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Introductory

SPECIES: Acacia constricta | Whitethorn Acacia
ABBREVIATION : ACACON SYNONYMS : Acaciopsis constricta SCS PLANT CODE : ACCO2 COMMON NAMES : whitethorn acacia whitethorn white-thorn mescat acacia largancillo TAXONOMY : The currently accepted scientific name of whitethorn acacia is Acacia constricta Benth. in Gray [3,16,17]. LIFE FORM : Tree, Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Ronald Uchytil/August 1990 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Uchytil, Ronald J. 1990. Acacia constricta. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Acacia constricta | Whitethorn Acacia
GENERAL DISTRIBUTION : Whitethorn acacia occurs in the Trans-Pecos region of Texas, southern New Mexico, southern and central Arizona, and northern Mexico [12,17]. ECOSYSTEMS : FRES30 Desert shrub FRES33 Southwestern shrubsteppe FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES40 Desert grasslands STATES : AZ NM TX MEXICO ADMINISTRATIVE UNITS : BIBE CACA CORO FOBO GUMO ORPI SAGU BLM PHYSIOGRAPHIC REGIONS : 7 Lower Basin and Range 12 Colorado Plateau 13 Rocky Mountain Piedmont KUCHLER PLANT ASSOCIATIONS : K023 Juniper - pinyon woodland K027 Mesquite bosque K031 Oak - juniper woodlands K032 Transition between K031 and K037 K042 Creosotebush - bursage K043 Paloverde - cactus shrub K044 Creosotebush - tarbush K054 Grama - tobosa prairie K058 Grama - tobosa shrubsteppe K059 Trans-Pecos shrub savanna K065 Grama - buffalograss SAF COVER TYPES : 68 Mesquite 239 Pinyon - juniper 241 Western live oak 242 Mesquite SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Whitethorn acacia is generally not a dominant plant but occurs as scattered individuals in many plant communities. Within Sonoran Desert uplands, it attains its highest densities in washes [11]. It sometimes codominates desert shrub types in the Chihuahuan Desert with creosotebush (Larea tridentata) on gravelly loam [8]. Whitethorn acacia is not listed as a dominant plant or habitat type indicator in any published classifications.

