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Introductory

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
ABBREVIATION : CERLED SYNONYMS : NO-ENTRY SCS PLANT CODE : CELE3 CELEI CELEI4 CELEL COMMON NAMES : curlleaf mountain-mahogany TAXONOMY : The currently accepted scientific name for curlleaf mountain-mahogany is Cercocarpus ledifolius Nutt. [20]. It is a member of the rose family (Rosaceae). Kartesz [29] recognized the following three varieties: C. ledifolius var. intercedens Schneid. C. ledifolius var. intermontanus N. Holmgren C. ledifolius var. ledifolius Nutt. Curlleaf mountain-mahogany hybridizes with true mountain-mahogany (C. montanus) and littleleaf mountain-mahogany (C. intricatus) where their ranges overlap [9,57,64]. LIFE FORM : Tree, Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : COMPILED BY AND DATE : Nancy E. McMurray, August 1986 LAST REVISED BY AND DATE : K. Anna Marshall, February 1995 AUTHORSHIP AND CITATION : Marshall, K. Anna. 1995. McMurray, Nancy E. 1986. Cercocarpus ledifolius. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
GENERAL DISTRIBUTION : The distribution of curlleaf mountain-mahogany ranges from the Sierra Nevada and the Cascade Range east to Montana and south to Colorado, northern Arizona, and Baja California, Mexico [10,12,64]. ECOSYSTEMS : FRES20 Douglas-fir FRES21 Ponderosa pine FRES23 Fir-spruce FRES26 Lodgepole pine FRES28 Western hardwoods FRES29 Sagebrush FRES34 Chaparral-mountain shrub FRES35 Pinyon-juniper STATES : AZ CA CO ID MT NV OR UT WA WY MEXICO ADMINISTRATIVE UNITS : BICA BRCA CANY CEBR COLM DEVA DINO GRBA JODA LAVO LABE NABR SEQU YOSE ZION BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 3 Southern Pacific Border 4 Sierra Mountains 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K008 Lodgepole pine-subalpine forest K010 Ponderosa shrub forest K011 Western ponderosa forest K012 Douglas-fir forest K015 Western spruce-fir forest K018 Pine-Douglas-fir forest K019 Arizona pine forest K020 Spruce-fir-Douglas-fir forest K021 Southwestern spruce-fir forest K022 Great Basin pine forest K023 Juniper-pinyon woodland K024 Juniper steppe woodland K037 Mountain-mahogany-oak scrub K038 Great Basin sagebrush K055 Sagebrush steppe SAF COVER TYPES : 206 Engelmann spruce-subalpine fir 208 Whitebark pine 209 Bristlecone pine 210 Interior Douglas-fir 211 White fir 213 Grand fir 217 Aspen 218 Lodgepole pine 219 Limber pine 220 Rocky Mountain juniper 237 Interior ponderosa pine 238 Western juniper 239 Pinyon-juniper 240 Arizona cypress 241 Western live oak 247 Jeffrey pine SRM (RANGELAND) COVER TYPES : 104 Antelope bitterbrush-bluebunch wheatgrass 105 Antelope bitterbrush-Idaho fescue 107 Western juniper/big sagebrush/bluebunch wheatgrass 109 Ponderosa pine shrubland 209 Montane shrubland 210 Bitterbrush 314 Big sagebrush-bluebunch wheatgrass 315 Big sagebrush-Idaho fescue 317 Bitterbrush-bluebunch wheatgrass 318 Bitterbrush-Idaho fescue 319 Bitterbrush-rough fescue 322 Curlleaf mountain-mahogany-bluebunch wheatgrass 401 Basin big sagebrush 402 Mountain big sagebrush 404 Threetip sagebrush 406 Low sagebrush 409 Tall forb 412 Juniper-pinyon woodland 413 Gambel oak 415 Curlleaf mountain-mahogany 416 True mountain-mahogany 417 Littleleaf mountain-mahogany 420 Snowbrush 421 Chokecherry-serviceberry-rose 504 Juniper-pinyon pine woodland 509 Transition between oak-juniper woodland and mahogany-oak association HABITAT TYPES AND PLANT COMMUNITIES : Curlleaf mountain-mahogany occurs throughout the Rocky Mountains and Intermountain West in shrub ecotones or mountain brush communities, in open forests, on ridgetops, and on rock outcrops [4,5,9,33,52]. Curlleaf mountain-mahogany usually occurs in isolated, pure patches that are often very dense. In the Great Basin, curlleaf mountain-mahogany may form a distinct belt on mountain slopes and ridgetops above pinyon-juniper woodland [9,33,52]. In the Rocky Mountain Region, curlleaf mountain-mahogany is associated with skunkbush sumac (Rhus trilobata), antelope bitterbrush (Purshia tridentata), serviceberry (Amelanchier spp.), and western juniper (Juniperus occidentalis) [14]. As a codominant member of the sagebrush-forest ecotone in Idaho, curlleaf mountain mahogany is associated with snowberry (Symphoricarpos spp.), mountain big sagebrush (Artemisia tridentata ssp. vaseyana), green rabbitbrush (Chrysothamnus viscidiflorus), bluebunch wheatgrass (Pseudoroegneria spicata), Sandberg bluegrass (Poa secunda), Idaho fescue (Festuca idahoensis), and Columbia needlegrass (Stipa columbiana) [47]. Curlleaf mountain-mahogany occurs in mid-elevation forests that do not develop dense canopies. It is commonly associated with limber pine (Pinus flexilis), lodgepole pine (P. contorta), ponderosa pine (P. ponderosa), Douglas-fir (Pseudotsuga menziesii), Englemann spruce (Picea engelmannii), subalpine fir (Abies lasiocarpa), and white fir (A. concolor) [4,5,10,33]; it may also occur with quaking aspen (Populus tremuloides) and whitebark pine (Pinus albicaulus) above 9,000 feet in the Great Basin [33]. In Utah curlleaf mountain-mahogany is associated with maple (Acer spp.)-oak (Quercus spp.) woodlands [5]. Publications listing curlleaf mountain-mahogany as a dominant or codominant species include: Habitat characteristics of the Silver Lake mule deer range [11] Sagebrush-grass habitat types of southern Idaho [22] Preliminary descriptions of the terrestrial natural communities of California [23] Plant associations of the Fremont National Forest [24] Plant associations of the Wallowa-Snake Province: Wallowa-Whitman National Forest [28] Flora and major plant communities of the Ruby-East Humboldt Mountains with special emphasis on Lamoille Canyon [36] Grassland and shrubland habitat types of western Montana [41] Forest habitat types of the South Warner Mountains, Modoc County, California [46] Forest habitat types of central Idaho [53] Coniferous forest habitat types of central and southern Utah [63]

