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Introductory

SPECIES: Salix amygdaloides | Peachleaf Willow
ABBREVIATION : SALAMY SYNONYMS : Salix amygdaloides var. wrightii (Anderss.) Schneid. Salix nigra Marsh. var. amygdaloides (Anderss.) Anderss. Salix wrightii Anderss. SCS PLANT CODE : SAAM2 COMMON NAMES : peachleaf willow peach-leaved willow peach willow southwestern peach willow almond willow almondleaf willow Wright willow TAXONOMY : The currently accepted scientific name of peachleaf willow is Salix amygdaloides Anderss. [27]. No subspecies, varieties, or forms are recognized. LIFE FORM : Tree FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Ronald Uchytil/March 1989 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Uchytil, Ronald J. 1989. Salix amygdaloides. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Salix amygdaloides | Peachleaf Willow
GENERAL DISTRIBUTION : Peachleaf willow is distributed from Quebec and New York west across southern Canada to British Columbia and south to Washington, Oregon, Utah, Arizona, Texas, Kentucky, and Vermont [26,27,37]. ECOSYSTEMS : FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES21 Ponderosa pine FRES28 Western hardwoods FRES29 Sagebrush FRES35 Pinyon - juniper FRES38 Plains grasslands FRES39 Prairie STATES : AZ CO ID IL IN IA KS KY MI MN MO MT NE NV NM NY ND OH OK OR PA SD TX UT VT WA WI WY AB BC MB ON PQ SK MEXICO ADMINISTRATIVE UNITS : AGFO ARCH BADL BAND BICA CACH CANY CODA CUVA DETO GLAC GLCA INDU LAME MOCA PIPE SLBE THRO VOYA WICA BLM PHYSIOGRAPHIC REGIONS : 2 Cascade Mountains 5 Columbia Plateau 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K023 Juniper - pinyon woodland K037 Mountain mahogany - oak scrub K038 Great Basin sagebrush K039 Blackbrush K040 Saltbush - greasewood K041 Creosotebush K051 Wheatgrass - bluegrass K055 Sagebrush steppe K057 Galleta - threeawn shrubsteppe K063 Foothills prairie K065 Grama - needlegrass - wheatgrass K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem - needlegrass K070 Sandsage - bluestem prairie K074 Bluestem prairie K075 Nebraska Sandhills prairie K081 Oak savanna K098 Northern floodplain forest K101 Elm - ash forest SAF COVER TYPES : 39 Black ash - American elm - red maple 61 River birch - sycamore 63 Cottonwood 93 Sugarberry - American elm - green ash 94 Sycamore - sweetgum - American elm 95 Black willow 235 Cottonwood - willow SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Peachleaf willow is normally found in early seral riparian communities. It typically occurs scattered along the edge sof streams, rivers, ponds, or lakes. Published classification schemes listing peachleaf willow as an indicator or dominant in community types (cts), or dominance types (dts) are presented below: Area Classification Authority e CO: South Platte & Arkansas Rivers riparian cts Lindauer 1983 MT, WY: Bighorn Canyon Natl. Recr. Area general veg. cts Knight & others 1987 MT riparian dts Hansen & others 1988 sw US wetland cts Brown 1979

