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Wildlife, Animals, and Plants |
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INTRODUCTORY
ABBREVIATION:PAXMYR
SYNONYMS:Pachistima myrsinites (Pursh) Raf. [24,58]
NRCS PLANT CODE [51]:PAMY
COMMON NAMES:Oregon boxwood paxistima pachistima myrtle boxwood boxwood mountain-lover Montana-box mountain-box myrtle-bush myrtle-boxleaf false-box myrtle pachistima
TAXONOMY:The currently accepted scientific name of Oregon boxwood is Paxistima myrsinites (Pursh) Raf. (Celastraceae) [58,59].
LIFE FORM:Shrub
FEDERAL LEGAL STATUS:No special status
OTHER STATUS:No entry
AUTHORSHIP AND CITATION:Snyder, S. A. 1991. Paxistima myrsinites. In: Remainder of citation DISTRIBUTION AND OCCURRENCE
GENERAL DISTRIBUTION:
ECOSYSTEMS [18]:
STATES:
BLM PHYSIOGRAPHIC REGIONS [5]:
KUCHLER [30] PLANT ASSOCIATIONS:
SAF COVER TYPES [17]:
HABITAT TYPES AND PLANT COMMUNITIES:
Publications listing Oregon boxwood as an
indicator are listed below.
VALUE AND USE
IMPORTANCE TO LIVESTOCK AND WILDLIFE:Oregon boxwood is considered important forage for deer, elk, and moose [9,13,19,43,56]. Mountain sheep and grouse also browse Oregon boxwood [52]. Livestock occasionally eat Oregon boxwood, but it is not considered an important forage species [52,56].
PALATABILITY:The degree of use shown by livestock for Oregon boxwood has been rated as follows [14]: UT CO MT Cattle poor poor poor Sheep fair poor poor Horse poor poor poor
NUTRITIONAL VALUE:Stark [46] has reported on the nutrient content of Oregon boxwood following harvest and burn treatments in western Montana. The nutritional value of Oregon boxwood has been rated as follows [14]: UT WY MT Elk fair --- poor Mule deer fair poor --- White-tailed deer --- fair --- Antelope poor --- --- Upland game bird poor poor --- Waterfowl poor --- --- Nongame bird poor poor --- Small mammal poor --- ---
COVER VALUE:The degree to which Oregon boxwood provides environmental protection during one or more seasons for wildlife species has been rated as follows [14]: MT UT WY Pronghorn --- poor --- Elk poor poor --- Mule deer poor poor --- White-tailed deer --- --- poor Small mammals poor fair --- Nongame birds poor fair --- Upland game birds poor fair --- Waterfowl --- poor ---
VALUE FOR REHABILITATION OF DISTURBED SITES:Oregon boxwood root cuttings or 2-0 stock can be used to revegetate disturbed sites [38].
OTHER USES AND VALUES:Oregon boxwood is easily shaped and adapts well to both sunny and shady spots, making it ideal for an ornamental and ground cover [23,29,52].
MANAGEMENT CONSIDERATIONS:Oregon boxwood is not easily controlled with herbicides, possibly because of its leathery, evergreen leaves [3,21,37]. Oregon boxwood appears to increase in logged areas compared to uncut areas in grand fir (Abies grandis) types of western Montana [1]. In general, Oregon boxwood seems to increase following logging; however, it may not really benefit from management treatments because undisturbed shrubs may have the same growth rate as disturbed shrubs [32]. In logged grand fir/Oregon boxwood sites of northern Idaho, Oregon boxwood decreased for the first 7 years, then increased after 25 years to higher cover values than in unlogged areas [55]. BOTANICAL AND ECOLOGICAL CHARACTERISTICS
GENERAL BOTANICAL CHARACTERISTICS:Oregon boxwood is a native, cool-season, evergreen shrub, with maroon flowers [24]. It is low growing, reaching heights of 1 to 3 feet (0.3-1 m), sometimes spreading, and densely branched. Its leaves are oblong and glabrous. Its fruit is a one- to two-seeded capsule [40,52].
RAUNKIAER [44] LIFE FORM:Phanerophyte
REGENERATION PROCESSES:Oregon boxwood stems can layer and root, and the shrub can be propagated easily through stem cuttings [29]. Seeds are dispersed by gravity, with no evidence to suggest they are dispersed by any other means [42].
