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Introductory

SPECIES: Dalea purpurea | Purple Prairie Clover
ABBREVIATION : DALPUR SYNONYMS : Petalostemon purpureus (Vent.) Rydb. SCS PLANT CODE : DAPU5 COMMON NAMES : purple prairie clover TAXONOMY : The currently accepted scientific name for purple prairie clover is Dalea purpurea Vent. Formerly, the species was regarded as Petalostemon purpureus, but a 1970 taxonomic revision of the genus Petalostemon placed purple prairie clover in the genus Dalea. Recognized varieties are as follows [14]: D. purpurea var. purpurea D. purpurea var. arenicola (Wemple) Barneby LIFE FORM : Forb FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Milo Coladonato, February 1993 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Coladonato, Milo. Dalea purpurea. 1993. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Dalea purpurea | Purple Prairie Clover
GENERAL DISTRIBUTION : Purple prairie clover is distributed throughout the Great Plains. It is indigenous from southeastern Alberta, southern Saskatchewan, and Manitoba south to Texas and New Mexico and east to Illinois, Arkansas, and Missouri. It occurs infrequently westward to eastern Montana, Colorado, and Wyoming [14,18,30]. ECOSYSTEMS : FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES36 Mountain grasslands FRES38 Plains grasslands FRES39 Prairie STATES : AR CO IL IN IA KS MN MS MO MT NE NM ND OK SD TN TX WY AB MB SK ADMINISTRATIVE UNITS : BADL BICA BUFF CACA DETO EFMO FLFO GUMO INDU JECA NATR OZAR PIPE ROMO THRO WICA BLM PHYSIOGRAPHIC REGIONS : 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K063 Foothills prairie K064 Grama - needlegrass - wheatgrass K065 Grama - buffalograss K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem - needlegrass K068 Wheatgrass - grama - buffalograss K069 Bluestem - grama prairie K074 Bluestem prairie K075 Nebraska Sandhills prairie K082 Mosaic of K074 and K100 K084 Cross Timbers K098 Northern floodplain forest SAF COVER TYPES : 14 Northern pin oak 40 Post oak - blackjack oak 42 Bur oak 46 Eastern redcedar 53 White oak 62 Silver maple - American elm 64 Sassafras - persimmon 93 Sugarberry - American elm - green ash 94 Sycamore - sweetgum - American elm 95 Black willow 110 Black oak 242 Mesquite SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : NO-ENTRY

