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Introductory

WILDLIFE SPECIES: Accipiter gentilis | Northern Goshawk
ABBREVIATION : ACGE COMMON NAMES : northern goshawk goshawk American goshawk bluehen hawk blue darter partridge hawk dove hawk chicken hawk Apache northern goshawk TAXONOMY : The currently accepted scientific name for the northern goshawk is Accipiter gentilis (Linnaeus). There are two recognized subspecies in North America: A. g. ssp. atricapillus (Wilson) and A. g. ssp. laingi (Taverner) [1]. Accipiter genitilis ssp. apache (Rossem), formerly recognized as a distinct subspecies, has been included in the taxon A. g. ssp. atricapillus by the American Ornithologist's Union [16]. ORDER : Falconiformes CLASS : Bird FEDERAL LEGAL STATUS : Notice of Review, Category 2 OTHER STATUS : Region 3 of the U.S. Forest Service has listed northern goshawk as a sensitive species [7]. South Dakota has listed it as an uncommon species that merits monitoring [9]. COMPILED BY AND DATE : Randy Scott Griffith, July 1993. LAST REVISED BY AND DATE : AUTHORSHIP AND CITATION : Griffith, Randy S. 1993. Accipiter gentilis. In: Remainder of Citation

WILDLIFE DISTRIBUTION AND OCCURRENCE

WILDLIFE SPECIES: Accipiter gentilis | Northern Goshawk
GENERAL DISTRIBUTION : The range of the northern goshawk is circumpolar [11]. Its year-round range in North America extends from northern Alaska and Canada south to northern Mexico, Colorado, Minnesota, and western Maryland [1,19]. Winter sightings of northern goshawk have been recorded in Missouri, Tennessee, West Virginia, Virginia, and Florida [1]. Accipiter g. ssp. atricapillus occupies most of the above range. Accipiter g. ssp. laingi occupies a limited range on Masset, Queen Charlotte, and Vancover islands, British Columbia [1]. ECOSYSTEMS : FRES10 White-red-jack pine FRES11 Spruce-fir FRES15 Oak-hickory FRES17 Elm-ash-cottonwood FRES18 Maple-beech-birch FRES19 Aspen-birch FRES20 Douglas-fir FRES21 Ponderosa pine FRES22 Western white pine FRES23 Fir-spruce FRES24 Hemlock-Sitka spruce FRES25 Larch FRES27 Redwood FRES28 Western hardwoods STATES :
AK AZ CA CO CT FL ID
IL IN IA KS KY ME MA
MI MN MO MT NC NE NV
NH NJ NM NY ND OH OK
OR PA RI SD TN TX UT
VT VA WA WV WI WY