VALUE AND USE

SPECIES: Salix amygdaloides | Peachleaf Willow
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Willows (Salix spp.) in general are a preferred food of moose and beaver; peachleaf willow occurs in riparian and floodplain habitats which these animals frequent [6,42]. Stands in Montana typically have a good moisture regime throughout the growing season, and the understory associates provide quality forage for both livestock and wildlife [10]. PALATABILITY : In the West, willowsin general are considered to be more palatable to sheep than to catle, but cattle may make greater use of willow because they tend to frequent riparian areas [40]. The palatability of peachleaf willow has been rated as follows [10]: CO MT ND UT WY Cattle fair fair fair poor fair Sheep fair good good fair fair Horses fair fair fair poor fair NUTRITIONAL VALUE : Peachleaf willow's energy value has been rated as fair and its protein value as poor [10]. COVER VALUE : Peachleaf willow provides cover for numerous wildlife species. The degree to which it provides environmental protection during one or more seasons has been rated as follows [10]: CO ND UT WY Pronghorn good ---- ---- poor Elk ---- ---- fair fair Mule deer ---- ---- fair good White-tailed deer good good ---- good Small mammals ---- ---- fair good Small nongame birds good ---- good good Upland game birds ---- ---- good good Waterfowl ---- ---- ---- fair VALUE FOR REHABILITATION OF DISTURBED SITES : Cuttings of pachleaf willow can be used to revegetate disturbed riparian areas. Cuttings quickly stabilize disturbed alluvium, allowing other plants to become established. Unrooted willow stem cuttings (slips) should be planted on sites that provide sufficient moisture to start and maintain growth throughout the growing season [43]. Since willows are sensitive to both competition and shading, dense tall grasses will reduce transplant survival [31] and may need to be removed by cutting or by herbicide application [29]. Although harder to plant, rooted stock is recommended for use because it has higher survival rates [31,43]. A 20-inch (50 cm) cutting should be planted to a depth of 12 inches (30 cm), with 8 inches (20 cm) left aboveground [31]. This deep planting allows for more rooting surface to extract soil moisture and higher amounts of carbohydrates as stored food reserves [31,43]. Peachleaf willow cuttings root at the base of the cut; roots appear in about 10 to 20 days, and stems in about 10 days [31]. If serious streambank erosion has resulted in a nearly vertical cut bank, slope reshaping may be necessary to enhance success of transplants. Reshaping may not be necessary if, through protective measures, existing vegetation is able to stabilize the site [31,43]. With any method of revegetation, sites should be fenced to protect them from grazing and trampling. OTHER USES AND VALUES : All willows produce salicin, which chemically is closely related to acetylsalicylic acid, commonly known as aspirin. This is probably why Native Americans used various preparations from willows to treat toothache, stomach ache, diarrhea, dysentery, and dandruff [30]. Native Americans also used the stems for basketry and bow making, and the bark for tea and fabric making [24]. MANAGEMENT CONSIDERATIONS : Overuse by livestock can decrease vigor and eventually kill individuals; however, degraded stands recover rapidly after they are fenced to exclude livestock [34]. Because of its soil-binding properties, peachleaf willow helps stabilize streambanks and protect them from erosion; stands should therefore be maintained.

