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Introductory

SPECIES: Quercus laurifolia | Laurel Oak
ABBREVIATION : QUELAU SYNONYMS : Quercus laurifolia Michx. Quercus hemisphaerica Bartr. Quercus hemisphaerica var. maritima (Michx.) Quercus obtusa (Willd.) Ashe Quercus phellos L. var. laurifolia (Michx.) Chapman Quercus succulenta Small Quercus virginiana P. Mill. var. maritima (Michx.) Sarg. SCS PLANT CODE : QULA3 COMMON NAMES : laurel oak Darlington oak diamond-leaf oak swamp laurel oak laurel-leaf oak water oak obtusa oak spotted oak coastal laurel oak TAXONOMY : The historical nomenclature of laurel oak is complicated. In the past, most authorities, including Little [24], treated laurel oaks as a single species but differed on the appropriate scientific name [43]. More recent authorities [4,8,40,43] recognize two species, Quercus laurifolia and Q. hemisphaerica, based on anatomical differences and vast differences in site preferences. Diamond-leaf oak, swamp laurel oak, and laurel oak are common names for the wetland form, currently called Q. laurifolia and formerly called Q. obtusa. The upland form, currently called Q. hemisphaerica by some authorities, has acute leaf tips and flowers 2 weeks later than the wetland form in the same area [8,12,27]. This report recognizes one species of laurel oak under the scientific name of Quercus laurifolia L. [24,27]. Information from authors that recognize and discuss Q. hemisphaerica as a separate species is included and noted as such. Laurel oak has been placed within the subgenus Erythrobalanus, or black (red) oak group. Laurel oak is difficult to identify and is often confused with willow oak (Q. phellos) and water oak (Q. nigra) [40]. It has been suggested that laurel oak is a hybrid between these two species, but that may not be the case because willow oak is absent from southeastern Georgia and peninsular Florida where laurel oak is abundant [27]. Laurel oak hybridizes with the following species [24,27]: x Q. falcata (southern red oak): Q. X. beaumontiana Sarg. x Q. incana (bluejack oak): Q. X. atlantica Ashe x Q. laevis (turkey oak): Q. X. mellichampii Trel. x Q. marilandica (blackjack oak): Q. X. diversiloba Tharp ex A. Camus LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Jennifer H. Carey, Sept. 1992 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Carey, Jennifer H. 1992. Quercus laurifolia. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Quercus laurifolia | Laurel Oak
GENERAL DISTRIBUTION : Laurel oak occurs on the Atlantic and Gulf coastal plains of the southeastern United States from southeastern Virginia to southern Florida and west to the extreme southeastern Gulf Coast of Texas. Disjunct populations occur north of its contiguous coastal range in Texas, Arkansas, Louisiana, Alabama, Mississippi, Georgia, and South Carolina [27]. ECOSYSTEMS : FRES12 Longleaf - slash pine FRES13 Loblolly - shortleaf pine FRES14 Oak - pine FRES16 Oak - gum - cypress STATES : AL AR FL GA LA MS NC SC VA TN TX ADMINISTRATIVE UNITS : BICY BITH CAHA CALO COLO COSW CUIS EVER FOCA NATR BLM PHYSIOGRAPHIC REGIONS : NO-ENTRY KUCHLER PLANT ASSOCIATIONS : K111 Oak - hickory - pine forest K112 Southern mixed forest K113 Southern floodplain forest SAF COVER TYPES : 74 Cabbage palmetto 81 Loblolly pine 83 Longleaf pine - slash pine 84 Slash pine 88 Willow oak - water oak - diamondleaf oak 92 Sweetgum - willow oak 102 Baldcypress - tupelo 103 Water tupelo - swamp tupelo 104 Sweetbay - swamp tupelo - redbay SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Laurel oak is often present in forested wetlands, a transitional community between swamps and pine (Pinus spp.) flatlands or mesic hammocks. It grows throughout hydric hammocks, from the swamp margin to the drier sections, being replaced at the very dry end by live oak (Q. virginiana) and water oak [43]. Q. hemisphaerica is present in high hammocks which are situated between sand or clay hills and midslope hammocks [34]. The following published classifications list laurel oak as a dominant species: The natural communities of South Carolina [16] Eastern deciduous forest [45] Forest vegetation of the Big Thicket, southeast Texas [26] Forest associations in the uplands of the lower Gulf Coastal Plain [33] The natural features of southern Florida [6] Ecological processes and vegetation of the maritime strand in the southeastern United States [32]

