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Introductory

SPECIES: Menziesia ferruginea | Menziesia
ABBREVIATION : MENFER SYNONYMS : Menziesia glabella (Gray) Peck SCS PLANT CODE : MEFE COMMON NAMES : menziesia false huckleberry fool's huckleberry false azalea mock azalea rusty menziesia smooth menziesia Pacific menziesia skunkbrush rustyleaf TAXONOMY : The currently accepted scientific name of menziesia is Menziesia ferruginea (Smith) [26]. In existence are two varieties distinguished by morphological variations and geographic location [25,26,32,33]: M. ferruginea var. ferruginea M. ferruginea var. glabella (Gray) Peck LIFE FORM : Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : R. J. Habeck, January 1992 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Habeck, R. J. 1992. Menziesia ferruginea. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Menziesia ferruginea | Menziesia
GENERAL DISTRIBUTION : M. ferruginea var. ferruginea is found north along the Alaskan Coast, and south along the Pacific Coast to northern California, inland across the Cascades and Rocky Mountains to Wyoming [25,33]. M. ferruginea var. glabella is distributed from British Columbia to Alberta, south to Montana, Wyoming, Idaho, eastern Washington, and Oregon; down the Columbia River to Mt. Hood and Mt. Adams, where the two varieties freely interbreed [25,27]. ECOSYSTEMS : FRES20 Douglas-fir FRES22 Western white pine FRES23 Fir - spruce FRES24 Hemlock - Sitka spruce FRES25 Larch STATES : AK CA ID MT OR WA WY AB BC ADMINISTRATIVE UNITS : BIHO DENA GLBA GLAC GRTE LACL MORA NOCA OLYM REDW BLM PHYSIOGRAPHIC REGIONS : 1 Northern Pacific Border 2 Cascade Mountains 4 Sierra Mountains 5 Columbia Plateau 8 Northern Rocky Mountains 9 Middle Rocky Mountains KUCHLER PLANT ASSOCIATIONS : K001 Spruce - cedar - hemlock forest K002 Cedar - hemlock - Douglas-fir forest K003 Silver fir - Douglas-fir forest K004 Fir - hemlock forest K012 Douglas-fir forest K013 Cedar - hemlock - pine forest K014 Grand fir - Douglas-fir forest K015 Western spruce - fir forest SAF COVER TYPES : 205 Mountain hemlock 206 Engelmann spruce - subalpine fir 210 Interior Douglas-fir 212 Western larch 213 Grand fir 215 Western white pine SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Menziesia is an understory dominant on moist, wooded north- and east-facing slopes of the montane to upper subalpine zones. Overstory components mostly include subalpine fir (Abies lasiocarpa), Pacific silver fir (A. amabilis), western redcedar (Thuja plicata), mountain hemlock (Tsuga mertensiana), western hemlock (T. heterophylla), and Engelmann spruce (Picea engelmannii) [25]. Menziesia's associates generally include blue huckleberry (Vaccinium globulare), smooth woodrush (Luzula hitchcockii), and on higher sites, white rhododendron (Rhododendron albiflorum) [33]. In Alaska, menziesia is a common understory shrub in the coastal spruce-hemlock forest, often under a dense canopy. It also grows in the southern part of the boreal forest in white spruce (Picea glauca) and white spruce-paper birch (Betula papyrifera) stands [53]. Publications listing menziesia as an indicator or dominant species in habitat types (hts), community types (cts), or plant associations (pas) are presented below: Area Classification Authority AK forest (cts) Reynolds 1990 c ID forest (hts) Steele & others 1981 n ID forest (hts) Cooper & others 1991 e ID and w WY forest (hts) Steele & others 1983 MT forest (hts) Pfister & others 1977 OR: Mt. Hood & Willamette N.F. forest (pas) Hemstrom & others 1982 OR: Wallowa-Whitman National Forest forest (pas) Johnson & Simon 1987 WA: Mount Rainier N.P. forest (pas) Franklin & others 1988 WA: Gifford Pinchot National Forest forest (pas) Brockway & others 1983