AB BC MB NB NF NT NS ON PE PQ
SK YT

MEXICO
BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K002 Cedar - hemlock - Douglas-fir forest K003 Silver fir - Douglas-fir forest K004 Fir - hemlock forest K005 Mixed conifer forest K006 Redwood forest K007 Red fir forest K010 Ponderosa shrub forest K011 Western ponderosa forest K012 Douglas-fir forest K013 Cedar - hemlock - pine forest K014 Grand fir - Douglas-fir forest K015 Western spruce - fir forest K016 Eastern ponderosa forest K017 Black Hills pine forest K018 Pine - Douglas-fir forest K019 Arizona pine forest K020 Spruce - fir - Douglas-fir forest K021 Southwestern spruce - fir forest K025 Alder - ash forest K026 Oregon oakwoods K028 Mosaic of K002 and K026 K081 Oak savanna K082 Mosaic of K074 and K100 K084 Cross Timbers K093 Great Lakes spruce - fir forest K094 Conifer bog K095 Great Lakes pine forest K096 Northeastern spruce - fir forest K097 Southeastern spruce - fir forest K098 Northern floodplain forest K099 Maple - basswood forest K100 Oak - hickory forest K101 Elm - ash forest K102 Beech - maple forest K103 Mixed mesophytic forest K106 Northern hardwoods K107 Northern hardwoods - fir forest K108 Northern hardwoods - spruce forest SAF COVER TYPES : 1 Jack pine 14 Northern pin oak 16 Aspen 17 Pin cherry 18 Paper birch 24 Hemlock - yellow birch 25 Sugar maple - beech - yellow birch 26 Sugar maple - basswood 27 Sugar maple 28 Black cherry - maple 39 Black ash - American elm - red maple 42 Bur oak 43 Bear oak 44 Chestnut oak 46 Eastern redcedar 52 White oak - black oak - northern red oak 53 White oak 55 Northern red oak 57 Yellow-poplar 59 Yellow-poplar - white oak - northern red oak 60 Beech - sugar maple 62 Silver maple - American elm 201 White spruce 202 White spruce - paper birch 203 Balsam poplar 204 Black spruce 205 Mountain hemlock 206 Engelmann spruce - subalpine fir 207 Red fir 210 Interior Douglas-fir 211 White fir 212 Western larch 213 Grand fir 215 Western white pine 216 Blue spruce 217 Aspen 221 Red alder 221 Red alder 222 Black cottonwood - willow 223 Sitka spruce 224 Western hemlock 225 Western hemlock - Sitka spruce 226 Coastal true fir - hemlock 227 Western redcedar - western hemlock 228 Western redcedar 230 Douglas-fir - western hemlock 231 Port-Orford-cedar 232 Redwood 233 Oregon white oak 234 Douglas-fir - tanoak - Pacific madrone 235 Cottonwood - willow 236 Bur oak 237 Interior ponderosa pine 238 Western juniper 243 Sierra Nevada mixed conifer 244 Pacific ponderosa pine - Douglas-fir 245 Pacific ponderosa pine 246 California black oak 247 Jeffrey pine 250 Blue oak - Digger pine 251 White spruce - aspen 252 Paper birch 253 Black spruce - white spruce 254 Black spruce - paper birch 256 California mixed subalpine SRM (RANGELAND) COVER TYPES : NO-ENTRY PLANT COMMUNITIES : The northern goshawk is a forest dweller. In the East, it usually selects coniferous forests of pine (Pinus spp.), spruce (Picea spp.), and/or fir (Abies spp.) with scattered patches of willow (Salix spp.), alder (Alnus spp.), or other shrub species. However, its habitat is not limited to coniferous forests. It also selects stands of birch (Betula spp.), beech (Fagus spp.), and quaking aspen (Populus tremuloides) in deciduous forests [16]. In the West, the northern goshawk often dwells in ponderosa pine (Pinus ponderosa), mixed-species, and spruce-fir forests. Douglas-fir (Pseudotsuga menziesii), white fir (Abies concolor), and grand fir (A. grandis) frequently dominate the mixed-species forests. Dominants in spruce-fir forests include Engelmann spruce (Picea engelmannii) and subalpine fir (A. lasiocarpa) [18]. REFERENCES : NO-ENTRY