VALUE AND USE

SPECIES: Quercus laurifolia | Laurel Oak
WOOD PRODUCTS VALUE : Although laurel oak has hard, heavy, and strong wood, it is not good quality lumber. It is marketed for pulp wood and also used for firewood [8,27]. IMPORTANCE TO LIVESTOCK AND WILDLIFE : The consistent and abundant acorn crops are an important food source for many animals, including white-tailed deer, raccoon, squirrels, wild turkey, ducks, quail, smaller birds, and rodents. Laurel oak ranked second in quantity and frequency of acorns consumed by wild turkey in Florida. In a study of the 10 most heavily used winter foods of deer in Florida, laurel oak acorns rated fifth, sixth, and seventh in a 6-year period. Acorns of the black oak group do not germinate until spring, unlike those of the white oak group, and are an important winter food source [27]. PALATABILITY : Laurel oak is considered a good deer browse [38]. NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : Laurel oak has attractive leaves and is often planted as an ornamental [27]. MANAGEMENT CONSIDERATIONS : Laurel oak is susceptible to oak leaf blister (Taphrina caerulescens), actinopelte spot (Actinopelte dryina), and canker rots by various fungi. Although not damaged itself, laurel oak is a very susceptible host for the alternative stage of fusiform rust (Cronartium quercuum f. sp. fusiforme), a serious disease of southern pines. Weevils (Curculio spp.) infest the acorns of laurel oak [27]. Laurel oak will grow well on moderately drained sites using several silvicultural systems. Natural regeneration is possible using the shelterwood system but requires a relatively high density of 35 to 40 square feet basal area per acre (3.7-4.2 sq m/ha). If clearcutting is used, direct seeding is the best method to regenerate laurel oak. Planting seedlings may be difficult because of poor drainage and difficult access into bottomland sites [15].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Quercus laurifolia | Laurel Oak
GENERAL BOTANICAL CHARACTERISTICS : Laurel oak is a rapidly growing, short-lived, semi-evergreen tree. It can reach 148 feet (45 m) in height and 6.6 feet (2 m) in d.b.h. Laurel oak identified as Q. hemisphaerica is slightly smaller at 131 feet (40 m) [8], and a geographic or climatic form in east Texas grows to only 30 feet (9.1 m) [40]. Poor site conditions may be responsible for the smaller heights reported for Q. hemisphaerica. Laurel oak develops a large, well-defined taproot on upland sands, but little else is known about its rooting habit [27]. Roots of trees growing in wet areas are often buttressed, which provides stability in wet soils and may help aerate the root system [43]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Sexual: Laurel oak is monoecious. Acorn production begins when the tree reaches 15 to 20 years of age. Laurel oak produces abundant flowers almost every year and is a prolific seed bearer. Dissemination of the heavy acorns is mainly by squirrels, but gravity and water also play a role [27]. Germination is hypogeal. Although laurel oak acorns generally germinate in the spring, they exhibit only mild dormancy [27]. In one study, acorns showed a 50 percent germinative capacity without cold stratification [31]. Germination is unaffected or even slightly increased by soaking acorns in water, a condition frequently encountered in bottomland forests [48]. Seedlings grow rapidly [27]. Vegetative: If cut or burned, laurel oak sprouts from the base of the stump. Older trees do not sprout vigorously, and their sprouts are more susceptible to decay than those of younger trees [27]. SITE CHARACTERISTICS : Laurel oak is generally considered a bottomland [47] or facultative wetland species [1]. It is not as dependent on specific site conditions as other hardwood associates [30]. It generally grows in soils that are better drained than where water oak and willow oak grow, but grows in very wet sites as well [40]. Laurel oak commonly grows on alluvial flood plains and sandy soils near