VALUE AND USE

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Curlleaf mountain-mahogany is good forage for all classes of browsing animals in both summer and winter [9,14,52,65]; it is one of the few browse species that meets or exceeds the protein requirements for wintering big game animals [9]. In mature stands, much of curlleaf mountain-mahogany foliage is out of reach of browsing animals but provides excellent winter cover [52]. Deer mice in western Nevada consumed curlleaf mountain-mahogany seeds [15]. PALATABILITY : The palatability of curlleaf mountain-mahogany to livestock and wildlife is rated as follows [13]: CO ID MT OR UT WY Cattle ---- ---- Poor ---- Fair Fair Sheep ---- ---- Good Good Good Good Horses ---- ---- Fair ---- Poor Good Pronghorn ---- ---- ---- ---- Fair Poor Bighorn sheep ---- Good ---- Good ---- ---- Elk Good ---- Poor ---- Good Good Mountain goat ---- Good ---- ---- ---- ---- Mule deer Good ---- Good Good Good Good White-tailed deer ---- ---- ---- ---- ---- Good Small mammals ---- ---- ---- ---- Fair Good Small nongame birds ---- ---- ---- ---- Fair Good Upland game birds ---- ---- ---- ---- Fair Fair Waterfowl ---- ---- ---- ---- Poor Poor NUTRITIONAL VALUE : Curlleaf mountain-mahogany is highly nutritious and less seasonally variable than its associates. In southwest and central Montana, the crude protein content of current annual growth averaged 10.5, 10.4, 9.5, and 9.8 percent for spring, summer, fall, and winter, respectively [14]. Curlleaf mountain-mahogany in southeastern Oregon had a high calcium content and intermediate phosphorous content; the ratio of calcium to phosphorus ranged from 5:1 to 9:1. Fiber content was low in mid-spring and high in summer and fall. Curlleaf mountain-mahogany retained a high level of crude fat longer in the spring than its associates [21]. COVER VALUE : Curlleaf mountain-mahogany stands provide hiding and thermal cover for a variety of wildlife species. Cover values are as follows [13]: CO MT OR UT WY Pronghorn ---- ---- ---- Fair Poor Bighorn ---- ---- Good ---- Fair Elk Poor ---- Good Good Good Mule deer Fair Fair Good Good Good White-tailed deer ---- ---- ---- ---- Good Small mammals Good Fair ---- Fair Good Small nongame birds Good Fair ---- Good Good Upland game birds ---- Fair ---- Good Good Waterfowl ---- ---- ---- Poor Poor VALUE FOR REHABILITATION OF DISTURBED SITES : Curlleaf mountain-mahogany may be planted to help stabilize soil in disturbed areas such as roadcuts and mine spoils [25,68]. It was used to revegetate roadcuts in northwestern Montana (out of its normal range); established plants remained in good condition after 9 years [25]. Curlleaf mountain-mahogany may be seeded to improve range [54], although shrub seedlings may exhibit low seedling vigor and slow initial growth rates [40]. It is particularly recommended for southwestern exposures [54]. OTHER USES AND VALUES : Because curlleaf mountain-mahogany wood burns slowly, it was the preferred charcoal wood used for smelting ores in the nineteenth century [33]. It is also highly prized as a barbecue fuel [64]. Because of its tolerance to heat and drought, curlleaf mountain-mahogany can be used for water-efficient landscaping in arid environments [19]. MANAGEMENT CONSIDERATIONS : Pruning curlleaf mountain-mahogany can increase available browse by over 200 percent for 5 years, sustain increased forage production for up to 20 years, encourage the establishment of new seedlings, and increase understory vegetation [59]. On summer range in northern Utah, forage production of plants that were pruned by less than 50 percent increased and remained high for 13 years [2]. Thompson [58] recommended pruning in spring or fall because late fall or winter pruning in Emory County, Utah, resulted in complete loss of all pruned trees. Pruning of trees less than 12 feet tall, with stem diameters of 2 to 4 inches, yielded the best results. Large, mature trees were difficult to prune, had fewer branches suitable for pruning, and often died as a result of pruning.