VALUE AND USE

SPECIES: Acacia constricta | Whitethorn Acacia
WOOD PRODUCTS VALUE : Whitethorn acacia is probably used occasionally for firewood. IMPORTANCE TO LIVESTOCK AND WILDLIFE : Livestock: Whitethorn acacia is poor forage for livestock. It is browsed only occasionally in the early spring when the twigs are green, but is otherwise seldom eaten [14]. Cattle eat small amounts of the fruit [14,36]. Wildlife: Mule deer eat small amounts of whitethorn acacia leaves, twigs and fruit [29,31]. Jackrabbits regularly eat the leaves, bark, and twigs [10,37]. White-throated woodrats eat the flowers and leaves [10]. Whitethorn acacia seeds are important in the diets of numerous bird species, especially quail and white-winged doves [20,36]. Several species of desert rodents eat the seeds [21,27]. PALATABILITY : The palatability of whitethorn acacia for livestock is poor [14,32]. Palatability of pods is unclear. Humphrey [14] states that the pods are the most palatable part of the plant and furnish fair forage when green. Vines [36] states, however, that cattle eat the unpalatable legumes when nothing else is available. NUTRITIONAL VALUE : Nutritional information concerning whitethorn acacia is lacking. The seeds and pods of other Acacias are high in protein [5]. COVER VALUE : Whitethorn acacia presumably provides cover for birds and small to medium sized mammals. Some songbirds probably nest within the branches and quail may use the plants for roosting sites [35,36]. Plants provide shade for domestic and wild animals. VALUE FOR REHABILITATION OF DISTURBED SITES : Whitethorn acacia is drought tolerant and may be used for soil stabilization [32]. Near Tucson, Arizona, seedlings transplanted either directly onto copper mine tailings or onto overburden exhibited poor to moderate survival [24]. Seed is not commercially available but may be obtained by harvesting native plants [32]. Seed collection methods have been detailed [39]. OTHER USES AND VALUES : Whitethorn acacia flowers provide an important source of nectar for honey bees [25]. Native peoples ground the beans into a flour to make mush, breads and cakes [36]. MANAGEMENT CONSIDERATIONS : Whitethorn acacia is an invader of semidesert grasslands and has increased in density on grazing lands. It is considered a range pest because it reduces forage for livestock and makes moving and handling cattle more difficult. Numerous herbicides have been used for brush control in the Southwest. In general, whitethorn acacia is susceptible to aerial applications of phenoxy herbicides [30] and tebuthiuron pellets [4,13]. Mortality from mechanical brush control measures varies [22].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Acacia constricta | Whitethorn Acacia
GENERAL BOTANICAL CHARACTERISTICS : Whitethorn acacia frequently occurs as a 3- to 10-foot-tall (0.9-3 m) shrub, but may also develop into a small tree up to 19.5 feet (6 m) tall [3,15]. The brown or reddish-brown twigs contain 0.25 to 1.5 inches (0.6-3.8 cm) long thorns at the nodes [36]. The leaves are bipinnately compound with 3 to 6 pair of pinnae per leaf. Yellow or yellow-orange flowers occur in slender peduncles 0.75 to 1.75 inches (1.9-4.4 cm) long [36]. The reddish-brown to black legume-type fruits are 2 to 4.75 inches (5-12 cm) long, linear, straight to slightly curved, and constricted between the seeds [3,36]. RAUNKIAER LIFE FORM : Undisturbed State: Phanerophyte (microphanerophyte) Burned or Clipped State: Hemicryptophyte REGENERATION PROCESSES : Whitethorn acacia reproduces sexually by producing an abundance of seed. Some plants sprout from the root crown following damage to the aboveground portion of the plant [28]. Flowers are pollinated by insects. In general, Acacias begin to produce seed between 2 and 4 years of age [39]. Several, oblong, mottled, gray-black seeds are encased within a dehiscent legume-type fruit [3]. Seeds are dispersed by wild and domestic animals which eat the fruit. The seeds have a hard seed coat and can probably remain viable for several years [39]. Germination has been reported at 45 percent [36]. Over a 72-year period in desert shrub communities of the Sonoran Desert, whitethorn acacia seedling establishment was very erratic, with only a few new individuals appearing during unusually wet years [9]. Cattle may aid in establishment of new individuals in grasslands, as they eat the fruits and deposit the seeds in a nutrient-rich dung in grazed areas [38]. SITE CHARACTERISTICS : Whitethorn acacia generally inhabits dry slopes and flats, and canyons and washes within desert shrub and semidesert grassland communities. On Sonoran Desert uplands, it is largely confined to runnels and washes [11]. Away from washes, whitethorn acacia occurs as scattered individuals, such as on the slopes of a bajada. It it is often restricted to the upper portions of a bajada, where plant-available moisture is more abundant compared to middle or lower bajada situations [2]. Soils: Although adapted to a wide range of soil types, whitethorn acacia is usually found on sandy, gravelly-loam or caliche soils [8,36]. Associated species: Along washes in the Sonoran Desert, associated shrubs include catclaw acacia (Acacia greggii), velvet mesquite (Prosopis velutina), blue paloverde (Cercidium floridum), desert willow (Chilopsis linearis), and canyon ragweed (Franseria ambrosioides) [11]. On dry slopes and flats, and in grasslands it is often associated with mesquites (Prosopis spp.), creosotebush, tarbush (Flourencia cernua), catclaw acacia, and desertbroom (Baccharis sarothoides) [8,38]. Elevation: Whitethorn acacia occurs from 1,500 to 6,500 feet (457-1,981 m) but is rarely found above 4,500 feet (1,372 m) [32,36]. Regional elevational ranges are presented below [1,8,17]: from 2,500 to 5,000 feet (762-1,524 m) in AZ below 4,600 feet (1,402 m) Rincon Mtns, AZ from 3,600 to 5,000 feet (1,097-1,524 m) Guadalupe Escarpment, NM-TX SUCCESSIONAL STATUS : Catclaw acacia is an invader of semidesert grasslands. Its density has increased over the past 100 years, which probably resulted from a combination of overgrazing, dispersal of seed by livestock, and reduced fire frequencies [11,38]. Whitethorn acacia is often a member of climax desert shrub communities. In the Chihuahuan Desert, it often occurs in climax creosotebush-tarbush communities [23]. In desert shrub communities in the Sonoran Desert near Tucson, Arizona, whitethorn acacia was found to be a long-lived (greater than 72 years) species with a low density of mature individuals [9]. SEASONAL DEVELOPMENT : Whitethorn acacia generally flowers from May to September [3]. It does not produce leaves during the cold winter in the Sonoran Desert, but may foliate in the spring, summer, or autumn if soil moisture is sufficient [11].