VALUE AND USE

SPECIES: Menziesia ferruginea | Menziesia
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Menziesia's value to wildlife is primarily as food. In Idaho and Montana, elk used menziesia in considerable amounts in July, and to a lesser extent in October. Elk rarely eat menziesia when it occurs with more palatable browse species. Menziesia filled a browse requirement during July when elk were primarily on an herbaceous diet in western Montana and Idaho [14,54]. Mule deer use menziesia moderately in summer and fall in the western states [31]. Menziesia accounted for over 15 percent of the summer moose diet in the spruce-fir cover types of Jackson Hole, Wyoming [28]. Alaskan spruce grouse use small amounts of menziesia capsules in their October diet [15]. Following a burn treatment in western Montana, menziesia came back as an important source of fruits and seeds used by rodents such as deer mice, chipmunks, and voles [21]. PALATABILITY : The palatability of menziesia is generally rated poor. This shrub provides fair summer browse for deer and elk [35]. Menziesia's palatability was considered very low for moose in Jackson Hole, Wyoming [23]. NUTRITIONAL VALUE : Nutrient value of menziesia is considered poor to medium [20]. Menziesia contains toxic resinoids that are known to cause livestock losses when leaves and roots are eaten [43]. Mean values of percent composition of macro- and micronutrients for menziesia are as follows [22]: MACRO MICRO (percent dry weight) (ppm dry weight) --------------------------------------------------------------------- N P K Ca Mg Na Cu Mn Fe Zn --------------------------------------------------------------------- Leaves 2.48 0.24 1.19 0.62 0.40 0.47 10.0 16,656 62.0 86.6 Stems 1.18 0.15 0.85 0.35 0.17 0.09 11.8 7139.2 22.8 40.5 COVER VALUE : Menziesia provides shelter and cover for a variety of wildlife species. Dense thickets provide good cover for many small birds and mammals. VALUE FOR REHABILITATION OF DISTURBED SITES : NO-ENTRY OTHER USES AND VALUES : Menziesia is a desirable ornamental due its crimson-orange autumn foliage [26]. MANAGEMENT CONSIDERATIONS : Menziesia is mostly threatened by logging activities. Following clearcutting of Pacific silver fir and mountain hemlock, menziesia exhibits a difficult time regenerating [6]. In northwestern Montana, menziesia decreased considerably from its high abundance following a clearcut in an old-growth stand [2]. Soil layers restricting root growth and high water tables are common enough that compaction or erosion-causing activities should be carefully controlled [6]. There appears to be little destruction of mature menziesia plants in spruce-fir forests when selective logging is done during the winter [9]. If menziesia is a major component of the mature forest understory, it will continue to dominate the postharvest plant community [10]. Mechanical site preparation: Coates [8] found that recovery of menziesia on Engelmann spruce and subalpine fir sites after clipping and mechanical scarification was very limited and confined to the production of new shoots from buds on old stems rather than from basal sprouts or suckers. Two seasons after cutting, new shoots had not reached the height of precut stems, and did not overtop spruce or pine seedlings planted at the time of cutting. Menziesia control by Esteron brush killer was best achieved when this herbicide was mixed with diesel and sprayed by helicopter from 150 to 200 feet (46-61 m) high [40]. After 2 years, Tordon 101 and Banvel 720 killed 100 percent of menziesia when sprayed on leaf foliage [39].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Menziesia ferruginea | Menziesia