VALUE AND USE

SPECIES: Dalea purpurea | Purple Prairie Clover
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Purple prairie clover is excellent forage for livestock and wildlife and may be an important component of upland prairie hay [30,31]. PALATABILITY : Purple prairie clover is palatable to livestock as well as to white-tailed deer and wild turkey [6]. Palatability of purple prairie clover in two western states has been rated as follows [8]: CO ND cattle good fair sheep good good horses fair fair NUTRITIONAL VALUE : Purple prairie clover is highly nutritious. It has a high protein content, particularly in new growth [6]. Crude and digestible protein content of purple prairie clover are as follows [21]: crude protein 14.1 % digestible protein - cattle 9.9 % goats 9.7 % horses 9.5 % rabbits 9.6 % sheep 10.1 % COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Purple prairie clover is used in grass and forb seed mixtures for stabilization of disturbed sites such as mine spoils, road cuts, and overgrazed rangelands [4,27,28]. It is valuable for erosion control due to its deep, fibrous, branching root system. It is often seeded with grass mixtures because of its nitrogen-fixing ability [5]. OTHER USES AND VALUES : American Indians ate fresh and boiled purple prairie clover leaves. Bruised leaves were steeped in water and applied to open wounds. The Ponca chewed the leaves for their flavor and made tea from the leaves, and the Pawnee used the stems to make brooms [12,31]. MANAGEMENT CONSIDERATIONS : Purple prairie clover decreases in response to grazing [4,26]. The terminal growing points are within reach of the grazing animals soon after growth begins and are thus quite susceptible to injury by grazing or mowing [9]. Survival and growth of purple prairie clover seedlings is decreased in the presence of allelopathic species such as western ragweed (Ambrosia psilostachya), common lambsquarters (Chenopodium album), and smooth brome (Bromus inermis) [24].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Dalea purpurea | Purple Prairie Clover
GENERAL BOTANICAL CHARACTERISTICS : Purple prairie clover is a perennial forb 8 to 35 inches (20-90 cm) tall. Usually there are one to three branches per stem, but sometimes 10 to 12 branches may arise from a single stem. The mature purple prairie clover develops a strong taproot 5.5 to 6.5 feet (1.7-2.0 m) deep. The taproot gives rise to several minutely branched lateral roots. The fruit is a one-seeded pod enclosed in bracts [1,5,14,28]. RAUNKIAER LIFE FORM : Hemicryptophyte REGENERATION PROCESSES : Purple prairie clover reproduces by seed [7,15]. Good seed crops are produced about every 2 years, with some seeds produced every year [3]. Bjugstad and Whitman [3] reported that a temperature of 41 degrees Fahrenheit (5 deg C) is required to break seed dormancy. Documentation of this species' ability to sprout is scant. Nuzzo [22], however, reports that purple prairie clover seedlings that appeared dead due to water loss sent up new shoots within a few weeks when watered heavily. SITE CHARACTERISTICS : Purple prairie clover grows on a variety of sites including dry plains, prairies, hillsides, open woodlands, shaded ravines, and roadsides. It occurs on mesic sites in mixed and tallgrass prairies of the northern and central Great Plains and the shortgrass prairie of the southern Great Plains [4,14,23]. Where mean annual precipitation is less than 15 inches (380 mm) purple prairie clover occurs infrequently. It develops best where mean annual precipitation is 15 inches or more [33]. Some common associates of purple prairie clover include prairie gentian (Gentiana puberula), prairie compass plant (Silphium laciniatum), leadplant (Amorpha canescens), pitcher sage (Salvia azurea), blue false indigio (Baptisia minor), tall gray feather (Liatris scariosa), and yellow swetclover (Melilotus officinalis). Associated grasses include little bluestem (Schizachynum scoparium) switch grass (Pancium virgatum), Indian grass (Sorghastrum nutans), and big bluestem (Andropogon geradii) [16,36]. SUCCESSIONAL STATUS : Obligate Climax Species Purple prairie clover is part of the mixed prairie climax vegetation of several grassland communities [39]. It is moderately shade tolerant but can also grow on recently disturbed open areas [27,34]. SEASONAL DEVELOPMENT : Purple prairie clover begins vegetative growth at the same time as prairie grasses early in the spring, but purple prairie clover soon exceeds the grasses in height [37,38]. Purple prairie clover begins to flower in late May to July when the plant is about 2 feet (60 cm) high. Seeds mature in July and August [17,18,36].

FIRE ECOLOGY

SPECIES: Dalea purpurea | Purple Prairie Clover
FIRE ECOLOGY OR ADAPTATIONS : Purple prairie clover is generally favored by fire. Purple prairie clover is usually present in increased numbers following fire because fire stimulates the germination of soil-stored seed [2,7,32,33,36]. POSTFIRE REGENERATION STRATEGY : Ground residual colonizer (on-site, initial community) Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Dalea purpurea | Purple Prairie Clover
IMMEDIATE FIRE EFFECT ON PLANT : The aerial portions of purple prairie clover are damaged or killed by fire. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Purple prairie clover is generally considered an increaser following fire [2,13,23]. Fire stimulates seed germination; sprouting from the caudex is also possible [3,36]. Data pertaining to postfire density, frequency, or growth rates of purple prairie clover were not found in the literature. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : Biennial spring burning favors purple prairie clover, while annual burning favors prairie grasses and results in a decrease in purple prairie clover and other forbs [33].