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Salix amygdaloides | Peachleaf Willow
GENERAL BOTANICAL CHARACTERISTICS : Peachleaf willow is a rapidly growing, short-lived, small- to medium-sized deciduous tree with one to several trunks that is typically from 20 to 40 feet (6-12 m) tall [12,16,20,42,44] but occasionally reaches 65 to 80 feet (20-24 m) [3,37]. Trunk diameters are typically 8 to 20 inches (20-50 cm) [44]. The bark is thick, yellowish brown to dark brown, irregular, fissured, and has broad flat ridges [14,37]. The leaves are alternate, simple, pinnately veined, lanceolate to ovate-lanceolate, with finely serrate margins which have 15 to 17 teeth per inch (6-7/cm) [16,20,12,37]. Leaves are 1 to 4 inches (2.5-10 cm) long and 0.4 to 1.2 inches (1-3 cm) wide [16,20,36,45]. Male and female flowers occur on separate trees as catkins. Pistillate catkins are 1 to 3 inches (3-8 cm) long, and staminate catkins are 1 to 2 inches (3-5 cm) long [14,16]. Due to their morphological similarities, peachleaf willow may be mistaken or confused with Pacific willow (Salix lasiandra), but they can be distinguished by differences in their bud scales [12] Peachleaf willow has bud scales with free overlapping margins and are pointed at the tip, while Pacific willow has bud scales without free overlapping margins and are rounded at the tip. RAUNKIAER LIFE FORM : Undisturbed State: Phanerophyte (mesophanerophyte) Undisturbed State: Phanerophyte (microphanerophyte) Undisturbed State: Phanerophyte (nanophanerophyte) Burned State: Hemicryptophyte REGENERATION PROCESSES : Peachleaf willow's regenerates primarily through the dispersal of thousands of small seeds. It is unable to produce suckers from lateral roots but will resprout from its root crown or stem base following fire or cutting [2,18,19]. Peachleaf willos It relies heavily on insect pollination, especially from bees [30]. After fertilization, a capsule develops which eventually splits open during spring or summer, dispersing the numerous tiny seeds (about 2,600,000 per pound [6,420,000/kg] [8]). Producing large quantities of seeds ensures that some will fall on favorable sites [3]. Seeds have a cottony down which allows them to float long distances in wind or on water. Seeds are non-dormant remaining viable for only a few days. They germinate rapidly, usually within 12 to 24 hours of dispersal if a moist seedbed is reached [8]. The seeds contain significant amounts of chlorophyll, and photosynthesis generally occurs as soon as the seed is moistened. Regeneration may also occur through broken pieces of stems or roots, which are transported and deposited by floodwaters, and later sprout [47]. SITE CHARACTERISTICS : Peachleaf willow is found along stream and river banks, pond and lake borders, moist ravines and ditches, oxbows, roadside gullies, and prairie sloughs [12,14,16,19,37]. It is shade intolerant and requires canopy openings to survive [42,44]. It is tolerant of poor drainage and prolonged flooding [19] but extended immersion in water for a year or longer will cause most plants to die [17]. Soils: Peachleaf willow occurs on a wide variety of soil textures but is most commonly found on sandy or silty alluvium [14,19,44]. It can tolerate weakly saline or alkaline soils [19]. Its growth on gravel or dense clay is poor, on clay is fair, and on sand, sandy-loam, loam, clay-loam is good [10]. Associates: Peachleaf willow is often found bordering water in riparian or floodplain forests that contain eastern cottonwood (Populus deltoides), green ash (Fraxinus pennsylvanica), boxelder (Acer negundo), American elm (Ulmus americana), Woods rose (Rosa woodsii), chokecherry (Prunus virginia), hackberry (Celtis occidentalis), and sandbar willow (Salix exigua) [4,19,33,39]. Elevation: Peachleaf willow is typically found along low elevation streams, rivers, and ponds in the Rocky Mountains [12,19]; along streambanks in the foothills and plains of dryland regions of the Pacific Northwest [20]; and along lowland floodplain forest of the Great Plains [35,37]. Elevational ranges for several western states are as follows [1,10,45]: from 3,500 to 9,500 feet (1,067-2,896 m) in CO from 3,500 to 5,600 feet (1,067-1,707 m) in UT from 3,500 to 7,000 feet (1,067-2,134 m) in WY SUCCESSIONAL STATUS : Peachleaf willow is an early successional species which pioneers floodplain alluvium [4,44]. In Kansas, it occurrs only as a pioneer on newly deposited alluvium; stands maintain themselves for about 30 years, until other riparian forest trees shade them out [4]. It is shade intolerant and can therefore persist along a river's edge, where repeated flooding prevents other species from becoming established. SEASONAL DEVELOPMENT : Peachleaf willow is a deciduous tree. Its catkins emerge with the leaves in the spring [16]. After fruits ripen, seeds are dispersed from spring to early summer [8]. The flowering times for several states are presented below [8,10]: State Flowering Begins Flowering Ends CO April June MN May June ND April May UT May June WY June July

FIRE ECOLOGY

SPECIES: Salix amygdaloides | Peachleaf Willow
FIRE ECOLOGY OR ADAPTATIONS : Peachleaf willow sprouts from its roots following fire [19]. The high soil and fuel moisture content chracteristic of its streamside habitat reduces the chance of fire ignition and spread. Its numerous wind-dispersed seeds are also important in revegetating areas following fire [47]. POSTFIRE REGENERATION STRATEGY : survivor species; on-site surviving root crown or caudex off-site colonizer; seed carried by wind; postfire years one and two off-site colonizer; seed carried by animals or water; postfire yr 1&2