GENERAL BOTANICAL CHARACTERISTICS : Menziesia is an erect, decidous shrub 3.3 to 6.6 feet (1-2 m) tall. Ovate-elliptic to elliptic-obovate leaves are 1.5 to 2.5 inches (4-6 cm) long and exude a skunky oder when crushed. Menziesia has scaly bark on older branches, while the young twigs are covered with fine hair [20]. The fruits are oval capsules containing many seeds 0.20 to 0.28 inch (5-7 mm) long. The leaves and stems of this shrub are sticky to the touch [25,26,44]. Menziesia decreases in height with increasing elevation. This would imply that menziesia's height is generally at its maximum on western hemlock sites, and at its minimum on mountain hemlock and subalpine fir sites [34]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Seed production and dispersal: Numerous small seeds are produced in capsules which dehisce naturally at maturity. Seeds are wind or gravity dispersed. Vegetative Reproduction: Menziesia resprouts from root crowns and adventitious stem buds after destruction of aboveground parts. When covered with ash from Mt. St. Helen's, menziesia produced adventitious roots [3]. This plant also appears to be capable of layering. Layering is an effective means of growth and expansion at high elevations where heavy snowpacks press menziesia branches close to the ground [10]. On steep slopes, menziesia tends to grow downhill with the tips of its branches ascending [35]. SITE CHARACTERISTICS : Menziesia is a common shade-tolerant shrub, found predominantly on steep, northerly exposures with abundant moisture [17,20,25,26]. Menziesia generally occurs from 3,300 to 7,500 feet (1,000-2,300 m) in elevation on 15 to 60 percent slopes [11,29,50]. Absence from extreme northern distributions may imply a limitation to cold temperatures. Menziesia may be restricted to areas with adequate snow cover, which protects plants from winter desiccation [25]. In Oregon, menziesia is typically found on high-elevation sites where frequent frost, heavy snowpacks, and short growing seasons are common [6]. The presence of menziesia has been reported as being a particularly good indicator of site potential [13]. Menziesia often grows under dense canopies, in openings, and on cut-over forest land, especially on well-drained slopes [53]. Mueggler [42] found the distribution of menziesia in cedar-hemlock stands to be positively associated with low soil potassium content, and high organic matter content (5.6 to 8.0 percent). Soils: Soil depth usually ranges from 17 to 70 inches (43-178 cm). Soils are generally derived from loam to sandy-loam pumice layers to a depth of 15 to 30 inches (38-76 cm) [6,11,24]. In central Idaho, soil acidity ranges from pH 5.1 to 6.2; litter depth can average at least 3.5 inches (9 cm) [6,11]. Effective rooting depth in Oregon is approximately 46 inches (117 cm) [6]. SUCCESSIONAL STATUS : Menziesia is most always associated with cold, wet habitat types. Menziesia is very shade tolerant. It can persist in heavily shaded forests where light levels are typically 5 percent of full sunlight. Alaback [1] reported that menziesia decreased in biomass productivity in Alaskan spruce-hemlock forests over 30 years of age. Menziesia biomass was lowest in stands 31 to 130 years old, but after this period increased. In western Montana, burning increased menziesia fourfold from immature stands (less than 90 years) to old stands (greater than 150 years) [2]. SEASONAL DEVELOPMENT : Phenology: Menziesia flowers from late May through July; capsules mature in July and August [53]. Leaves expand rapidly in early June at the time flowers open and begin to senesce in mid-August and September at high elevations in southern British Columbia [10]. Menziesia has been found to live at least 30 years in northern forests [1].