BIOLOGICAL DATA AND HABITAT REQUIREMENTS

WILDLIFE SPECIES: Accipiter gentilis | Northern Goshawk
TIMING OF MAJOR LIFE HISTORY EVENTS : Age at first reproduction: Some yearling females breed with older males; yearling males are typically incapable of breeding [16]. Generally, 25 percent of northern goshawks breed as yearlings, another 25 percent in their second year, and the remainder in their third year [5]. Pair formation: Northern goshawks are monogomous. If a mate is lost the northern goshawk will form another pair bond [16]. Nesting: The northern goshawk selects a preexisting nest or builds a platform nest [11]. Nesting occurs from early April to mid-June, depending on latitude [16]. Clutch/incubation and fledging: The clutch size of the northern goshawk varies from one to five eggs, with a mean of three eggs per clutch. One egg is laid every 2 days. If a nest is destroyed early in the breeding season, the northern goshawk will nest again. Incubation period is 32 to 34 days. The young fledge in 37 to 41 days with the smaller, faster-developing males fledging a few days before the females [16]. Migration: Winter migration begins in late August to early September. Spring migration begins in early Feburary and is usually complete by mid-March. Many birds, however, remain in their territories year-round, only moving when prey is limited [16]. Molting: The female northern goshawk begins molting at the start of incubation, and pauses when the young are about ready to fledge. The male molts during this pause. The female resumes molting when the young are flying and hunting on their own [16]. PREFERRED HABITAT : The northern goshawk uses a wide variety of forest ages, structural conditions, and successional stages [17,18]. For hunting habitat, the northern goshawk prefers the transitional zones from bog to forest and forest to shrubland. Riparian zones and mosaics of forested and open areas are also important hunting habitats [4,16]. The northern goshawk uses stands of old-growth forest as nesting sites [4]. COVER REQUIREMENTS : The home range size for a pair of northern goshawks is about 6,000 acres (2,400 ha). Within the home range there are three areas of use: the nest area, the post-fledgling family area, and the foraging area. Greater than 20 percent of the home range is old-growth forest [17,18,20]. The nest area is about 30 acres (12 ha) in size, and it is usually located on north aspects in drainages or canyons with streams [17,18]. The nest area is usually in a mature forest stand that has a multilayered canopy with dense to open understory [12]. Within a home range there are usually two to four alternative nest areas [17,18]. The tree selected for nesting has a crotch, fork, or several limbs on one side to support the platform nest [16]. The post-fledgling family area is about 420 acres (168 ha). This area is a mosaic of forest types that provide hiding cover for the fledglings and habitat for abundant prey [17,18]. The foraging area is about 5,400 acres (2,160 ha). This area consists of a mosaic of shrublands, forests, and openings with perching trees to observe prey [16,17,18]. FOOD HABITS : The northern goshawk selects two or three species of the prey species available within its hunting territory. It searches the area from the forest floor to the lower canopy from a perch. Once prey is observed the northern goshawk will take flight and pursue its prey. Once pursuit is started and the cover becomes too thick for flight, the northern goshawk will pursue prey on foot. The diet of the northern goshawk changes with the season. In the spring and summer, it is mainly birds, with a few small mammals. In the winter, the diet consists of prey species that do not migrate of hibernate. In northern portions of their range, northern goshawk prey upon ptarmigans (Lagopus spp.) in the winter. In southern portions of their range, they consume ruffed grouse (Bonasa umbellus), blue grouse (Dendragapus obscurus), hares (Lepus spp.), and red squirrel (Tamiasciurus hudsonicus) [16]. The wide distribution of the northern goshawk results in a varied prey base. Stomach and pellet contents show the following prey animals: robin (Turdus migratorius), Stellar's jay (Cyanocitta stelleri), mallard (Anas platyrhynchos), black duck (Anas rubripes), sparrows (Ploceidae), warblers (Parulidae), kestrel (Falco tinnunculus), crows (Corvus spp.), hares, cottontails (Sylvilagus spp.), chipmunks (Eutamias spp.), ground squirrels (Spermophilus spp.), woodchuck (Marmota monax), muskrat (Ondantra zibethicus), mice (Heteromyidae), and shrews (Soricidae spp.) [16]. PREDATORS : The northern goshawk is fairly high in the food web; its predators are large avians such as the bald eagle (Haliaeetus leucocephalus) and golden eagle (Aquila chrysaetos) [3]. MANAGEMENT CONSIDERATIONS : Timber harvest can be part of a management plan to perpetuate northern goshawk habitat providing the plan calls for retention of a minimum of 20 percent old growth in the northern goshhawk's home range. Rotation time for old-growth stands should be 150 years or more [17,18,20]. Reynolds and others [18] provided detailed information on the types of management practices that enhance northern goshawk habitat. During harvest, the snags within the harvest area should be retained. When hunting, the northern goshawk usually observes its prey while perched in a snag, and then pursues the prey animal. Few pursuits are started while already in flight [2,16,18]. When using spruce-fir forests as grazing areas, recommended utilization of herbaceous cover is 20 percent. Heavy grazing in these forests reduces herbaceous ground cover, resulting in dense stands of trees. This reduces the quality of the northern goshawk's habitat by reducing populations of its prey species [18]. Forest management activities should not take place in nesting areas during the nesting period [10,18]. Roads in the nesting area should be kept at a minimum and should be closed during nesting. Use of skid trails is preferable to construction of permanent roads [18]. REFERENCES : NO-ENTRY

FIRE EFFECTS AND USE

WILDLIFE SPECIES: Accipiter gentilis | Northern Goshawk
DIRECT FIRE EFFECTS ON ANIMALS : Adult birds rarely are killed by fire [14]. Fires in the early spring, before fledging, may result in mortality of juveniles. HABITAT RELATED FIRE EFFECTS : Large, stand-replacing fires can reduce northern goshawk recruitment by destroying nesting and foraging areas [14]. Fire suppression in ponderosa pine, mixed species, and spruce-fir forests has resulted in dense, single-story forests with few openings. Fuel accumulation may result in stand-replacing fires. Large, stand-replacing fires can destroy large blocks of northern goshawk habitat, resulting in an uniform habitat instead of a mosiac of open and forested areas[17,18]. FIRE USE : Fire can be beneficial to the northern goshawk by perpetuating forest seres, which provide habitat for its prey. The fire-maintained game preserves in the Southwest provide habitat for quail and other passerines on which the northern goshawk preys [14]. Reynolds and others [17] recommended prescribed fire in ponderosa pine and mixed-species forests to perpetuate northern goshawk habitat and to reduce fuel loading. REFERENCES : NO-ENTRY