SITE CHARACTERISTICS:Oregon boxwood grows on dry to moist sites in shaded mountain areas as high as subalpine habitats, but can be found at sea level in California [29,31]. It can grow in frost pockets in steep ravines or in open woods, ridgetops, and glades [12,22,28,29]. Oregon boxwood can occur on well-drained, shallow, gravelly soils, in clay and silt loams, and cobbly clay [22]. In British Columbia Oregon boxwood occurs on Podzols and Regosols [36]. Elevational ranges have been listed for some western states and provinces [2,14,36,39,52]: from 5,000 to 10,500 feet (1,524-3,200 m) in Utah from 6,600 to 11,000 feet (2,012-3,353 m) in Colorado from 6,700 to 9,500 feet (2,042-2,896 m) in Wyoming from 3,500 to 7,600 feet (1,067-2,317 m) in Montana from 6,900 to 8,200 feet (2,103-2,499 m) in Idaho from 6,000 to 10,000 feet (1,829-3,048 m) in Arizona/New Mexico from 4,020 to 5,160 feet (1,279-1,600 m) in California/Oregon from 3,950 to 4,950 feet (1,200-1,500 m) in British Columbia
SUCCESSIONAL STATUS:Oregon boxwood is an indicator species in several western habitat types and plant communities. It is a climax shrub and can tolerate both sun and shade [23], but it usually indicates dry to moist, cool sites and well-drained soils [19]. Quaking aspen/Saskatoon serviceberry (Amelanchier alnifolia)-Oregon boxwood community types in Idaho appear to be stable but may be slowly successional to quaking aspen-Douglas-fir/ Saskatoon serviceberry community types which might be in the Oregon boxwood phase of the Douglas-fir/pinegrass habitat type or the subalpine/pinegrass habitat type [39].
SEASONAL DEVELOPMENT:The following months have been noted for budding, flowering, and fruiting of Oregon boxwood. State Buds Flowers Fruits Source CA May-July [40] AZ, MX April-June June-Sept [52] OR, WA April-June [19] ID, MT March-May March-June July-Sept [15,42] UT April-July [14] CO May-July [14] WY June-August [14] FIRE ECOLOGY
FIRE ECOLOGY OR ADAPTATIONS:Following fire, Oregon boxwood can sprout from buds on the taproot or from the root crown [10,35,42]. Some seedling establishment via short-term viablity seed stored on-site may also occur [49].
POSTFIRE REGENERATION STRATEGY [33]:Small shrub, adventitious bud/root crown FIRE EFFECTS
IMMEDIATE FIRE EFFECT ON PLANT:Oregon boxwood can survive low- to moderate-severity fires that do not consume the duff or raise the soil temperature too high [10]. It can, however, be killed by severe fires [9].
DISCUSSION AND QUALIFICATION OF FIRE EFFECT:No entry
PLANT RESPONSE TO FIRE:Oregon boxwood usually sprouts from its root crown or from buds on its taproot following low- to moderate-severity fires [10,42].
DISCUSSION AND QUALIFICATION OF PLANT RESPONSE:The development of Oregon boxwood cover following wildfire and clearcutting/ broadcast burning has been recorded for western larch (Larix occidentalis) and Douglas-fir forests in Montana [47]. Oregon boxwood appears to have a varied response to both wildfire and broadcast burning, depending on site [8,16,47]. Stickney tracked first decade postfire succession following a severe fire in western hemlock/ Oregon boxwood habitat type. Oregon boxwood exhibited a steady-state frequency pattern throughout the decade, with little expansion or reduction in distribution within the study site [48,49]. Some have classified Oregon boxwood as "neutral" in its resistance to fire, meaning that it has less than a 12.5 percent frequency increase or decrease when compared to average frequencies of those shrubs in unburned areas [53].