FIRE ECOLOGY

SPECIES: Menziesia ferruginea | Menziesia
FIRE ECOLOGY OR ADAPTATIONS : Menziesia is a fire-sensitive species [51]. It responds to fire by sprouting from the root crown [38]. Since menziesia is indicative of cool, moist sites, fire plays a minor role in its development. Barrett and Arno [4] found that the subalpine fir/menziesia cover type in the Selway-Bitterroot Wilderness, Idaho, had a mean fire interval of 174 to 181 years, achieving lethal/nonuniform fire behavior. POSTFIRE REGENERATION STRATEGY : survivor species; on-site surviving root crown or caudex

FIRE EFFECTS

SPECIES: Menziesia ferruginea | Menziesia
IMMEDIATE FIRE EFFECT ON PLANT : Menziesia is very susceptible to fire-kill. Moderate to severe fires reduce survival and slow redevelopment [44]. Wildfires apparently removed menziesia totally from experimental plots in Washington and Oregon for up to 16 years [58]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : In western Montana, Stickney [51] found that postburn survival of menziesia was poor; menziesia was very slow to redevelop after fire, especially in areas that had been intensley burned. The average percent cover of menziesia doubled from unburned plots to lightly burned plots, and tripled from moderately burned plots in western Montana [48]. Another study found menziesia to increase its density of aboveground parts the first year after burning [37]. Model projections predict that menziesia will reach a height of 2.4 feet (.73 m) 2 years after a burn, and that the average height of menziesia will peak at 5.1 to 5.6 feet (1.5-1.7 m) 20 to 25 years after burning [34]. Mueggler [42], however, found no significant difference in cover and frequency between burned and unburned sites in northern Idaho. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Menziesia ferruginea | Menziesia
REFERENCES : 1. Alaback, Paul B. 1982. Dynamics of understory biomass in Sitka spruce-western hemlock forests of southeast Alaska. Ecology. 63(6): 1932-1948. [7305] 2. Antos, J. A.; Habeck, J. R. 1981. Successional development in Abies grandis (Dougl.) Forbes forests in the Swan Valley, western Montana. Northwest Science. 55(1): 26-39. [12445] 3. Antos, Joseph A.; Zobel, Donald B. 1985. Plant form, developmental plasticity and survival following burial by volcanic tephra. Canadian Journal of Botany. 63: 2083-2090. [12553] 4. Barrett, Stephen W.; Arno, Stephen F. 1991. Classifying fire regimes and defining their topographic controls in the Selway-Bitterroot Wilderness. In: Andrews, Patricia L.; Potts, Donald F., eds. Proceedings, 11th annual conference on fire and forest meteorology; 1991 April 16-19; Missoula, MT. SAF Publication 91-04. Bethesda, MD: Society of American Foresters: 299-307. [16179] 5. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 6. Brockway, Dale G.; Topik, Christopher; Hemstrom, Miles A.; Emmingham, William H. 1985. Plant association and management guide for the Pacific silver fir zone: Gifford Pinchot National Forest. R6-Ecol-130a. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 122 p. [525] 7. Cates, Rex G.; Orians, Gordon H. 1975. Successional status and the palatability of plants to generalized herbivores. Ecology. 56: 410-418. [15989] 8. Coates, K. David. 1987. Effects of shrubs and herbs on conifer regeneration and microclimate in the Rhododendron-Vaccinium-Menziesia community of south-central BC. Vancouver, BC: University of British Columbia. Thesis. Abstract. [17445] 9. Coates, D.; Haeussler, S. 1986. A preliminary guide to the response of major species of competing vegetation to silvicultural treatments. Victoria, BC: Ministry of Forests, Information Services Branch; Land Management Handbook Number 9. 88 p. [17453] 10. Comeau, Philip G.; Watts, Susan B.; Caza, Caroline L.; [and others]. 1989. Autecology, biology, competetive status and response to treatment of seven southern interior weed species. FRDA Report 093; ISSN 0835 0572. Victoria, BC: BC Ministry of Forests, Research Branch. 