rivers, swamps, and hammocks. It grows best on soils of the Ultisol and Inceptisol orders [27]. Laurel oak is moderately tolerant of flooding [11,43]. It is more tolerant of prolonged soil saturation than is water oak or live oak, but it cannot survive inundation during the entire growing season. Short periods of deep inundation have been known to kill laurel oak [43]. Laurel oak grows on high lands surrounding swamps and major rivers which flood deeply and frequently but drain rapidly because of relief. It also grows on wet flats which are better drained than swamps [18]. In addition to bottomland forests, laurel oak is found in bay swamps, mixed hardwood swamps, river swamps, hydric hammocks, and cypress (Taxodium spp.) ponds and strands [11,39]. Laurel oak also grows on barrier islands off the Atlantic Coast [17,21]. On an island off South Carolina, laurel and live oaks grow 0.3 mile (0.5 km) inland from the ocean on the southern end of the island and to the beach on the northern end [17]. Q. hemisphaerica grows on dry, sandy sites [11] including stable dunes of beaches and islands [20], mesic and xeric hammocks, and sandhills [7,42]. In east Texas, a form of laurel oak, described as Q. hemisphaerica, forms extensive mottes on sandy hammocks and prairies [40]. In addition to trees mentioned in SAF Cover Types; Habitat Types and Plant Communities; and Taxonomy, overstory associates include Nuttall oak (Q. nuttallii), white oak (Q. alba), yellow-poplar (Liriodendron tulipifera), red maple (Acer rubrum), green ash (Fraxinus pennsylvanica), swamp hickory (Carya glabra), and honeylocust (Gleditsia triacanthos); and on wetter sites water hickory (Carya aquatica), waterlocust (Gleditsia aquatica), and overcup oak (Q. lyrata). On better-drained sites, laurel oak is associated with spruce pine (Pinus glabra), swamp chestnut oak (Q. michauxii), and cherrybark oak (Q. falcata var. pagodifolia). Common associates in Florida are southern magnolia (Magnolia grandifolia), pignut hickory (Carya glabra), scrub hickory (C. floridana), and Carolina basswood (Tilia caroliniana). Associated shrubs and small trees include American hornbeam (Carpinus caroliniana), Virgnia-willow (Itea virginica), poison-sumac (Toxicodendron vernix), swamp cyrilla (Cyrilla racemiflora), littleleaf cyrilla (C. racemiflora var. parvifolia), sebastian bush (Sebastiana ligustrina), dahoon (Ilex cassine), possumhaw (I. decidua), swamp dogwood (Cornus stricta), sweet pepperbush (Clethra alnifolia), tree lyonia (lyonia ferruginea), buttonbush (Cephalanthus occidentalis), pinckneya (Pinckneya pubens), and rhododendron (Rhododendron spp.). Associated vines include coral greenbrier (Smilax walteri), laurelleaf greenbrier (S. laurifolia), crossvine (Bignonia capreolata), muscadine grape (Vitis rotundifolia), and Alabama supplejack (Berchemia scandens) [27]. SUCCESSIONAL STATUS : Facultative Seral Species Laurel oak is shade tolerant and often becomes established under and grows up through a dense canopy [27]. Monk [28] considers laurel oak a climax pioneer species because it invades successional communities and is instrumental in the conversion to climax. It has produced successful seedlings beneath a laurel oak canopy [5]. Laurel oak is also an early invader on some sites. It will invade early seral wetlands if a seed source is nearby [9]. With fire suppression, laurel and willow oaks invaded a wetland savanna in the Big Thicket area of east Texas. The savanna eventually became an oak flat with very little growing beneath the oaks and standing water much of the year [46]. In the absence of fire, laurel oak (including Q. hemisphaerica of [10,29,42]) invades and becomes established on former longleaf pine (Pinus palustris) sites and sandhill sites [5,13,36]. Laurel oak was found on upland longleaf sites with colored sand, but not on harsher white sand soils which are leached and well sorted [5]. SEASONAL DEVELOPMENT : Laurel oak sustains high leaf fall production from October to March [43]. It flowers in February or March, at the same time that the last of the previous year's leaves are shed. Acorns mature after 2 years and fall in late September and October. Germination occurs in the spring [27].