FIRE MANAGEMENT CONSIDERATIONS:Nalley [41] developed models for predicting fuel loading in western redcedar/Oregon boxwood types in northern Idaho. Brown [6] lists bulk densities of some Montana and Idaho habitat types (in which Oregon boxwood is an indicator) for determining fuel depth. Fuel loadings and fire ratings for quaking aspen/Oregon boxwood community types have also been listed [7]. Paxistima myrsinites: References1. Antos, Joseph A.; Shearer, Raymond C. 1980. Vegetation development on disturbed grand fir sites, Swan Valley, northwestern Montana. Res. Pap. INT-251. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 26 p. [7269] 2. Atzet, Thomas; Wheeler, David L. 1984. Preliminary plant associations of the Siskiyou Mountain Province. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 278 p. [9351] 3. Balfour, Patty M. 1989. Effects of forest herbicides on some important wildlife forage species. Victoria, BC: British Columbia Ministry of Forests, Research Branch. 58 p. [12148] 4. Beetle, Alan A. 1962. Range survey in Teton County, Wyoming: Part 2. Utilization and condition classes. Bull. 400. Laramie, WY: University of Wyoming, Agricultural Experiment Station. 38 p. [418] 5. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 6. Brown, James K. 1981. Bulk densities of nonuniform surface fuels and their application to fire modeling. Forest Science. 27(4): 667-683. [13269] 7. Brown, James K.; DeByle, Norbert V. 1989. Effects of prescribed fire on biomass and plant succession in western aspen. Res. Pap. INT-412. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 16 p. [9286] 8. Brown, James K.; Simmerman, Dennis G. 1986. Appraising fuels and flammability in western aspen: a prescribed fire guide. Gen. Tech. Rep. INT-205. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 48 p. [544] 9. Canon, S. K.; Urness, P. J.; DeByle, N. V. 1987. Habitat selection, foraging behavior, and dietary nutrition of elk in burned aspen forest. Journal of Range Management. 40(5): 443-438. [3453] 10. Crane, M. F.; Fischer, William C. 1986. Fire ecology of the forest habitat types of central Idaho. Gen. Tech. Rep. INT-218. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 85 p. [5297] 11. Crouch, Glenn L. 1985. Effects of clearcutting a subalpine forest in central Colorado on wildlife habitat. Res. Pap. RM-258. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 12 p. [8225] 12. Daubenmire, Rexford F.; Daubenmire, Jean B. 1968. Forest vegetation of eastern Washington and northern Idaho. Technical Bulletin 60. Pullman, WA: Washington State University, Agricultural Experiment Station. 104 p. [749] 13. Daubenmire, Rexford. 1980. Mountain topography and vegetation patterns. Northwest Science. 54(2): 146-152. [7896] 14. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 15. Drew, Larry Albert. 1967. Comparative phenology of seral shrub communities in the cedar/hemlock zone. Moscow, ID: University of Idaho. 108 p. Thesis. [9654] 16. Edgerton, Paul J. 1987. Influence of ungulates on the development of the shrub understory of an upper slope mixed conifer forest. In: Provenza, Frederick D.; Flinders, Jerran T.; McArthur, E. Durant, compilers. Proceedings--symposium on plant-herbivore interactions; 1985 August 7-9; Snowbird, UT. Gen. Tech. Rep. INT-222. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station: 162-167. [7411] 17. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 18. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 19. Halverson, Nancy M., compiler. 1986. Major indicator shrubs and herbs on National Forests of western Oregon and southwestern Washington. R6-TM-229. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 180 p. [3233] 20. Hanley, Donald P. 1976. Tree biomass and productivity estimated for three habitat types of northern Idaho. Bull. No. 14. Moscow, ID: University of Idaho, Forest Wildlife and Range Experiment Station. 15 p. [12923] 21. Hann, Wendel J. 1986. Evaluation of site preparation and conifer release treatments in north Idaho shrubfields. In: Baumgartner, David M.; Boyd, Raymond J.; Breuer, David W.; Miller, Daniel L., compilers and eds. Weed control for forest poductivity in the Interior West: Symposium proceedings; 1985 February 5-7; Spokane, WA. Pullman, WA: Washington State University, Cooperative Extension: 115-119. [1074] 22. Hess, Karl; Wasser, Clinton H. 1982. Grassland, shrubland, and forestland habitat types of the White River-Arapaho National Forest. Final Report. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 335 p. [1142] 23. Hitchcock, C. Leo; Cronquist, Arthur. 1961. Vascular plants of the Pacific Northwest. Part 3: Saxifragaceae to Ericaceae. Seattle, WA: University of Washington Press. 614 p. [1167] 24. Hitchcock, C. Leo; Cronquist, Arthur. 1973. Flora of the Pacific Northwest. Seattle, WA: University of Washington Press. 730 p. [1168] 25. Hoffman, George R.; Alexander, Robert R. 1980. Forest vegetation of the Routt National Forest in northwestern Colorado: a habitat classification. Res. Pap. RM-221. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 41 p. [1179] 26. Hoffman, George R.; Alexander, Robert R. 1983. Forest vegetation of the White River National Forest in western Colorado: a habitat type classification. Res. Pap. RM-249. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 36 p. [1178] 27. Hopkins, William E. 1979. Plant associations of south Chiloquin and Klamath Ranger Districts--Winema National Forest. R6-Ecol-79-005. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 96 p. [7339] 28. Komarkova, Vera. 1986. Habitat types on selected parts of the Gunnison and Uncompahgre National Forests. Final Report Contract No. 28-K2-234. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 270 p. [1369] 29. Kruckeberg, A. R. 1982. Gardening with native plants of the Pacific Northwest. Seattle: University of Washington Press. 252 p. [9980] 30. Kuchler, A. W. 1964. United States [Potential natural vegetation of the conterminous United States]. Special Publication No. 36. New York: American Geographical Society. 1:3,168,000; colored. [3455] 31. Lackschewitz, Klaus. 1991. Vascular plants of west-central Montana--identification guidebook. Gen. Tech. Rep. INT-227. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 648 p. [13798] 32. Laursen, Steven B. 1984. Predicting shrub community composition and structure following management disturbance in forest ecosystems of the Intermountain West. Moscow, ID: University of Idaho. 261 p. Dissertation. [6717] 33. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 34. Mauk, Ronald L.; Henderson, Jan A. 1984. Coniferous forest habitat types of northern Utah. Gen. Tech. Rep. INT-170. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 89 p. [1553] 35. McLean, Alastair. 1968. Fire resistance of forest species as influenced by root systems. Journal of Range Management. 22: 120-122. [1621] 36. McLean, Alastair. 1970. Plant communities of the Similkameen Valley, British Columbia. Ecological Monographs. 40(4): 403-424. [1620] 37. Miller, Daniel L.; Kidd, Frank A. 1982. How to write a herbicide prescription for shrub control. Forestry Technical Paper TP-82-6. Lewiston, ID: Potlatch Corporation, Wood Products, Western Division. 12 p. [3390] 38. Monsen, Stephen B. 1983. Plants for revegetation of riparian sites within the Intermountain region. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings of symposia; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 83-89. [9652] 39. Mueggler, Walter F.; Campbell, Robert B., Jr. 1982. Aspen community types on the Caribou and Targhee National Forests in southeastern Idaho. Res. Pap. INT-294. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 32 p. [1713] 40. Munz, Philip A. 1973. A California flora and supplement. Berkeley, CA: University of California Press. 1905 p. [6155] 41. Nalley, Roque Nolan. 1982. Relationships of dead downed woody fuels to site and stand characteristics within the Thuja plicata-Pachistima myrsinites ht. Moscow, ID: University of Idaho. 86 p. Thesis. [10015] 42. Noste, Nonan V.; Bushey, Charles L. 1987. Fire response of shrubs of dry forest habitat types in Montana and Idaho. Gen. Tech. Rep. INT-239. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 22 p. [255] 43. Peek, J. M. 1974. A review of moose food habits studies in North America. Le Naturaliste Canadien. 101: 195-215. [7420] 44. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 45. Rickard, W. H. 1960. The distribution of small mammals in relation to the climax vegetation mosaic in eastern Washington and northern Idaho. Ecology. 41(1): 99-106. [8454] 46. Stark, N. 1983. The nutrient content of Rocky Mountain vegetation: a handbook for estimating nutrients lost through harvest and burning. Misc. Publ. 14. Missoula, MT: University of Montana, School of Forestry, Montana Forest and Conservation Experiment Station. 81 p. [8617] 47. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 48. Stickney, Peter F. 1985. Data base for early postfire succession on the Sundance Burn, northern Idaho. Gen. Tech. Rep. INT-189. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 121 p. [7223] 49. Stickney, Peter F. 1986. First decade plant succession following the Sundance Forest Fire, northern Idaho. Gen. Tech. Rep. INT-197. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 26 p. [2255] 50. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. [2387] 51. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 52. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest. Austin, TX: University of Texas Press. 1104 p. [7707] 53. Vogl, Richard J.; Ryder, Calvin. 1969. Effects of slash burning on conifer reproduction in Montana's Mission Range. Northwest Science. 43(3): 135-147. [8546] 54. Williams, Clinton K.; Lillybridge, Terry R. 1983. Forested plant associations of the Okanogan National Forest. R6-Ecol-132b. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Region. 116 p. [2566] 55. Wittinger, W. T.; Pengelly, W. L.; Irwin, L. L.; Peek, J. M. 1977. A 20-year record of shrub succession in logged areas in the cedar- hemlock zone of northern Idaho. Northwest Science. 51(3): 161-171. [6828] 56. Young, Vernon A.; Payne, Gene F. 1948. Utilization of "key" browse species in relation to proper grazing practices in cutover western white pine lands in northern Idaho. Journal of Forestry. 46: 35-40. [2683] 57. Pengelly, William Leslie. 1966. Ecological effects of slash-disposal fires on the Coeur d'Alene National Forest, Idaho. Missoula, MT: U.S. Department of Agriculture, Forest Service, Northern Region. 23 p. [174] 58. Hickman, James C., ed. 1993. The Jepson manual: Higher plants of California. [21992] 59. Kartesz, John T.; Meacham, Christopher A (1999). Synthesis of the North American flora (Windows Version 1.0), [CD-ROM] [36745] Paxistima myrsinites Index
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