46 p. [9471] 11. Cooper, Stephen V.; Neiman, Kenneth E.; Roberts, David W. 1991. (Rev.) Forest habitat types of northern Idaho: a second approximation. Gen. Tech. Rep. INT-236. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 143 p. [14792] 12. Crane, M. F.; Fischer, William C. 1986. Fire ecology of the forest habitat types of central Idaho. Gen. Tech. Rep. INT-218. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 85 p. [5297] 13. Daubenmire, Rexford F.; Daubenmire, Jean B. 1968. Forest vegetation of eastern Washington and northern Idaho. Technical Bulletin 60. Pullman, WA: Washington State University, Agricultural Experiment Station. 104 p. [749] 14. Edge, W. Daniel; Marcum, C. Les; Olson-Edge, Sally L. 1988. Summer forage and feeding site selection by elk. Journal of Wildlife Management. 52(4): 573-577. [6778] 15. Ellison, Laurence. 1966. Seasonal foods and chemical analysis of winter diet of Alaskan spruce grouse. Journal of Wildlife Management. 30(4): 729-735. [9735] 16. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 17. Forsythe, Warren Louis. 1975. Site influence on the post-fire composition of a Rocky Mountain forest. Missoula, MT: University of Montana. 173 p. Dissertation. [6723] 18. Franklin, Jerry F.; Moir, William H.; Hemstrom, Miles A.; [and others]. 1988. The forest communities of Mount Rainier National Park. Scientific Monograph Series No 19. Washington, DC: U.S. Department of the Interior, National Park Service. 194 p. [12392] 19. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 20. Green, R. N.; Courtin, P. J.; Klinka, K.; [and others]. 1984. Site diagnosis, tree species selection, and slashburning guidelines for the Vancouver Forest Region. Land Management Handbook Number 8. Abridged version. Burnaby, BC: Ministry of Forests, Vancouver Forest Region. 143 p. [9475] 21. Halvorson, Curtis H. 1982. Rodent occurrence, habitat disturbance, and seed fall in a larch-fir forest. Ecology. 63(2): 423-433. [8522] 22. Hanley, Thomas A.; McKendrick, Jay D. 1983. Seasonal changes in chemical composition and nutritive values of native forages in a spruce-hemlock forests, southeastern Alaska. Res. Pap. PNW-312. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station. 41 p. [8770] 23. Harry, G. Bryan. 1957. Winter food habits of moose in Jackson Hole, Wyoming. Journal of Wildlife Management. 21(1): 53-57. [8429] 24. Hemstrom, Miles A.; Emmingham, W. H.; Halverson, Nancy M.; [and others]. 1982. Plant association and management guide for the Pacific silver fir zone, Mt. Hood and Willamette National Forests. R6-Ecol 100-1982a. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 104 p. [5784] 25. Hickman, James C.; Johnson, Michael P. 1969. An analysis of geographical variation in western North American Menziesia (Ericaceae). Madrono. 20(1): 1-32. [17109] 26. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 27. Hitchcock, C. Leo; Cronquist, Arthur; Ownbey, Marion. 1959. Vascular plants of the Pacific Northwest. Part 4: Ericaceae through Campanulaceae. Seattle, WA: University of Washington Press. 510 p. [1170] 28. Houston, Douglas B. 1968. The Shiras Moose in Jackson Hole, Wyoming. Tech. Bull. No. 1. [Place of publication unknown]: The Grand Teton Natural History Association. 110 p. [7824] 29. Johnson, Charles G., Jr.; Simon, Steven A. 1987. Plant associations of the Wallowa-Snake Province: Wallowa-Whitman National Forest. R6-ECOL-TP-255A-86. Baker, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region, Wallowa-Whitman National Forest. 399 p. [9600] 30. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 31. Kufeld, Roland C.; Wallmo, O. C.; Feddema, Charles. 1973. Foods of the Rocky Mountain mule deer. Res. Pap. RM-111. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 31 p. [1387] 32. Lackschewitz, Klaus. 1986. Plants of west-central Montana--identification and ecology: annotated checklist. Gen. Tech. Rep. INT-217. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 128 p. [2955] 33. Lackschewitz, Klaus. 1991. Vascular plants of west-central Montana--identification guidebook. Gen. Tech. Rep. INT-227. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 648 p. [13798] 34. Laursen, Steven B. 1984. Predicting shrub community composition and structure following management disturbance in forest ecosystems of the Intermountain West. Moscow, ID: University of Idaho. 261 p. Dissertation. [6717] 35. Lee, Lyndon C.; Pfister, Robert D. 1978. A training manual for Montana forest habitat types. Missoula, MT: University of Montana, School of Forestry, Montana Forest and Conservation Experiment Station. 142 p. [1434] 36. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 37. Miller, Melanie. 1976. Shrub sprouting response to fire in a Douglas-fir/western larch ecosystem. Missoula, MT: University of Montana. 124 p. Thesis. [8945] 38. Miller, Melanie. 1977. Response of blue huckleberry to prescribed fires in a western Montana larch-fir forest. Gen. Tech. Rep. INT-188. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 33 p. [6334] 39. Miller, Daniel L.; Kidd, Frank A. 1983. Shrub control in the Inland Northwest--a summary of herbicide test results. Forestry Research Note RN-83-4. Lewiston, ID: Potlatch Corporation. 49 p. [7861] 40. Miller, Daniel L.; Robinson, Vernon S. 1983. The influence of spray altitude on shrub control. Forestry Research Note RN-83-1. Lewiston, ID: Potlach Corporation, Wood Products, Western Division. 6 p. [3399] 41. Morris, William G. 1970. Effects of slash burning in overmature stands of the Douglas-fir region. Forest Science. 16(3): 258-270. [4810] 42. French, N. R. 1978. Reexamination of permanent vegetation plots on the Idaho National Engineering Laboratory Site. In: Markham, O. D., ed. Ecological studies on the Idaho National Engineering Laboratory Site: 1978 Progress Report. IDO-12087. Idaho Falls, ID: U.S.Dept. of Energy, Environ. Sciences Branch, Radiological and Environmental Sciences Lab:167-170. [969] 43. Mueggler, W. F. 1970. Objectionable characteristics of range plants. In: Range and wildlife habitat evaluation--a research symposium: Proceedings; 1968 May; Flagstaff; Tempe, AZ. Misc. Publ. 1147. Washington, DC: U.S. Department of Agriculture, Forest Service: 63-70. [12986] 44. Patterson, Patricia A.; Neiman, Kenneth E.; Tonn, Jonalea. 1985. Field guide to forest plants of northern Idaho. Gen. Tech. Rep. INT-180. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 246 p. [1839] 45. Pfister, Robert D.; Kovalchik, Bernard L.; Arno, Stephen F.; Presby, Richard C. 1977. Forest habitat types of Montana. Gen. Tech. Rep. INT-34. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 174 p. [1878] 46. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 47. Reynolds, Keith M. 1990. Preliminary classification of forest vegetation of the Kenai Penninsula, Alaska. Res. Pap. PNW-RP-424. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 67 p. [14581] 48. Stark, N.; Steele, R. 1977. Nutrient content of forest shrubs following burning. American Journal of Botany. 64(10): 1218-1224. [2224] 49. Steele, Robert; Cooper, Stephen V.; Ondov, David M.; [and others]. 1983. Forest habitat types of eastern Idaho-western Wyoming. Gen. Tech. Rep. INT-144. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 122 p. [2230] 50. Steele, Robert; Pfister, Robert D.; Ryker, Russell A.; Kittams, Jay A. 1981. Forest habitat types of central Idaho. Gen. Tech. Rep. INT-114. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 138 p. [2231] 51. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 52. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 53. Viereck, Leslie A.; Little, Elbert L., Jr. 1972. Alaska trees and shrubs. Agric. Handb. 410. Washington, DC: U.S. Department of Agriculture, Forest Service. 265 p. [6884] 54. Young, Vernon A.; Robinette, W. Leslie. 1939. A study of the range habits of elk on the Selway Game Preserve. Bull. No. 9. Moscow, ID: University of Idaho, School of Forestry. 47 p. [6831] 55. Zager, Peter Edward. 1980. The influence of logging and wildfire on grizzly bear habitat in northwestern Montana. Missoula, MT: University of Montana. 131 p. Dissertation. [5032]

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