REFERENCES

SPECIES: Dalea purpurea | Purple Prairie Clover
REFERENCES : 1. Bare, Janet E. 1979. Wildflowers and weeds of Kansas. Lawrence, KS: The Regents Press of Kansas. 509 p. [3801] 2. Bidwell, Terrence G.; Engle, David M.; Claypool, P. Larry. 1990. Effects of spring headfires and backfires on tallgrass prairie. Journal of Range Management. 43(3): 209-212. [11141] 3. Bjugstad, Ardell J.; Whitman, Warren C. 1982. Perennial forbs for wildlife habitat restoration on mined lands in the northern Great Plains. In: Western proceedings, 62nd annual conference of the Western Association of Fish and Wildlife Agencies; 1982 July 19-22; Las Vegas, Nevada. [Place of publication unknown]: [Publisher unknown]: 257-271. On file with: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Lab, Missoula, MT. [2932] 4. Brown, Lauren. 1985. The Audobon Society nature guides: Grasslands. New York: Alfred A. Knopf, Inc. 606 p. [4561] 5. Burton, Joseph C. 1972. Nodulation and symbiotic nitrogen fixation by prairie legumes. In: Zimmerman, James H., ed. Proceedings, 2nd Midwest prairie conference; 1970 September 18-20; Madison, WI. Madison, WI: University of Wisconsin Arboretum: 116-121. [2909] 6. Crawford, Hewlette S.; Kucera, Clair L.; Ehrenreich, John H. 1969. Ozark range and wildlife plants. Agric. Handb. 356. Washington, DC: U.S. Department of Agriculture, Forest Service. 236 p. [18602] 7. Cull, Margaret Irene. 1978. Establishing prairie vegetation along highways in the Peoria area. In: Glenn-Lewin, David C.; Landers, Roger Q., Jr., eds. Proceedings, 5th Midwest prairie conference; 1976 August 22-24; Ames, IA. Ames, IA: Iowa State University: 172-177. [3378] 8. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 9. Ehrenreich, John H.; Aikman, John M. 1963. An ecological study of the effect on certain management practices on native prairie in Iowa. Ecological Monographs. 33(2): 113-130. [9] 10. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 11. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 12. Gilmore, Melvin Randolph. 1919. Uses of plants by the Indians of the Missouri River region. 33rd Annual Report. Washington, DC: Bureau of American Ethnology. 154 p. [6928] 13. Glenn-Lewin, David C.; Johnson, Louise A.; Jurik, Thomas W.; [and others]. 1990. Fire in central North American grasslands: vegetative reproduction, seed germination, and seedling establishment. In: Collins, Scott L.; Wallace, Linda L., eds. Fire in North American tallgrass prairies. Norman, OK: University of Oklahoma Press: 28-45. [14194] 14. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 15. Halinar, Marlene. 1981. Germination studies and purity determinations on native Wisconsin prairie seeds. In: Stuckey, Ronald L.; Reese, Karen J., eds. The Prairie Peninsula--in the "shadow" of Transeau: Proceedings, 6th North American prairie conference; 1978 August 12-17; Columbus, OH. Ohio Biological Survey Biological Notes No. 15. Columbus, OH: Ohio State University, College of Biological Sciences: 227-231. [3433] 16. Harrington, George; Capel, Stephen. 1978. Effects of a water resource development project on native prairie in the Flint Hills of Kansas. In: Glenn-Lewin, David C.; Landers, Roger Q., Jr., eds. Proceedings, 5th Midwest prairie conference; 1976 August 22-24; Ames, IA. Ames, IA: Iowa State University: 166-168. [3371] 17. Hurd, Richard M.; Christisen, Donald M. 1975. Ecology study of Friendly Prairie, Missouri. In: Wali, Mohan K., ed. Prairie: a multiple view. Grand Forks, ND: University of North Dakota Press: 89-102. [4432] 18. Johnson, James R.; Nichols, James T. 1970. Plants of South Dakota grasslands: A photographic study. Bull. 566. Brookings, SD: South Dakota State University, Agricultural Experiment Station. 163 p. [18501] 19. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 20. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 21. National Academy of Sciences. 