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
GENERAL BOTANICAL CHARACTERISTICS : Curlleaf mountain-mahogany is a native, xerophytic, evergreen shrub or small tree growing up to 35 feet (10.6 m) tall and 3 feet (0.9 m) in diameter [9,33,57]. The thick, tortuous, leaf-scarred branches [37] arise from a short trunk [33] and form a round or umbrella-shaped crown [14,33]. Leaves are broadly elliptic to lanceolate, 0.48 to 1 inch (12-25 mm) long, leathery, somewhat resinous, and curled under at the margins [64,33,37,57]. Flowers are borne singly or in rows of three in the leaf axils [37,64]. Achenes retain their long, plumose styles [30]. The roots of curlleaf mountain-mahogany play a key role in its ability to inhabit water- and nutrient-deficient substrates. Dealy [66] suggested that a combination of initial rapid root growth and slow top growth may help curlleaf mountain-mahogany outcompete its associates. Average root growth rates of 24 seedlings in a growth chamber were as follows: Average Root Extension Number of Days 13.6 inches (0.34 m) 25 23.2 inches (0.58 m) 62 30.4 inches (0.76 m) 92 39.2 inches (0.98 m) 120 The six longest roots reached a mean depth of 3.7 feet (1.13 m). Curlleaf mountain-mahogany is able to extend its taproot into cracks in bedrock, tapping into otherwise unavailable water supplies [26]. Curlleaf mountain-mahogany roots form nodules in association with nitrogen-fixing, filamentous bacteria which increase available soil nitrogen [33,35,62]. In the San Bernadino Mountains of California, the soil in limber pine stands associated with curlleaf mountain-mahogany contained higher nitrogen concentrations than soils beneath similar stands lacking curlleaf mountain-mahogany [35]. Curlleaf mountain-mahogany can be extremely long-lived. On western and central Nevada sites, Schultz [49] discovered curlleaf mountain-mahogany trees that were approximately 1,350 years old. In Idaho curlleaf mountain-mahogany plants at least 150 years old were found; older stems had rotten cores which made accurate aging impossible [47]. The average age of curlleaf mountain-mahogany on sites in southwestern and central Montana was only 22 years but ranged from 5 to 85 years [14]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Curlleaf mountain-mahogany reproduces by seed. Curlleaf mountain-mahogany can grow vigorously after pruning, but its ability to sprout after top-kill has been described as weak or non-existent in the literature [4,5,8]. The minimum fruit-producing age of curlleaf mountain-mahogany is approximately 15 years [12]. Seed production may be sporadic or cyclic. Few seeds were found in central and southwest Montana curlleaf mountain-mahogany stands in 1973, but seed production was high in 1974 [14,75]. Observations of two stands in central Oregon over 12 years revealed only 3 years of high seed production [66]. High seed production occurred in different years in the two stands and the seed production of individual trees varied from year to year. Wind-drifted seeds occasionally piled up to 10 inches (25 cm) deep in rocky pockets under and around the edges of curlleaf mountain-mahogany stands. Curlleaf mountain-mahogany seeds are primarily wind dispersed [52]. The pointed basal end of the achene and corkscrew-like tail enable it to penetrate the ground [30]. Germination requirements for curlleaf mountain-mahogany are not well understood. Bradley and others [4] suggested that curlleaf mountain-mahogany is dependent on fire because it needs mineral soil for germination. However, Schultz [49] encountered curlleaf mountain-mahogany seedlings growing in deep plant litter under curlleaf mountain-mahogany canopies. Germination experiments on seed collected in central Oregon indicated that stratification increased germination [66]. Incubated seeds having no stratification did not germinate, but stratification for 170 days at 4 deg C resulted in 98 percent germination. Experiments performed on seed collected from sites in Utah, Idaho, and Nevada demonstrated that the chilling requirements for curlleaf mountain-mahogany seeds are variable [31]. Curlleaf mountain-mahogany seed may remain viable for a number of years; in Utah seeds stored in an open, unheated, uncooled warehouse for 7 years germinated at a rate of 76 percent [55]. Curlleaf mountain-mahogany seedlings are sensitive to drought, frost, and browsing [10,47,52]. First-year seedling survival in a curlleaf mountain-mahogany stand in north-central Idaho averaged only 29 percent in the