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Introductory

SPECIES: Fraxinus nigra | Black Ash
ABBREVIATION : FRANIG SYNONYMS : NO-ENTRY SCS PLANT CODE : FRNI COMMON NAMES : black ash TAXONOMY : The currently accepted scientific name for black ash is Fraxinus nigra Marsh. [26]. There are no recognized subspecies, varieties, or forms. LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Milo Coladonato, February 1994 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Coladonato, Milo 1994. Fraxinus nigra. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Fraxinus nigra | Black Ash
GENERAL DISTRIBUTION : Black ash ranges from western Newfoundland west to southeastern Manitoba and eastern North Dakota; south to Iowa; east to southern Indiana, Ohio, and West Virginia; and north from northern Virginia to Delaware and New Jersey [6,17,30,37]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES19 Aspen - birch STATES : CT DE IL IN IA KY ME MD MA MI MN NH NJ NY ND OH PA RI VT VA WV WI MB NB NF NS ON PE PQ ADMINISTRATIVE UNITS : ACAD APIS CATO CUVA DEWA EFMO INDU ISRO MACA MORR PIRO SHEN SLBE VOYA BLM PHYSIOGRAPHIC REGIONS : NO-ENTRY KUCHLER PLANT ASSOCIATIONS : K081 Oak savanna K082 Mosaic of K074 and K100 K093 Great Lakes spruce - fir forest K094 Conifer bog K095 Great Lakes pine forest K096 Northeastern spruce - fir forest K099 Maple - basswood forest K100 Oak - hickory forest K101 Elm - ash forest K102 Beech - maple forest K104 Appalachian oak forest K106 Northern hardwoods K107 Northern hardwoods - fir forest K108 Northern hardwoods - spruce forest SAF COVER TYPES : 1 Jack pine 5 Balsam fir 12 Black spruce 15 Red pine 16 Aspen 17 Pin cherry 24 Hemlock - yellow birch 25 Sugar maple - beech - yellow birch 26 Sugar maple - basswood 27 Sugar maple 33 Red spruce - balsam fir 35 Paper birch - red spruce - balsam fir 37 Northern white-cedar 38 Tamarack 39 Black ash - American elm - red maple SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Black ash is a major hardwood type on lowlands in the northern Great Lake States along with American elm (Ulmus americana) and red maple (Acer rubrum) [13]. Black ash is also a common component in the beech-maple (Fagus-Acer spp.) climax community [35]. It is typically a seral species with black spruce (Picea mariana) in bogs or where there is excess water [36]. Black ash is a dominant species in northern Wisconsin and the Upper Peninsula of Michigan [13]. Black ash is a principle plant associate in the Hardwood Canadian Forest in the Saint Lawrence Valley [12]. In the uplands of Michigan and Wisconsin, black ash mingles with white ash, red maple, yellow birch (Betula alleghaniensis), American basswood (Tilia americana), and sugar maple (Acer saccharum) [13,18]. Common understory associates of black ash include speckled alder (Alnus rugosa), red-osier dogwood (Cornus sericea), bog-laurel (Kalmia polifolia), Labrador-tea (Ledum groenlandicum), poison-sumac (Toxicodendron vernix), willows (Salix spp.), low sweet blueberry (Vaccinum angustifolium), highbush blueberry (V. corymbosum), small cranberry (V. oxycoccus), and common winterberry (Ilex verticillata) [15,22,23,29].