FIRE ECOLOGY

SPECIES: Quercus laurifolia | Laurel Oak
FIRE ECOLOGY OR ADAPTATIONS : Laurel oak is fire intolerant. It is frequently top-killed by even low-severity surface fires because it has relatively thin bark. It is also a poor natural pruner [27]. Many laurel oak stands such as those on hydric hammocks owe their existence to protection from fire [43]. Hardwood hammocks are extremely susceptible to fire damage, especially during the dry season. A dry-season surface fire may burn the organic soil down to the bedrock [44]. If fire is suppressed, laurel oak expands from hydric hammocks into adjacent communities [43]. Unlike the original hammock, expanding hammocks often have a dense saw-palmetto (Serenoa repens) understory. A dry-season fire in Myakka River State Park, Florida, killed many large laurel oaks in the expanding hammock but not in the original hammock. The dense saw-palmetto understory was, in part, responsible for the high mortality of laurel oak in the expanding hammock [19]. POSTFIRE REGENERATION STRATEGY : Tree with adventitious-bud root crown/root sucker Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Quercus laurifolia | Laurel Oak
IMMEDIATE FIRE EFFECT ON PLANT : Laurel oak smaller than 3 inches (7.6 cm) in d.b.h. can be top-killed by low-severity fire [2]. More severe fires may completely kill this fire-sensitive species [27,34]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Young laurel oak sprouts vigorously from the root crown if top-killed by fire. Older trees do not sprout as readily. Trees subject to occasional fires commonly develop heart rot where fire wounded [27]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Oak-dominated forests on elevated margins of wetland ecosystems are often converted to loblolly (Pinus taeda) and slash pine (P. elliottii) plantations because of the high productivity potential of these sites. Prescibed fire is used to prevent hardwood establishment in the plantations [3]. Laurel oak up to 3 inches (7.6 cm) in d.b.h. can be top-killed and sprouts kept small and controllable with prescribed winter fires. Summer fires are also effective at hardwood control, but do not enhance the wildlife food supply [2].

REFERENCES

SPECIES: Quercus laurifolia | Laurel Oak
REFERENCES : 1. Best, G. Ronnie; Segal, Debra S.; Wolfe, Charlotte. 1990. Soil-vegetation correlations in selected wetlands and uplands of north-central Florida. Biol. Rep. 90(9). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 51 p. [18161] 2. Harlow, Richard F.; Bielling, Paul. 1961. Controlled burning studies in longleaf pine-turkey oak association on the Ocala National Forest. Proceeding, Annual Conference of Southeastern Association of Game and Fish. 15: 9-24. [9905] 3. Christensen, Norman L. 1981. Fire regimes in southeastern ecosystems. In: Mooney, H. A.; Bonnicksen, T. M.; Christensen, N. L.; [and others], technical coordinators. Fire regimes and ecosystem properties: Proceedings of the conference; 1978 December 11-15; Honolulu, HI. Gen. Tech. Rep. WO-26. Washington, DC: U.S. Department of Agriculture, Forest Service: 112-136. [4391] 4. Clewell, Andre F. 1985. Guide to the vascular plants of the Florida Panhandle. Tallahassee, FL: Florida State University Press. 605 p. [13124] 5. Daubenmire, Rexford. 1990. The Magnolia grandiflora-Quercus virginiana forest of Florida. American Midland Naturalist. 123: 331-347. [10871] 6. Davis, John H., Jr. 1943. The natural features of southern Florida especially the vegetation, and the Everglades. Geological Bull. No. 25. Tallahassee, FL: State of Florida, Department of Conservation, Florida Geological Survey. 311 p. [17747] 7. Duever, Linda Conway. 