very dry summer of 1968. Winter mortality was also high; the average survival of seedlings protected from all browsing was 50 percent. For seedlings protected from big game and rabbits, survival was 45 percent. For completely unprotected seedlings, survival was 25 percent [47]. SITE CHARACTERISTICS : Curlleaf mountain-mahogany grows on shallow to moderately deep soil at middle to high elevations on gentle to steep slopes, rock outcrops, and ridges. Curlleaf mountain-mahogany commonly occurs on dry, rocky, sandy soils [8,11,14,26,44], although it sometimes occurs on clay or loam soils [10,52]. Elevational ranges for curlleaf mountain-mahogany vary from 2,013 to 4,528 feet (610-1,372 m) in the northern and northwestern parts of its range to 9,900 feet (3,000 m) or higher in Arizona, southern Utah, Nevada, and east-central California [10,30]. Precipitation and growing season information for curlleaf mountain-mahogany stands in Nevada is available. At Great Basin National Park in east-central Nevada, annual precipitation is evenly distributed throughout the year and averages 13.2 inches (330 mm). Average yearly minimum and maximum temperatures are 17.96 degrees Fahrenheit (-7.8 deg C) and 85.46 degrees Fahrenheit (29.7 deg C), respectively [26]. In the Humboldt National Forest, northeastern Nevada, the growing period is 49 days. The mean annual soil temperature is 42.8 degrees Fahrenheit (6 deg C) [27]. SUCCESSIONAL STATUS : Curlleaf mountain-mahogany is somewhat shade tolerant. It is able to grow in open coniferous forests; however, it occurs most often and grows most vigorously on sites without forest canopy. Seedlings establish under the canopies of mature curlleaf mountain-mahogany, but canopy gaps are necessary for long-term survival. The successional role of curlleaf mountain-mahogany varies with community type. Mountain brush communities in which curlleaf mountain-mahogany is either dominant or codominant are generally stable [7]. Changes in relative abundance of codominant species may occur; however, succession rates are extremely slow because vegetation changes depend on soil development which is also slow [10]. In coniferous forests and mesic mountain brush communities adjacent to forests, curlleaf mountain-mahogany is a seral species. Barring major disturbance such as fire, mature conifer stands develop and eventually shade out most curlleaf mountain-mahogany [4,5,7,39]. In western and central Nevada communities in which curlleaf mountain mahogany occurs, large-scale disturbance was infrequent in presettlement times. However, evidence of small-scale disturbance from lightning, low-severity fire, insects, wind, and snow is abundant. Small disturbances often create canopy gaps in dense stands of curlleaf mountain-mahogany. Gaps allow for the release of young curlleaf mountain-mahogany [49]. In western and central Nevada, Schultz and others [48] observed self-tolerance in curlleaf mountain-mahogany seedlings. Seedlings may establish under mature curlleaf mountain-mahogany where they remain as suppressed individuals, often dominating the understory, until canopy gaps release them. Suppressed plants eventually die; they are probably replaced by more seedlings. In mountain brush communities dominated by curlleaf mountain-mahogany, interspecific competition is minimal and curlleaf mountain-mahogany may grow large [48]. Over time, however, intraspecific competition may result in an overall reduction in the biomass production of the community [50]. As a result of intraspecific competition, many curlleaf mountain-mahogany communities are dominated by a few very large, mature individuals. Younger, smaller curlleaf mountain-mahogany may live as understory plants for 100 years or more [9]. SEASONAL DEVELOPMENT : Annual growth in curlleaf mountain-mahogany stands in Montana begins with formation of flower buds in April or May [14]. Flowering generally occurs in May [12,14,64] but may begin as early as April [26] and sometimes extends into July [12]. Branch elongation and leaf production occurred in May through July in eastern Nevada [26]. Seeds are dispersed during late summer (July or August through September) [12,26]. Curlleaf mountain-mahogany phenology at 4,700 feet (1,433 m) near Silver Lake, Oregon, was described as follows [21]: Month Growth Stage mid-April few buds early May many buds mid-May anthesis late June early milk (seeds are soft and immature) early July dough (seeds have a dough-like consistency) mid-July dough mid-August mature seed