VALUE AND USE

SPECIES: Fraxinus nigra | Black Ash
WOOD PRODUCTS VALUE : Black ash wood is not as strong nor as hard as that of other ashes. It is used for interior finishing, furniture, and cabinets [21]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Black ash twigs and leaves provide spring and summer browse for white-tailed deer and moose [11,19]. The seeds are an important food for game birds, songbirds, and small mammals [37]. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : Black ash grows very slowly on organic peats and mucks, attaining heights of only 30 to 45 feet (9-13 m) at 50 years and 50 to 60 feet (15-18 m) at 100 years. The best trees average only 10 inches (25 cm) d.b.h. at 110 years and 12 inches (30 cm) at 130 years. Black ash seedlings, saplings, and sprouts tend to dominate the regeneration layer where partial openings in the canopy have occurred. Clearcutting often results in inadequate natural regeneration of black ash or loss of advanced regeneration because of rising water tables or increased competition with grass and brush. Clearcuts are often winter habitat; deer browse heavily on black ash seedlings and stump sprouts [13]. Most swamp hardwood stands contain many defective and diseased trees. Many unmanaged, fully stocked hardwood stands contain no desirable advanced black ash regeneration. Trees on wet sites are shallowly rooted and subject to windthrow [13].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Fraxinus nigra | Black Ash
GENERAL BOTANICAL CHARACTERISTICS : Black ash is a small, native, deciduous tree ranging from 40 to 60 feet (12-18 m) in height. The tree has coarse ascending branches and a slender, sometimes bent or leaning trunk which extends almost to the top of a narrow crown. The compound leaves are 10 to 16 inches (25-40 cm) long. Black ash is polygamous; its flowers are small and inconspicuous [37]. The bark is shallowly fissured and divided into large irregular plates with thin, soft, papery scales that rub off easily. The fruit is an elongated, winged, single-seeded samara that is borne in terminal or axillary clusters. Black ash has a shallow, wide-spreading root system [8,9,21]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Black ash regenerates through sexual and vegetative reproduction. Good seed crops occur at 1- to 8-year intervals with most intervening years having poor seed crops [13,34]. Germination: Black ash produces an abundance of seed [4]. The seed contains a dormant embryo that requires stratification and a period of cold temperatures for good germination. Before embryos germinate, they must grow from about one-half to two-thirds the length of the seed, so most black ash seed do not germinate until the second spring after seedfall. Some seed may lie dormant in the litter for up to 8 years [13]. Seedling development: Black ash seed is capable of germinating in hardwood leaf litter or under 0.25- to 0.75-inch (0.6-1.8 cm) of soil; however, grass, brush, and advance hardwood reproduction must be controlled for successful black ash seedling establishment. Germinants are about 2 inches (5 cm) tall within 2 weeks. Under the best conditions they may average 6 inches (15 cm) in height by the end of their first growing season [13]. Black ash seedlings often grow more slowly than seedlings of associated species such as American elm and red maple [37]. Vegetative Reproduction: Black ash sprouts readily from stumps up to 12 inches (30 cm) in diameter. Sprouts originate from adventitious buds on the sides of the stump and at the root crown. Stump sprouts can exhibit fast growth. Black ash will also root sucker [13]. SITE CHARACTERISTICS : Black ash is most commonly found growing in moist to wet muck or shallow organic soils. It is found in swamps, along small streams in gullies, and in small poorly drained depressions [25,37]. It also grows on fine sands and loams underlain by clays and on other poorly drained sites with high water tables. In uplands black ash is restricted to sites with impeded drainage, where it grows on wetter than normal mineral soils [5,13]. SUCCESSIONAL STATUS : Facultative Seral Species In the Great Lake States black ash is a pioneer. It is also present but not abundant in mature forests dominated by elm (Ulmus spp.), cedar (Thuja spp.), larch (Larix spp.), spruce (Picea spp.), fir (Abies spp.), and birch (Betula spp.) [1,10]. It is an invader species in sedge meadows in southern Wisconsin [28]. In northeastern Minnesota, black ash seedlings invade open areas in maple-beech forests [7]. Black ash is shade intolerant [37]. SEASONAL DEVELOPMENT : The flowers of black ash appear in May or June, concurrent with or just before the leaves [37]. The fruits ripen from June to September and are dispersed from July to October [34,35].

FIRE ECOLOGY

SPECIES: Fraxinus nigra | Black Ash
FIRE ECOLOGY OR ADAPTATIONS : Black ash is easily damaged by fire [2,33]. Prior to 1871 in Quebec, ash forests containing black ash and American elm burned in mostly stand-replacing fires, with a fire cycle of about 63 years [2]. In northwestern Minnesota and west-central Canada, fires swept into boreal forests from open prairies, killing black ash and other fire-sensitive species [20]. Little information is available concerning black ash fire survival strategies. It probably survives fire by sprouting and colonizes burned sites through wind-dispersed seed. POSTFIRE REGENERATION STRATEGY : Tree with adventitious-bud root crown/soboliferous species root sucker Initial-offsite colonizer (off-site, initial community)

FIRE EFFECTS

SPECIES: Fraxinus nigra | Black Ash
IMMEDIATE FIRE EFFECT ON PLANT : Depending on fire severity, black ash is probably killed or top-killed by fire. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Black ash probably sprouts from the root crown following top-damage from fire. It is also probable that this tree, a prolific seeder, regenerates from seed [4]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Fire suppression has allowed the invasion of black ash into sedge meadows in southern Wisconsin [28].