1983. Natural communities of Florida's inland sand ridges. Palmetto. Winter Park, FL: Florida Native Plant Society; 3(3): 1-3, 10. [18775] 8. Duncan, Wilbur H.; Duncan, Marion B. 1988. Trees of the southeastern United States. Athens, GA: The University of Georgia Press. 322 p. [12764] 9. Dunn, William J. 1989. Wetland succession--what is the appropriate paradigm?. In: Fisk, David W., ed. Wetlands: concerns and successes: Proceedings of the symposium; 1989 September 17-22; Tampa, FL. Bethesda, MD: American Water Resources Association: 473-488. [17128] 10. Engstrom, R. Todd; Crawford, Robert L.; Baker, W. Wilson. 1984. Breeding bird populations in relation to changing forest structure following fire exclusion: a 15-year study. Wilson Bulletin. 96(3): 437-450. [9873] 11. Ewel, Katherine C. 1990. Swamps. In: Myers, Ronald L.; Ewel, John J., eds. Ecosystems of Florida. Orlando, FL: University of Central Florida Press: 281-322. [17392] 12. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 13. Garren, Kenneth H. 1943. Effects of fire on vegetation of the southeastern United States. Botanical Review. 9: 617-654. [9517] 14. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 15. Van Wagner, C. E. 1967. Seasonal variation in moisture content of eastern Canadian tree foliage and the possible effect on crown fires. Departmental Publ. No. 1204. Ottawa, Canada: Department of Forestry and Rural Development, Forestry Branch. 15 p. [15404] 16. Nelson, John B. 1986. The natural communities of South Carolina. Columbia, SC: South Carolina Wildlife & Marine Resources Department. 54 p. [15578] 17. Helm, A. C.; Nicholas, N. S.; Zedaker, S. M.; Young, S. T. 1991. Maritime forests on Bull Island, Cape Romain, South Carolina. Bulletin of the Torrey Botanical Club. 118(2): 170-175. [15686] 18. Hook, Donal D. 1978. Management of wetland hardwoods for timber production. In: Balmer, William E., ed. Proceedings--soil moisure...site productivity symposium; 1977 November 1-3; Myrtle Beach, SC. Atlanta, GA: U.S. Department of Agriculture, Forest Service, Southeastern Area, State and Private Forestry: 237-243. [4267] 19. Huffman, Jean M.; Blanchard, S. W. 1991. Changes in woody vegetation in Florida dry prairie and wetlands during a period of fire exclusion, and after dry-growing-season fire. In: Nodvin, Stephen C.; Waldrop, Thomas A., eds. Fire and the environment: ecological and cultural perspectives: Proceedings of an international symposium; 1990 March 20-24; Knoxville, TN. Gen. Tech. Rep. SE-69. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station: 75-83. [16636] 20. Johnson, Ann F.; Barbour, Michael G. 1990. Dunes and maritime forests. In: Myers, Ronald L.; Ewel, John J., eds. Ecosystems of Florida. Orlando, FL: University of Central Florida Press: 430-480. [17394] 21. Johnson, A. Sydney; Hillestad, Hilburn O.; Shanholtzer, Sheryl Fanning; Shanholtzer, G. Frederick. 1974. An ecological survey of the coastal region of Georgia. Scientific Monograph Series No 3, NPS 116. Washington, DC: U.S. Department of the Interior, National Park Service. 233 p. [16102] 22. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. Volume II: The biota of North America. Chapel Hill, NC: The University of North Carolina Press; in confederation with Anne H. Lindsey and C. Richie Bell, North Carolina Botanical Garden. 500 p. [6954] 23. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 24. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 25. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 26. Marks, P. L.; Harcombe, P. A. 1981. Forest vegetation of the Big Thicket, southeast Texas. Ecological Monographs. 51(3): 287-305. [9672] 27. Curtis, James D. 1946. Preliminary observations on northern white cedar in Maine. Ecology. 