER PLANT ASSOCIATIONS : K095 Great Lakes pine forest K096 Northeastern spruce - fir forest K097 Southeastern spruce - fir forest K100 Oak - hickory forest K104 Appalachian oak forest K108 Northern hardwoods - spruce forest K110 Northeastern oak - pine forest K111 Oak - hickory - pine forest K112 Southern mixed forest K113 Southern floodplain forest SAF COVER TYPES : 21 Eastern white pine 22 White pine - hemlock 23 Eastern hemlock 24 Hemlock - yellow birch 40 Post oak - blackjack oak 43 Bear oak 44 Chestnut oak 45 Pitch pine 46 Eastern redcedar 51 White pine - chestnut oak 52 White oak - black oak - northern red oak 53 White oak 55 Northern red oak 57 Yellow-poplar 58 Yellow-poplar - eastern hemlock 59 Yellow poplar - white oak - northern red oak 70 Longleaf pine 75 Shortleaf pine 76 Shortleaf pine - oak 78 Virginia pine - oak 79 Virginia pine 80 Loblolly pine - shortleaf pine 81 Loblolly pine 82 Loblolly pine - hardwood 83 Longleaf pine - slash pine 97 Atlantic white-cedar SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : NO-ENTRY

VALUE AND USE

SPECIES: Rhododendron maximum | Rosebay
WOOD PRODUCTS VALUE : Rosebay wood is occasionally used to make small craft items and as fuel for wood stoves [19]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Deer occasionally browse rosebay leaves and twigs, but it is not a major part of their diet [28]. In a Greenwhich, Connecticut study, only 17 seventeen percent of rosebay shoots examined showed moderate leaf or twig damage from browsing [3]. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : Although nutrient concentrations of rosebay leaves are generally lower than those of deciduous trees, leaf longevity allows significant storage of nutrients. Fresh leaf litter has lower concentrations of nitrogen, potassium, and phosphorus than do harvested leaves, indicating a resorption of these nutrients before leaf fall. The nutrient concentration (% dry weight) of leaves changes with age [17]: Leaf Age Nitrogen Phosphorus Potassium Calcium Magnesium 1 mo 1.60 0.17 1.34 0.36 0.17 6 mo 1.05 0.16 0.87 1.00 0.29 2 yr 0.93 0.13 0.41 1.20 0.20 3 yr 0.86 0.12 0.35 1.30 0.21 4 yr 0.88 0.12 0.34 1.40 0.16 COVER VALUE : Rosebay provides cover for the sharp-tailed grouse, white-tailed deer, and a variety of other birds and mammals [20,28]. VALUE FOR REHABILITATION OF DISTURBED SITES : Rosebay is useful in providing watershed protection and erosion control on steep slopes. OTHER USES AND VALUES : Rosebay is cultivated for its beautiful flowers [20]. MANAGEMENT CONSIDERATIONS : Rosebay presents timber management problems throughout the Appalachian Mountains [30]. Rosebay regenerates vigorously through stump sprouting, root suckers, or from layered stems after it is cut, burned, or otherwise disturbed [30]. As it grows, it forms dense thickets that cast so much shade over the forest floor that tree species cannot reproduce, and eventually the site becomes unable to support commercial timber production [24]. Rosebay is best controlled through chemical treatment. Basal spraying and mist blower applications of 2,4,5-T have been reported to kill 100 percent of new sprouts [18,24].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Rhododendron maximum | Rosebay
GENERAL BOTANICAL CHARACTERISTICS : Rosebay is a spreading, thicket-forming, evergreen shrub or small tree, 3 to 15 feet (1-5 m) tall [6]. The flowers are large, round clusters; the fruit is a capsule containing many seeds. The bark is thin. [2,8,21]. Rosebay exhibits an interesting adaptation to climatic change. During periods of cold weather, the leaves roll lengthwise. The lower the temperature, the tighter the roll gets, until the leaves become slender hollow cylinders [13,19]. RAUNKIAER LIFE FORM : Microphanerophyte Nanophanerophyte REGENERATION PROCESSES : Seed and Seedlings: Rosebay does not produce large seed crops, and seeds do not remain viable past the first growing season. Seedlings are limited to open sites where there is a moist seedbed and little shade covering. In closed thickets, rosebay reproduces vegetatively [21,24]. Flowering: The flowering potential of rosebay is extremely limited. Flowering and fertilization reaches it greatest capacity when the shrub is 2 years old. Rosebay stops producing flowers after the fifth growing season [4,16]. Vegetative: Rosebay reproduces almost exclusively by stump sprouting, root suckers, or layering [30]. SITE CHARACTERISTICS : Rosebay occurs mostly at elevations below 3,300 feet (1,000 m) but in the southern Appalachians occasionally occurs at elevations over 5,900 feet (1,800 m). Rosebay flourishes on well-drained acidic soils in cool moist locations. It grows on shaded mountain sides, in ravines, and along bottoms [5,6,28]. Some trees commonly associated with rosebay are pitch pine (Pinus rigida), white pine (P. strobus), table mountain pine (P. pungens), numerous oaks (Quercus spp.), and hickories (Carya spp.). Some understory associates include mountain-laurel (Kalmia latifolia), blueberry (Vaccinium vacillans), black huckleberry (Gaylussacia bacata), wild grape (Vitis spp.), greenbrier (Smilax rotundifolia), fetterbush (Pieris floribunda), and white-alder (Clethra acuminata) [1,5,12]. SUCCESSIONAL STATUS : Rosebay is very shade tolerant and survives even in the deep shade of closed canopies [30]; however, new thicket formation is generally restricted to openings created by logging, fire, or other types of disturbance [21,2]. SEASONAL DEVELOPMENT : Rosebay flowers open during a period of several weeks from May to July. The fruit ripens in September through October; seed dispersal occurs from October to November [20].