FIRE ECOLOGY

SPECIES: Acacia constricta | Whitethorn Acacia
FIRE ECOLOGY OR ADAPTATIONS : Whitethorn acacia is able to sprout from the root crown following top-kill by fire [28]. Seed could potentially be transported to burned sites by domestic and wild animals; however, this has not been documented. POSTFIRE REGENERATION STRATEGY : survivor species; on-site surviving root crown or caudex

FIRE EFFECTS

SPECIES: Acacia constricta | Whitethorn Acacia
IMMEDIATE FIRE EFFECT ON PLANT : Fire generally kills or top-kills whitethorn acacia [28]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Following top-kill by fire, some whitethorn acacia plants survive by sprouting from their root crown. Following a June wildfire in a desert shrub community in south-central Arizona, between 31 and 68 percent of whitethorn acacia plants sprouted [28]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Acacia constricta | Whitethorn Acacia
REFERENCES : 1. Bowers, Janice E.; McLaughlin, Steven P. 1987. Flora and vegetation of the Rincon Mountains, Pima County, Arizona. Desert Plants. 8(2): 50-94. [495] 2. Bowers, Michael A. 1988. Plant associations on a Sonoran Desert bajada: geographical correlates and evolutionary source pools. Vegetatio. 74: 107-112. [4408] 3. Clarke, H. David; Seigler, David S.; Ebinger, John E. 1990. Acacia constricta (Fabaceae: Mimosoideae) and related species from the southwestern U.S. and Mexico. American Journal of Botany. 77(3): 305-315. [10122] 4. Emmerich, W. E.; Helmer, J. D.; Renard, K. G.; Lane, L. J. 1984. Fate and effectiveness of tebuthiuron applied to a rangeland watershed. Journal of Environmental Quality. 13(3): 382-386. [3969] 5. Everitt, James H. 1986. Nutritive value of fruits or seeds of 14 shrub and herb species from south Texas. Southwestern Naturalist. 31(1): 101-137. [5273] 6. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 7. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 8. Gehlbach, Frederick R. 1967. Vegetation of the Guadalupe Escarpment, New Mexico-Texas. Ecology. 48(3): 404-419. [5149] 9. Goldberg, Deborah E.; Turner, Raymond M. 1986. Vegetation change and plant demography in permanent plots in the Sonoran Desert. Ecology. 67(3): 695-712. [4410] 10. Graham, Edward H. 1941. Legumes for erosion control and wildlife. Misc. Publ. 412. Washington, DC: U.S. Department of Agriculture. 153 p. [10234] 11. Hastings, James R.; Turner, Raymond M. 1965. The changing mile: An ecological study of vegetation change with time in the lower mile of an arid and semiarid region. Tuscon, AZ: University of Arizona Press. 317 p. [10533] 12. Hastings, James R.; Turner, Raymond M.; Warren, Douglas K. 1972. An atlas of some plant distributions in the Sonoran Desert. Technical Reports on the Meteorology and Climatology of Arid Regions No. 21. Tuscon, AZ: University of Arizona, Institute of Atmospheric Physics. 255 p. [10534] 13. Herbel, Carlton H.; Morton, Howard L.; Gibbens, Robert P. 1985. Controlling shrubs in the arid Southwest with tebuthiuron. Journal of Range Management. 38(5): 391-394. [10080] 14. Humphrey, R. R. 1950. Arizona range resources. II. Yavapai County. Bull. 229. Tucson, AZ: University of Arizona, Agricultural Experiment Station. 55 p. [5088] 15. Isely, D. 1973. Acacia. Memoirs of the New York Botanical Garden. 25(1): 10-74. [12229] 16. Kartesz, John T.; Kartesz, Rosemarie. 1980. A synonymized checklist of the vascular flora of the United States, Canada, and Greenland. Volume II: The biota of North America. Chapel Hill, NC: The University of North Carolina Press; in confederation with Anne H. Lindsey and C. Richie Bell, North Carolina Botanical Garden. 