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Introductory

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
ABBREVIATION : SYMLON SYNONYMS : none SCS PLANT CODE : SYLO COMMON NAMES : longflower snowberry desert snowberry TAXONOMY : The fully documented scientific name of longflower snowberry is Symphoricarpos longiflorus Gray. There are no recognized varieties or forms. LIFE FORM : Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Nancy E. McMurray/August 1986 LAST REVISED BY AND DATE : Nancy E. McMurray/December 1986 AUTHORSHIP AND CITATION : McMurray, Nancy E. 1986. Symphoricarpos longiflorus. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
GENERAL DISTRIBUTION : Longflower snowberry occurs throughout Nevada and adjacent southeastern Oregon, south to southern California and northern Baja, and eastward to eastern Utah, northwestern Colorado, New Mexico, and western Texas. In Utah, this species is found primarily south of the 40th parallel, except along the Green River where it extends north into Daggett County; in Nevada it is found everywhere except the northeast corner [4]. ECOSYSTEMS : FRES29 Sagebrush FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper STATES : AZ CA CO ID NV NM OR TX UT WY ADMINISTRATIVE UNITS : ARCH CANY CARE CACA DEVA DINO GLCA GRCA GUMO LAME NABR ZION BLM PHYSIOGRAPHIC REGIONS : 5 Columbia Plateau 6 Upper Basin and Range 7 Lower Basin and Range 10 Wyoming Basin 12 Colorado Plateau 13 Rocky Mountain Piedmont KUCHLER PLANT ASSOCIATIONS : K023 Juniper - pinyon woodland K037 Mountain-mahogany - oak scrub K038 Great Basin sagebrush K055 Sagebrush steppe SAF COVER TYPES : 220 Rocky Mountain juniper 237 Interior ponderosa pine 238 Western juniper 239 Pinyon - juniper SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Habitat types and community types using longflower snowberry as a climax indicator are limited. The following community types have been described for the northern desert shrub and pinyon-juniper zones in central Nevada [2,3]: (1) big sagebrush (Artemisia tridentata)/ serviceberry (Amelanchier pallida)/longflower snowberry; (2) big sagebrush/longflower snowberry/bluebunch wheatgrass (Pseudoroegneria spicata); (3) singleleaf pinyon (Pinus monophylla)/curlleaf mountain-mahogany (Cercocarpus ledifolius)/longflower snowberry/ arrowleaf balsamroot (Balsamorhiza sagittata); (3) longflower snowberry/ big sagebrush/cheatgrass (Bromus tectorum)/woolly wyethia (Wyethia mollis); (4) longflower snowberry/big sagebrush/serviceberry/ Idaho fescue (Festuca idahoensis); (5) longflower snowberry/Great Basin wildrye (Leymus cinereus).

VALUE AND USE

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Longflower snowberry is an important forage species on the xeric sites it occupies and is utilized by both big game and livestock. Over much of its range, it is often the first browse species to leaf out. Plants are easily accessible due to their low growth form and are moderately tolerant to grazing pressure [15,20]. The seeds are eaten by a number of species of birds, especially grouse, pheasant, and quail. The Rocky Mountain pika and various ground squirrels and chipmunks also eat the seeds [21]. PALATABILITY : Palatability of longflower snowberry is highest in the early spring and declines throughout the summer and fall. This species is usually more palatable in the Intermountain region than on more southern or western ranges [20]. Palatability of longflower snowberry to livestock and wildlife species in Utah has been rated as follows [5]: .NS Cattle Fair Sheep Good Horses Fair Elk Fair Mule deer Good Small mammals Good Small nongame birds Good Upland game birds Fair Waterfowl Poor NUTRITIONAL VALUE : Longflower snowberry has been rated as fair in both energy and protein value [5]. COVER VALUE : The degree to which longflower snowberry provides environmental protection during one or more seasons for wildlife species in Utah is as follows [5]: Elk Poor Mule deer Poor Small mammals Good Small nongame birds Fair Upland game birds Fair Waterfowl Poor VALUE FOR REHABILITATION OF DISTURBED SITES : Longflower snowberry is a very useful shrub for establishing cover on raw and severe sites. It grows well in areas where few other species are adapted, but it does not have a wide range of adaptability [13,15]. Once established longflower snowberry is very persistent. Sites can be revegetated using wildlings or transplants. Transplants are easily obtained by pulling up small rooted portions of plants in the early spring before leaf growth has begun. Propagation via stem cuttings has also been successful [7]. Direct seeding is generally recommended in the fall or winter in the following types: big sagebrush, blackbrush, pinyon-juniper, and aspen. Commercial seed may be dried or cleaned. Seeds have a pronounced domancy; acid treatments break down the seed coat. Stratification should be employed for spring planting [15]. OTHER USES AND VALUES : Snowberries are highly regarded for the beauty of their foliage and fruits; many species are widely grown as ornamentals [20]. MANAGEMENT CONSIDERATIONS : NO-ENTRY