FIRE ECOLOGY

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
FIRE ECOLOGY OR ADAPTATIONS : Curlleaf mountain-mahogany may depend on fire to reduce conifer competition and produce favorable soil conditions for seedling establishment [4,5]. However, individual curlleaf mountain-mahogany are severely damaged by fire [4,5,42]. Because many dead branches persist in the crown [14] and leaves are slightly resinous, curlleaf mountain-mahogany is probably very flammable. Curlleaf mountain-mahogany is occasionally a weak sprouter after fire [8]. The presettlement fire regime of curlleaf mountain-mahogany communities probably varied with community type and structure. Arno and Wilson [1] reported that the mean fire interval of curlleaf mountain-mahogany stands along the Salmon River in Idaho ranged from 13 to 22 years until the early 1900's, but lengthened considerably thereafter. However, Schultz [49] found large curlleaf mountain-mahogany up to 1,350 years old in western and central Nevada, indicating that severe fire has been infrequent in some curlleaf mountain-mahogany communities. Schultz also found fire scars on large, old-growth curlleaf mountain-mahogany in the Shoshone Range of central Nevada that suggested understory fuels were insufficient to carry severe fire. Some old-growth curlleaf mountain-mahogany avoid fire by growing on extremely rocky sites [1]. In northern California fire suppression has allowed curlleaf mountain-mahogany to proliferate in formerly open coniferous forests [67]. The presence of curlleaf mountain-mahogany and other shrub species has reduced pine reproduction and increased fuel loadings. When fires do occur on these sites, they may be more severe than in presettlement times. POSTFIRE REGENERATION STRATEGY : Shrub without adventitious-bud root crown Initial-offsite colonizer (off-site, initial community)

FIRE EFFECTS

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
IMMEDIATE FIRE EFFECT ON PLANT : Fire usually kills curlleaf mountain-mahogany. A wildfire occurred at Moose Creek in the Salmon National Forest, Idaho, in August of 1979. Curlleaf mountain-mahogany plants 40 to 80 years old growing on gentle to moderate slopes near the origin of the fire (which burned with "considerable" severity) were mostly killed. Only lightly seared curlleaf mountain-mahogany survived. Intense heat alone may cause mortality in curlleaf mountain-mahogany by searing green growth. Mortality of curlleaf mountain-mahogany was determined in postfire year 1 [8]: # tagged # alive # sprouting completely charred, crown burned 11 0 0 totally seared, crown partially burned 9 0 0 totally seared, crown unburned 11 1 0 partially seared, crown and trunk unburned 9 1 0 DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Curlleaf mountain-mahogany seedlings establish after fire, although establishment may be slow. A curlleaf mountain-mahogany stand near MacKay, Idaho, had burned around 1900. In 1968, it contained plants ranging from 8 to 54 years of age [47]. A stand that burned in 1965 showed no signs of regeneration by 1968. However, Collins [8] described excellent seedling emergence in postfire year 1 of a 1979 wildfire in the Salmon National Forest, possibly due to an unusually wet growing season. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : FIRE MANAGEMENT CONSIDERATIONS : In Montana prescribed fire can improve curlleaf mountain-mahogany forage in seral stands by killing conifers and promoting conditions conducive to seedling establishment. In most cases a fall prescribed fire of several hundred acres is recommended [18]. However, burning is not recommended for all curlleaf mountain-mahogany communities. Young vigorous stands, such as those that have regenerated since 1920, stands surrounded by volatile fuels such as sagebrush, and stands where curlleaf mountain-mahogany distribution is scattered (density of 0.5 plant per acre or less) are not candidates for prescribed burning to improve forage production [18]. Fuel loadings for limber pine/curlleaf mountain-mahogany, Douglas-fir/curlleaf mountain-mahogany, and mixed conifer communities in eastern Idaho and western Wyoming are given [4]. A general fire prescription for curlleaf mountain-mahogany range types in the Northwest is described [38,39].