27: 23-36. [19804] 28. Monk, Carl D. 1968. Successional and environmental relationships of the forest vegetation of north central Florida. American Midland Naturalist. 79(2): 441-457. [10847] 29. Myers, Ronald; White, Deborah L. 1987. Landscape history and changes in sandhill vegetation in north-central and south-central Florida. Bulletin of the Torrey Botanical Club. 114(1): 21-32. [9782] 30. Nixon, Elray S.; Willett, R. Larry; Cox, Paul W. 1977. Woody vegetation of a virgin forest in an eastern Texas river bottom. Castanea. 42: 227-236. [9898] 31. Olson, David F., Jr. 1974. Quercus L. oak. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 692-703. [7737] 32. Oosting, Henry J. 1954. Ecological processes and vegetation of the maritime strand in the southeastern United States. Botanical Review. 20: 226-262. [10730] 33. Pessin, L. J. 1933. Forest associations in the uplands of the lower Gulf Coastal Plain (longleaf pine belt). Ecology. 14(1): 1-14. [12389] 34. Platt, William J.; Schwartz, Mark W. 1990. Temperate hardwood forests. In: Myers, Ronald L.; Ewel, John J., eds. Ecosystems of Florida. Orlando, FL: University of Central Florida Press: 194-229. [17390] 35. Putnam, John A. 1951. Management of bottomland hardwoods. Occasional Paper 116. New Orleans, LA: U.S. Department of Agriculture, Forest Service, Southern Forest Experiment Station. 60 p. [6748] 36. Quarterman, Elsie; Keever, Catherine. 1962. Southern mixed hardwood forest: climax in the southeastern coastal plain, U.S.A. Ecological Monographs. 32: 167-185. [10801] 37. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 38. Reid, Vincent H.; Goodrum, Phil D. 1957. The effect of hardwood removal on wildlife. In: Proceedings of the Society of American Foresters meeting; 1957 November 10-13; Syracuse, NY. Washington, DC: Society of American Foresters: 141-147. [10477] 39. Richardson, Donald Robert. 1977. Vegetation of the Atlantic Coastal Ridge of Palm Beach County, Florida. Florida Scientist. 40(4): 281-330. [9644] 40. Simpson, Benny J. 1988. A field guide to Texas trees. Austin, TX: Texas Monthly Press. 372 p. [11708] 41. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 42. Veno, Patricia Ann. 1976. Successional relationships of five Florida plant communities. Ecology. 57: 498-508. [9659] 43. Vince, Susan W.; Humphrey, Stephen R.; Simons, Robert W. 1989. The ecology of hydric hammocks: A community profile. Biological Rep. 85(7.26). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service, Research and Development. 82 p. [17976] 44. Wade, Dale; Ewel, John; Hofstetter, Ronald. 1980. Fire in south Florida ecosystems. Gen. Tech. Rep. SE-17. Asheville, NC: U.S. Department of Agriculture, Forest Service, Southeastern Forest Experiment Station. 125 p. [10363] 45. Waggoner, Gary S. 1975. Eastern deciduous forest, Vol. 1: Southeastern evergreen and oak-pine region. Natural History Theme Studies No. 1, NPS 135. Washington, DC: U.S. Department of the Interior, National Park Service. 206 p. [16103] 46. Watson, Geraldine E. 1986. Influence of fire on the longleaf pine - bluestem range in the Big Thicket region. In: Kulhavy, D. L.; Conner, R. N., eds. Wilderness and natural areas in the eastern United States: a management challenge. Nacogdoches, TX: Stephen F. Austin University: 181-185. [10334] 47. Wyant, James G.; Alig, Ralph J.; Bechtold, William A. 1991. Physiographic position, disturbance and species composition in North Carolina coastal plain forests. Forest Ecology and Management. 41: 1-19. [15373] 48. Larsen, Harry S. 1963. Effects of soaking in water on acorn germination of four southern oaks. Forest Science. 9(2): 236-241. [18903]

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