FIRE EFFECTS

SPECIES: Salix amygdaloides | Peachleaf Willow
IMMEDIATE FIRE EFFECT ON PLANT : Most fires kill only aboveground plant parts. However, severe fires can completely remove soil organic layers, leaving willow roots exposed and charred, and thus eliminating basal sprouting [22,36,47]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Generally peachleaf willow will resprout from its roots following fire [19]. Peachleaf willow is a prolific seeder, and off-site plants are important seed sources for revegetating burned areas [47]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Due to their proximity to water or high water tables, peachleaf willow communities may serve as natural fire breaks [19].

REFERENCES

SPECIES: Salix amygdaloides | Peachleaf Willow
REFERENCES : 1. Argus, George W. 1957. The willows of Wyoming. University of Wyoming Publications. 21(1). Laramie, WY: University of Wyoming, Publications in Science. 63 p. [4962] 2. Argus, George W. 1973. The genus Salix in Alaska and the Yukon. Publications in Botany, No. 2. Ottowa, ON: National Museums of Canada, National Museum of Natural Sciences. 279 p. [6167] 3. Arno, Stephen F.; Hammerly, Ramona P. 1977. Northwest trees. Seattle, WA: The Mountaineers. 222 p. [4208] 4. Bellah, R. Glenn; Hulbert, Lloyd C. 1974. Forest succession on the Republican River floodplain in Clay County, Kansas. Southwestern Naturalist. 19(2): 155-166. [241] 5. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 6. Boyd, Raymond J.; Cooperrider, Allen Y.; Lent, Peter C.; Bailey, James A. 1986. Ungulates. In: Cooperrider, Allen Y.; Boyd, Raymond J.; Stuart, Hanson R., eds. Inventory and monitoring of wildlife habitat. Denver, CO: U.S. Department of the Interior, Bureau of Land Management, Service Center: 519-564. [10856] 7. Brayshaw, T. Christopher. 1976. Catkin bearing plants of British Columbia. Occas. Pap. No. 18. Victoria, BC: The British Columbia Provincial Museum. 176 p. [6170] 8. Brinkman, Kenneth A. 1974. Salix L. willow. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 746-750. [5412] 9. Brown, David E. 1979. Southwestern wetlands - their classification and characteristics. In: Johnson, R. Roy; McCormick, J. Frank, technical coordinators. Strategies for protection and management of floodplain wetlands & other riparian ecosystems: Proc. of the symposium; 1978 December 11-13; Callaway Gardens, GA. Gen. Tech. Rep. WO-12. Washington, DC: U.S. Department of Agriculture, Forest Service: 269-282. [4366] 10. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 11. Dorn, Robert D. 1970. The willows of Montana. Bozeman, MT: Montana State University, Department of Botany and Microbiology. 18 p. [6176] 12. Dorn, Robert D. 1977. Willows of the Rocky Mountain States. Rhodora. 79: 390-429. [6000] 13. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 14. Froiland, Sven G. 1962. The genus Salix (Willows) in the Black Hills of South Dakota. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 75 p. [5992] 15. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 16. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 17. Green, William E. 1947. Effect of water impoundment on tree mortality and growth. Journal of Forestry. 45(2): 118-120. [3718] 18. Haeussler, S.; Coates, D. 1986. Autecological characteristics of selected species that compete with conifers in British Columbia: a literature review. Land Management Report No. 33. Victoria, BC: Ministry of Forests, Information Services Branch. 180 p. [1055] 19. Hansen, Paul L.; Chadde, Steve W.; Pfister, Robert D. 1988. Riparian dominance types of Montana. Misc. Publ. No. 49. Missoula, MT: University of Montana, School of Forestry, Montana Forest and Conservation Experiment Station. 411 p. [5660] 20. Hitchcock, C. Leo; Cronquist, Arthur. 1964. Vascular plants of the Pacific Northwest. Part 2: Salicaceae to Saxifragaceae. Seattle, WA: University of Washington Press. 