REFERENCES

SPECIES: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany
REFERENCES : 1. Arno, Stephen F.; Wilson, Andrew E. 1986. Dating past fires in curlleaf mountain-mahogany communities. Journal of Range Management. 39(3): 241-243. [350] 2. Austin, Dennis D. 1990. Response of curlleaf mountain mahogany to pruning in northern Utah 13 years following treatments. In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 61-65. [16096] 3. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 4. Bradley, Anne F.; Noste, Nonan V.; Fischer, William C. 1991. Fire ecology of forests and woodlands in Utah. Gen. Tech. Rep. INT-287. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 128 p. [18211] 5. Bradley, Anne F.; Fischer, William C.; Noste, Nonan V. 1992. Fire ecology of the forest habitat types of eastern Idaho and western Wyoming. Gen. Tech. Rep. INT-290. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 92 p. [19557] 6. Britton, Carlton M.; Wright, Henry A. 1983. Brush management with fire. In: McDaniel, Kirk C., ed. Proceedings--brush management symposium; 1983 February 16; Albuquerque, NM. Denver, CO: Society for Range Management: 61-68. [521] 7. Christensen, Earl M. 1964. Succession in a mountain brush community in central Utah. Utah Academy Proceedings. 41(1): 10-13. [6913] 8. Collins, Thomas C. 1980. A report on the Moose Creek Fire of August, 1979. Unpublished report on file with: U.S. Department of Agriculture, Forest Service, Salmon National Forest, North Fork Ranger District, North Fork, ID. 27+ p. [666] 9. Davis, James N. 1990. General ecology, wildlife use, and management of the mountain mahoganies in the Intermountain West. In: Johnson, Kendall L., ed. Proceedings, 5th Utah shrub ecology workshop: The genus Cercocarpus; 1988 July 13-14; Logan, UT. Logan, UT: Utah State University, College of Natural Resources: 1-13. [16092] 10. Davis, James N.; Brotherson, Jack D. 1991. Ecological relationships of curlleaf mountain-mahogany (Cercocarpus ledifolius Nutt.) communities in Utah and implications for management. Great Basin Naturalist. 51(2): 153-166. [15471] 11. Dealy, J. Edward. 1971. Habitat characteristics of the Silver Lake mule deer range. Res. Pap. PNW-125. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 99 p. [782] 12. Deitschman, Glenn H.; Jorgensen, Kent R.; Plummer, A. Perry. 1974. Cercocarpus H.B.K. cercocarpus (mountain-mahogany). In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 309-312. [7583] 13. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 14. Duncan, Elizabeth Ann. 1975. The ecology of curl-leaf mountain mahogany (Cercocarpus ledifolius Nutt.) in southwestern Montana with special reference to use by mule. Bozeman, MT: Montana State University. 87 p. Thesis. [12585] 15. Everett, Richard L.; Meeuwig, Richard O.; Stevens, Richard. 1978. Deer mouse preference for seed of commonly planted species, indigenous weed seed, and sacrifice foods. Journal of Range Management. 31(1): 70-73. [896] 16. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 17. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 18. Gruell, George E.; Brown, James K.; Bushey, Charles L. 1986. Prescribed fire opportunities in grasslands invaded by Douglas-fir: state-of-the-art guidelines. Gen. Tech. Rep. INT-198. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 19 p. [1050] 19. Gutknecht, Kurt W. 1989. Xeriscaping: an alternative to thirsty landscapes. Utah Science. 50(4): 142-146. [10166] 20. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. Berkeley, CA: University of California Press. 1400 p. [21992] 21. Hickman, O. Eugene. 1975. Seasonal trends in the nutritive content of important range forage species near Silver Lake, Oregon. Research Paper PNW-187. 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Index

Related categories for Species: Cercocarpus ledifolius | Curlleaf Mountain-Mahogany

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