REFERENCES

SPECIES: Fraxinus nigra | Black Ash
REFERENCES : 1. Barnes, Burton V. 1976. Succession in deciduous swamp communities of southeastern Michigan formerly dominated by American elm. Canadian Journal of Botany. 54: 19-24. [4914] 2. Bergeron, Yves. 1991. The influence of island and mainland lakeshore landscapes on boreal forest fire regimes. Ecology. 72(6): 1980-1992. [17186] 3. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 4. Bonner, F. T. 1974. Fraxinus ash. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United States. Agriculture Handbook No. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 411-416. [7668] 5. Brand, Gary J. 1985. Environmental indices for common Michigan trees and shrubs. Res. Pap. NC-261. St. Paul, MN: U.S. Department of Agriculture, Forest Service, Northcentral Forest Experiment Station. 5 p. [14465] 6. Braun, E. Lucy. 1961. The woody plants of Ohio. Columbus, OH: Ohio State University Press. 362 p. [12914] 7. Bray, J. Roger. 1956. Gap phase replacement in a maple-basswood forest. Ecology. 37(3): 598-600. [13003] 8. Brundrett, Mark; Murase, Gracia; Kendrick, Bryce. 1990. Comparative anatomy of roots and mycorrhizae of common Ontario trees. Canadian Journal of Botany. 68: 551-578. [11380] 9. Collingwood, G. H. 1937. Knowing your trees. Washington, DC: The American Forestry Association. 213 p. [6316] 10. Cooper, William S. 1913. The climax forest of Isle Royale, Lake Superior, and its development. III. Botanical Gazette. 55(3): 189-235. [11539] 11. Cumming, H. G. 1987. Sixteen years of moose browse surveys in Ontario. Alces. 23: 125-156. [8859] 12. Dansereau, Pierre. 1959. The principal plant associations of the Saint Lawrence Valley. No. 75. Montreal, Canada: Contrib. Inst. Bot. Univ. Montreal. 147 p. [8925] 13. Erdmann, Gayne G.; Crow, Thomas R.; Peterson, Ralph M., Jr.; Wilson, Curtis D. 1987. Managing black ash in the Lake States. Gen. Tech. Rep. NC-115. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Forest Experiment Station. 10 p. [22296] 14. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 15. Frank, Robert M. 1990. Abies balsamea (L.) Mill. balsam fir. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 26-35. [13365] 16. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 17. Gleason, Henry A.; Cronquist, Arthur. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. 2nd ed. New York: New York Botanical Garden. 910 p. [20329] 18. Godman, Richard M.; Mattson, Gilbert A. 1976. Seed crops and regeneration problems of 19 species in northeastern Wisconsin. Res. Pap. NC-123. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Forest Experiment Station. 5 p. [3715] 19. Habeck, J. R.; Curtis, J. T. 1959. Forest cover and deer population densities in early northern Wisconsin. Transactions, Wisconsin Academy of Sciences, Arts and Letters. 48: 49-56. [16405] 20. Heinselman, Miron L. 1981. Fire intensity and frequency as factors in the distribution and structure of northern ecosystems. In: Mooney, H. A.; Bonnicksen, T. M.; Christensen, N. L.; [and others], technical coordinators. Fire regimes and ecosystem properties: Proceedings of the conference; 1978 December 11-15; Honolulu, HI. Gen. Tech. Rep. WO-26. Washington, DC: U.S. Department of Agriculture, Forest Service: 7-57. [4390] 21. Hosie, R. C. 1969. Native trees of Canada. 7th ed. Ottawa, ON: Canadian Forestry Service, Department of Fisheries and Forestry. 380 p. [3375] 22. Johnston, William F. 1990. Larix laricina (Du Roi) K. Koch tamarack. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 141-151. [13379] 23. Johnston, William F. 1990. Thuja occidentalis L. northern white-cedar. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 580-589. [13418] 24. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 25. Levy, Gerald F. 1970. The phytosociology of northern Wisconsin upland openings. American Midland Naturalist. 83: 213-237. [9986] 26. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 27. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 28. Reuter, D. Dayton. 1986. Effects of prescribed burning, cutting and torching on shrubs in a sedge meadow wetland. In: Koonce, Andrea L., ed. Prescribed burning in the Midwest: state-of-the-art: Proceedings of a symposium; 1986 March 3-6; Stevens Point, WI. Stevens Point, WI: University of Wisconsin, College of Natural Resources, Fire Science Center: 108-115. [16278] 29. Rudolf, Paul O. 1990. Pinus resinosa Ait. red pine. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America. Volume 1. Conifers. Agric. Handb. 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 442-455. [13246] 30. Stephens, H. A. 1973. Woody plants of the North Central Plains. Lawrence, KS: The University Press of Kansas. 530 p. [3804] 31. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 32. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 33. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 34. Wang, B. S. P. 1974. Tree-seed storage. Publication No. 1335. Ottawa, Canada: Department of the Environment, Canadian Forestry Service. 32 p. [17267] 35. Williams, Arthur B. 1936. The composition and dynamics of a beech-maple climax community. Ecological Monographs. 6(3): 318-408. [8346] 36. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. [2620] 37. Wright, Jonathan W.; Rauscher, H. Michael. 1990. Fraxinus nigra Marsh. black ash. In: Burns, Russell M.; Honkala, Barbara H., technical coordinators. Silvics of North America: Vol. 2. Hardwoods. Agriculture Handbook 654. Washington, DC: U.S. Department of Agriculture, Forest Service: 344-347. [22499]

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