597 p. [1166] 21. Knight, Dennis H.; Jones, George P.; Akashi, Yoshiko; Myers, Richard W. 1987. Vegetation ecology in the Bighorn Canyon National Recreation Area: Wyoming and Montana. Final Report. Laramie, WY: University of Wyoming, National Park Service Research Center. 114 p. [12498] 22. Kovalchik, Bernard L. 1987. Riparian zone associations: Deschutes, Ochoco, Fremont, and Winema National Forests. R6 ECOL TP-279-87. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 171 p. [9632] 23. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 24. Lanner, Ronald M. 1983. Trees of the Great Basin: A natural history. Reno, NV: University of Nevada Press. 215 p. [1401] 25. Lindauer, Ivo E. 1983. A comparison of the plant communities of the South Platte and Arkansas River drainages in eastern Colorado. Southwestern Naturalist. 28(3): 249-259. [5886] 26. Little, Elbert L., Jr. 1971. Atlas of the United States trees. Volume 1. Conifers and important hardwoods. Misc. Publ. 1146. Washington, DC: U.S. Department of Agriculture, Forest Service. 320 p. [1462] 27. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 28. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 29. Monsen, Stephen B. 1983. Plants for revegetation of riparian sites within the Intermountain region. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings of symposia; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 83-89. [9652] 30. Mozingo, Hugh N. 1987. Shrubs of the Great Basin: A natural history. Reno, NV: University of Nevada Press. 342 p. [1702] 31. Platts, William S.; Armour, Carl; Booth, Gordon D.; [and others]. 1987. Methods for evaluating riparian habitats with applications to management. Gen. Tech. Rep. INT-221. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 177 p. [6171] 32. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 33. Read, D. J. 1983. The biology of mycorrhiza in the Ericales. Canadian Journal of Botany. 61: 985-1004. [10602] 34. Rickard, W. H.; Cushing, C. E. 1982. Recovery of streamside woody vegetation after exclusion of livestock grazing. Journal of Range Management. 35(3): 360-361. [5888] 35. Rothenberger, Steven J. 1985. Community analysis of the forest vegetation in the lower Platte River Valley, eastern Nebraska. Prairie Naturalist. 17(1): 1-14. [2031] 36. Rowe, J. S.; Scotter, G. W. 1973. Fire in the boreal forest. Quaternary Research. 3: 444-464. [72] 37. Stephens, H. A. 1973. Woody plants of the North Central Plains. Lawrence, KS: The University Press of Kansas. 530 p. [3804] 38. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 39. Tolstead, W. L. 1941. Plant communities and secondary succession in south-central South Dakota. Ecology. 22(3): 322-328. [5887] 40. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. [2387] 41. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 42. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 43. Ward, Don; Thompson, Robert; Kelly, Dennis. 1986. Willow planting guide. R-4 Hydrograph No. 54. Ogden, UT: U.S. Department of Agriculture, Forest Service, Range and Watershed Management. 12 p. [2936] 44. Weaver, J. E. 1960. Flood plain vegetation of the central Missouri Valley and contacts of woodland with prairie. Ecological Monographs. 30(1): 37-64. [275] 45. Welsh, Stanley L.; Atwood, N. Duane; Goodrich, Sherel; Higgins, Larry C., eds. 1987. A Utah flora. Great Basin Naturalist Memoir No. 9. Provo, UT: Brigham Young University. 894 p. [2944] 46. White, W. W. 1956. Native willows found in Montana. Proceedings of the Montana Academy of Science. 16: 21-35. [6001] 47. Zasada, J. 1986. Natural regeneration of trees and tall shrubs on forest sites in interior Alaska. In: Van Cleve, K.; Chapin, F. S., III; Flanagan, P. W.; [and others], eds. Forest ecosystems in the Alaska taiga: A synthesis of structure and function. New York: Springer-Verlag: 44-73. [2291]

Index

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