1971. Atlas of nutritional data on United States and Canadian feeds. Washington, DC: National Academy of Sciences. 772 p. [1731] 22. Nuzzo, Victoria. 1978. Propagation and planting of prairie forbs and grasses in southern Wisconsin. In: Glenn-Lewin, David C.; Landers, Roger Q., Jr., eds. Proceedings, 5th Midwest prairie conference; 1976 August 22-24; Ames, IA. Ames, IA: Iowa State University: 182-189. [3379] 23. Pemble, R. H.; Van Amburg, G. L.; Mattson, Lyle. 1981. Intraspecific variation in flowering activity following a spring burn on a northwestern Minnesota prairie. In: Stuckey, Ronald L.; Reese, Karen J., eds. The prairie peninsula--in the "shadow" of Transeau: Proceedings, 6th North American prairie conference; 1978 August 12-17; Columbus, OH. Ohio Biological Survey: Biological Notes No. 15. Columbus, OH: Ohio State University, College of Biological Sciences: 235-240. [3435] 24. Peterson, Gary. 1981. Allelopathic associations and their effect on native legumes. In: Stuckey, Ronald L.; Reese, Karen J., eds. The Prairie Peninsula--in the "shadow" of Transeau: Proceedings of the sixth North American prairie conference; 1978 August 12-17; Columbus, OH. Ohio Biological Survey Biological Notes No. 15. Columbus, OH: Ohio State University, College of Biological Sciences: 211-212. [3409] 25. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 26. Risser, P. G.; Birney, E. C.; Blocker, H. D.; [and others]. 1981. The true prairie ecosystem. US/IBP Synthesis Series 16. Stroudsburg, PA: Hutchinson Ross Publishing Company. 557 p. [16874] 27. Schramm, Peter; Kalvin, Richard L. 1978. The use of prairie in strip mine reclamation. In: Glenn-Lewin, David C.; Landers, Roger Q., Jr., eds. Proceedings, 5th Midwest prairie conference; 1976 August 22-24; Ames, IA. Ames, IA: Iowa State University: 151-153. [3369] 28. Sharp Bros. Seed Co. 1989. Catalog of wildflowers and forbs. Amarillo, TX: Sharp Bros. Seed Co. 20 p. [18001] 29. Sharp Bros. Seed Co. 1989. Grasses and forbs for erosion control. Fact Sheet. Amarillo, TX: Sharp Bros. Seed Co. 2 p. [18015] 30. Stubbendiek, James; Conard, Elverne C. 1989. Common legumes of the Great Plains: an illustrated guide. Lincoln, NE: University of Nebraska Press. 330 p. [11049] 31. Stubbendieck, James; Nichols, James T.; Butterfield, Charles H. 1989. Nebraska range and pasture forbs and shrubs (including succulent plants). Extension Circular 89-118. Lincoln, NE: University of Nebraska, Nebraska Cooperative Extension. 153 p. [10168] 32. Bulinska-Radomska, Z.; Lester, R. N. 1986. Species relationships in Festuca (Sect. Ovinae, Poaceae). Plant Systematics and Evolution. 154(3-4): 175-182. [7532] 33. Svedarsky, W. D.; Buckley, P. E.; Feiro, T. A. 1986. The effect of 13 years of annual burning on an aspen-prairie ecotone in northwestern Minnesota. In: Clambey, Gary K.; Pemble, Richard H., eds. The prairie: past, present and future: Proceedings, 9th North American prairie conference; 1984 July 29 - August 1; Moorhead, MN. Fargo, ND: Tri-College University Center for Environmental Studies: 118-122. [3540] 34. Tilman, David. 1986. Nitrogen-limited growth in plants from different successional stages. Ecology. 67(2): 555-563. [2809] 35. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 36. Wasser, Clinton H. 1982. Ecology and culture of selected species useful in revegetating disturbed lands in the West. FWS/OBS-82/56. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 347 p. [4837] 37. Weaver, J. E. 1968. Prairie plants and their environment: A fifty-year study in the Midwest. Lincoln, NE: University of Nebraska Press. 276 p. [17546] 38. Weaver, J. E.; Albertson, F. W. 1956. Grasslands of the Great Plains. Lincoln, NE: Johnsen Publishing Company. 395 p. [2463] 39. Weaver, J. E.; Fitzpatrick, T. J. 1934. The prairie. Ecological Monographs. 4(2): 111-295. [2464]

Index

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