500 p. [6954] 17. Kearney, Thomas H.; Peebles, Robert H.; Howell, John Thomas; McClintock, Elizabeth. 1960. Arizona flora. 2d ed. Berkeley, CA: University of California Press. 1085 p. [6563] 18. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 19. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 20. Martin, Alexander C.; Zim, Herbert S.; Nelson, Arnold L. 1951. American wildlife and plants. New York: McGraw-Hill Book Company, Inc. 500 p. [4021] 21. Monson, Gale; Kessler, Wayne. 1940. Life history notes on the banner-tailed kangaroo rat, Merriam's kangaroo rat, and white-throated wood rat in Arizona and New Mexico. Journal of Wildlife Management. 4(1): 37-43. [12166] 22. Morton, Howard L.; Ibarra-F., Fernando A.; Martin-R., Martha H.; Cox, Jerry R. 1990. Creosotebush control and forage production in the Chihuahuan and Sonoran Deserts. Journal of Range Management. 43(1): 43-48. [12228] 23. Muller, Cornelius H. 1940. Plant succession in the Larrea-Flourensia climax. Ecology. 21: 206-212. [4244] 24. Norem, M. A.; Day, A. D.; Ludeke, K. L. 1982. An evaluation of shrub and tree species used for revegetating copper mine wastes in the south-western United States. Journal of Arid Environments. 5: 99-304. [1776] 25. Powell, A. Michael. 1988. Trees & shrubs of Trans-Pecos Texas including Big Bend and Guadalupe Mountains National Parks. Big Bend National Park, TX: Big Bend Natural History Association. 536 p. [6130] 26. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 27. Reichman, O. J. 1975. Relation of desert rodent diets to available resources. Journal of Mammalogy. 56(4): 731-751. [4572] 28. Rogers, Garry F.; Steele, Jeff. 1980. Sonoran Desert fire ecology. In: Stokes, Marvin A.; Dieterich, John H., technical coordinators. Proceedings of the fire history workshop; 1980 October 20-24; Tucson, AZ. Gen. Tech. Rep. RM-81. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station: 15-19. [16036] 29. Severson, Kieth E.; Medina, Alvin L. 1983. Deer and elk habitat management in the Southwest. Journal of Range Management Monograph No. 2. Denver: Society for Range Management. 64 p. [2110] 30. Schmutz, Ervin M. 1967. Chemical control of three Chihuahuan Desert shrubs. Weeds. 15: 62-67. [522] 31. Short, Henry L. 1977. Food habits of mule deer in a semi-desert grass-shrub habitat. Journal of Range Management. 30: 206-209. [9895] 32. Thornburg, Ashley A. 1982. Plant materials for use on surface-mined lands. SCS-TP-157. Washington, DC: U.S. Department of Agriculture, Soil Conservation Service. 88 p. [3769] 33. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 34. Vallentine, John F. 1961. Important Utah range grasses. Extension Circular 281. Logan, UT: Utah State University. 48 p. [2937] 35. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 36. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest. Austin, TX: University of Texas Press. 1104 p. [7707] 37. Vorhies, Charles T.; Taylor, Walter P. 1933. The life histories and ecology of jack rabbits, Lepus alleni and Lepus californicus ssp., in relation to grazing in Arizona. Technical Bulletin No. 49. Tucson, AZ: University of Arizona, Agricultural Experiment Station. 117 p. [9933] 38. Whitfield, Charles J.; Anderson, Hugh L. 1938. Secondary succession in the desert plains grassland. Ecology. 19(2): 171-180. [5252] 39. Whitesell, Craig D. 1974. Acacia Mill. acacia. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 184-186. [66]

Index

Related categories for Species: Acacia constricta | Whitethorn Acacia

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