REFERENCES

WILDLIFE SPECIES: Accipiter gentilis | Northern Goshawk
REFERENCES : 1. American Ornithologists' Union. 1957. Checklist of North American birds. 5th ed. Baltimore, MD: The Lord Baltimore Press, Inc. 691 p. [21235] 2. Benson, Patrick C. 1979. Land use and wildlife with emphasis on raptors. [Ogden, UT]: U.S. Department of Agriculture, Forest Service, Intermountain Region. 32 p. On file with: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT. [17208] 3. Bosakowski, T.; Smith, D. G.; Speiser, R. 1992. Niche overlap of two sympatric-nesting hawks Accipiter spp. in the New Jersey--New York Highlands. Ecography. 15(4): 358-372. [19978] 4. DuBois, Kristi; Becker, Dale; Thornbrugh, Joe. 1987. Identification of Montana's birds of prey. Montana Outdoors. 18(6): 11-31. [3606] 5. Ehrlich, Paul R.; Dobkin, David S.; Wheye, Darryl. 1988. The birder's handbook: a field guide to the natural history of North American birds. New York: Simon & Schuster, Inc. 785 p. [21559] 6. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 7. Finch, Deborah M. 1992. Threatened, endangered, and vulnerable species of terrestrial vertebrates in the Rocky Mountain Region. Gen. Tech. Rep. RM-215. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 38 p. [18440] 8. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 9. Houtcooper, Wayne C.; Ode, David J.; Pearson, John A.; Vandell, George M., III. 1985. Rare animals and plants of South Dakota. Prairie Naturalist. 17(3): 143-165. [17792] 10. Kennedy, Patricia L.; Stahlecker, Dale W. 1993. Responsiveness of nesting northern goshawks to taped broadcasts of 3 conspecific calls. Journal of Wildlife Management. 57(2): 249-257. [21463] 11. Knopf, A. A. 1977. The Audubon Society field guide to North American birds, western region. New York: Chanticleer Press. 852 p. [21560] 12. Kochert, Michael N. 1986. Raptors. In: Cooperrider, Allan Y.; Boyd, Raymond J.; Stuart, Hanson R., eds. Inventory and monitoring of wildlife habitat. Denver, CO: U.S. Department of the Interior, Bureau of Land Management, Denver Service Center: 313-349. [13527] 13. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 14. Landers, J. Larry. 1987. Prescribed burning for managing wildlife in southeastern pine forests. In: Dickson, James G.; Maughan, O. Eugene, eds. Managing southern forests for wildlife and fish: a proceedings; [Date of conference unknown]; [Location of conference unknown]. Gen. Tech. Rep. SO-65. New Orleans, LA: U.S. Department of Agriculture, Forest Service, Southern Forest Experiment Station: 19-27. [11562] 15. Moseley, Robert; Groves, Craig, compilers. 1990. Rare, threatened and endangered plants and animals of Idaho. Boise, ID: Idaho Department of Fish and Game, Nongame and Endangered Wildlife Program, Natural Heritage Section. 33 p. [19328] 16. Palmer, R. S., ed. 1988. Handbook of North American birds. New York: Yale University Press. 5 vol p. [21561] 17. Reynolds, Richard T.; Graham, Russell T.; Reiser, M. Hildegard; [and others]. 1992. Management recommendations for the northern goshawk in the southwestern United States. In: Proceedings, 10th Intermountain Region silvicultural workshop; 1991 December 3-5; Ogden, UT. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Region: 54-59. [21324] 18. Reynolds, Richard T.; Graham, Russell T.; Reiser, M. Hildegard; [and others]. 1992. Management recommendations for the northern goshawk in the southwestern United States. Gen. Tech. Rep. RM-217. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 90 p. [19721] 19. Scott, S. L. 1987. National Geographic Society field guide to the birds of North America. 2d ed. Washington D. C: National Geographic Society. 464 p. [21562] 20. St. Clair, Jeffrey. 1992. Goshawk guidelines threaten old-growth. Forest Watch. 13(3): 10-11. [20793] 21. Tate, James, Jr. 1986. The Blue List for 1986. American Birds. 40(2): 227-235. [24324] 22. U.S. Department of the Interior, Fish and Wildlife Service. 1994. Endangered and threatened wildlife and plants; animal candidate review for listing as endangered or threatened species; proposed rule. 50 CFR Part 17. Tuesday, November 15, 1994. Federal Register. 59(219): 58982-59028. [24357]

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