FIRE ECOLOGY

SPECIES: Rhododendron maximum | Rosebay
FIRE ECOLOGY OR ADAPTATIONS : Rosebay is well adapted to fire. It typically survives by sprouting from the root crown or stump [10,23]. POSTFIRE REGENERATION STRATEGY : survivor species; on-site surviving root crown or caudex

FIRE EFFECTS

SPECIES: Rhododendron maximum | Rosebay
IMMEDIATE FIRE EFFECT ON PLANT : Fire usually kills aboveground stems smaller than 1.5 inches (4 cm) d.b.h. Larger stems often survive fire [21,24]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Rosebay typically sprouts vigorously after the aboveground portions are killed by fire. A fall prescribed burn in North Carolina top-killed 70 percent of rosebay less than 1 inch (2.5 cm) d.b.h., but 17 months after the burn, nearly all the top-killed rosebay had resprouted [10]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Rosebay is susceptible to annual burning. Annual intense burning for extended periods has eliminated rosebay populations on many sites in the Appalachians [10,24].

REFERENCES

SPECIES: Rhododendron maximum | Rosebay
REFERENCES : 1. Cain, Stanley A. 1931. Ecological studies of the vegetation of the Great Smoky Mountains of North Carolina and Tennessee. Botanical Gazette. 91: 22-41. [10340] 2. Chapman, William K.; Bessette, Alan E. 1990. Trees and shrubs of the Adirondacks. Utica, NY: North Country Books, Inc. 131 p. [12766] 3. Conover, M. R.; Kania, G. S. 1988. Browsing preference of white-tailed deer for different ornamental species. Wildlife Society Bulletin. 16: 175-179. [8933] 4. Cooper, S. D.; McGraw, J. B. 1988. Constraints on reproductive potential at the level of the shoot module in three ericaceous shrubs. Functional Ecology. 2: 97-108. [9039] 5. Della-Bianca, Lino. 1990. Pinus pungens Lamb. table mountain pine. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 425-432. [13400] 6. Duncan, Wilbur H.; Duncan, Marion B. 1988. Trees of the southeastern United States. Athens, GA: The University of Georgia Press. 322 p. [12764] 7. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 8. Fernald, Merritt Lyndon. 1950. Gray's manual of botany. [Corrections supplied by R. C. Rollins]. Portland, OR: Dioscorides Press. 1632 p. (Dudley, Theodore R., gen. ed.; Biosystematics, Floristic & Phylogeny Series; vol. 2). [14935] 9. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 10. Hooper, Ralph M. 1969. Prescribed burning for laurel and rhododendron control in the southern Appalachians. Res. Note SE-116. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station. 6 p. [10699] 11. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 12. Laderman, Aimlee D.; Golet, Francis C.; Sorrie, Bruce A.; Woolsey, Henry L. 1987. Atlantic white cedar in the glaciated Northeast. In: Laderman, Aimlee D., ed. Atlantic white cedar wetlands. [Place of publication unknown]: Westview Press: 19-34. [15872] 13. Leach, David G. 1963. Rhododendrons of the world and how to grow them. New York: Charles Scribner's Sons. 544 p. [10688] 14. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 15. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 16. McGraw, James B. 1989. Effects of age and size on life histories and population growth of Rhododendron maximum shoots. American Journal of Botany. 76(1): 113-123. [13488] 17. Monk, Carl D.; McGinty, Douglas T.; Day, Frank P., Jr. 1985. The ecological importance of Kalmia latifolia and Rhododendron maximum in the deciduous forest of the southern Appalachians. Bulletin of the Torrey Botanical Club. 112(2): 187-193. [16504] 18. Neary, D. G.; Douglass, J. E.; Ruehle, J. L.; Fox, W. 1984. Converting rhododendron-laurel thickets to white pine with picloram and mycorrhizae-inoculated seedlings. Southern Journal of Applied Forestry. 8(3): 163-168. [10697] 19. Nilsen, E. T. 1987. Influence of water relations and temperature on leaf movements of Rhododendron species. Plant Physiology. 83: 607-612. [10395] 20. Olson, David F., Jr. 1974. Rhododendron L. rhododendron. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 709-712. [7739] 21. Plocher, Allen E.; Carvell, Kenneth L. 1987. Population dynamics of rosebay rhododendron thickets in the southern Appalachians. Bulletin of the Torrey Botanical Club. 114(2): 121-126. [16505] 22. Radford, Albert E.; Ahles, Harry E.; Bell, C. Ritchie. 1968. Manual of the vascular flora of the Carolinas. Chapel Hill, NC: The University of North Carolina Press. 1183 p. [7606] 23. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 24. Romancier, Robert M. 1971. Combining fire and chemicals for the control of rhododondron thickets. Res. Note SE-149. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest and Range Experiment Station. 7 p. [10698] 25. Runkle, James Reade. 1982. Patterns of disturbance in some old-growth mesic forests of eastern North American. Ecology. 63(5): 1533-1546. [9261] 26. Sluder, Earl R. 1958. Control of cull trees and weed species in hardwood stands. Station Paper 95. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station. 13 p. [16503] 27. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 28. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 29. Whittaker, R. H. 1956. Vegetation of the Great Smoky Mountains. Ecological Monographs. 26(1): 1-79. [11108] 30. Yawney, Harry W. 1932. Control of rhododendron by basal spray. Res. Note No. 132. Upper Darby, PA: U.S. Department of Agriculture, Forest Service, Northeast Forest Experiment Station. 7 p. [16502]

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