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
GENERAL BOTANICAL CHARACTERISTICS : Longflower snowberry is a low, spreading, deciduous shrub with somewhat drooping branchlets [21]. Heights range from 3.3 to 5 feet (1-1.5 m). Persistent, smaller branches tend to give plants a thorny appearance [4]. This species exhibits wide ecotypic variation in leaf and twig pubescence [9]. Longflower snowberry is monoecious. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Information concerning regenerative processes in longflower snowberry is scant. Morphologically similar mountain snowberry regenerates via rhizomes, basal sprouts, stolons, and seed [17,18]. Although many members of the Symphoricarpos genus are highly rhizomatous, the extent to which this habit is developed in longflower snowberry is unclear. The general pattern for seed production in the Symphoricarpos genus has been described by Pelton [14] for western snowberry (S. occidentalis) and apparently also applies to longflower snowberry [15]. Fruits are white, berrylike drupes containing two, seedlike nutlets which ripen in the fall. Since no abscision layer is formed on the pedicle, fruits can persist on shrubs for up to two seasons before falling to the ground. Nutlets have a double dormancy which delays germination and establishment [6]. Seed coat restrictions are eliminated through storage in warm, moist soil; afterripening and development of the embryo occur during one or two winters in the soil. Seeds do not remain viable for extended periods in nature. Birds or mammals are probably the main dispersal agents; field mice scatter seeds, and plants are widely distributed from their caches. Germination begins in the early spring as soon as the soil thaws. First-year seedlings are quite sensitive to drought. SITE CHARACTERISTICS : Longflower snowberry typically occurs on open, rocky slopes and washes at elevations ranging between 4,500 and 7,500 feet (1,372 and 2,287 m). It occurs on sites more xeric than those occupied by mountain snowberry [17]. Longflower snowberry is usually a component of pinyon-juniper communities [15] but extends down into the sagebrush-grassland and cold desert on sites receiving 8 to 9 inches (20-23 cm) minimum annual precipitation. In the mountain brush and ponderosa pine zones, the ranges of longflower snowberry and mountain snowberry sometimes overlap [4]. Elevational ranges for some western states are listed below [4]. From 5,000 to 5,000 feet (1,524-1,524 m) in CO 5,000 to 7,400 feet (1,524-2,256 m) in UT SUCCESSIONAL STATUS : Longflower snowberry is present in nonforested and forested climax communities throughout its range. Self-perpetuating stands are indicative of climax conditions [3]. SEASONAL DEVELOPMENT : Flowering generally occurs from May through July [5].

FIRE ECOLOGY

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
FIRE ECOLOGY OR ADAPTATIONS : Information concerning the fire ecology of longflower snowberry is scant; however, resprouting has been documented following burning in Nevada [10]. As a genus, snowberries are moderately resistent to fire mortality. Most species resprout via basal buds or rhizomes [19,23,24]. Although plant response is variable, postfire recovery rates are moderately rapid on most sites. POSTFIRE REGENERATION STRATEGY : Small shrub, adventitious-bud root crown Rhizomatous shrub, rhizome in soil Secondary colonizer - off-site seed

FIRE EFFECTS

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
IMMEDIATE FIRE EFFECT ON PLANT : Fire usually completely top-kills longflower snowberry. It is probably moderately resistant to fire mortality due to perennating buds located on root crowns and rhizomes. Survival may be highly variable, however, and appears to be significantly influenced by fire intensity and severity [18.25]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Fire response of longflower snowberry has not been well documented. Following a spring burn in a sagebrush-grassland community in Nevada, all reexamined plants sprouted; however, sprout origin and vigor were not discussed [10]. (For fire response information on a morphologically similar species, see the FEIS write-up on mountain snowberry). DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Symphoricarpos longiflorus | Longflower Snowberry
REFERENCES : 1. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 2. Blackburn, Wilbert H.; Eckert, Richard E., Jr.; Tueller, Paul T. 1969. Vegetation and soils of the Coils Creek Watershed. R-48. Reno, NV: University of Nevada, Agricultural Experiment Station. 80 p. In cooperation with: U.S. Department of the Interior, Bureau of Land Management. [455] 3. Blackburn, Wilbert H.; Eckert, Richard E., Jr.; Tueller, Paul T. 1971. Vegetation and soils of the Rock Springs Watershed. R-83. Reno, NV: University of Nevada, Agricultural Experiment Station. 116 p. In cooperation with: U.S. Department of the Interior, Bureau of Land Management. [457] 4. Cronquist, Arthur; Holmgren, Arthur H.; Holmgren, Noel H.; [and others]. 1984. Intermountain flora: Vascular plants of the Intermountain West, U.S.A. Vol. 4. Subclass Asteridae, (except Asteraceae). New York: The New York Botanical Garden. 573 p. [718] 5. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 6. Evans, Keith E. 1974. Symphoricarpos Duham. snowberry. In: Schopmeyer, C. S., ed. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 787-790. [7759] 7. Everett, Richard L.; Meeuwig, Richard O.; Robertson, Joseph H. 1978. Propagation of Nevada shrubs by stem cutting. Journal of Range Management. 31(6): 426-429. [894] 8. Goodrich, Sherel. 1986. Vascular plants of the Desert Experimental Range, Millard County, Utah. Gen. Tech. Rep. INT-209. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 72 p. [1033] 9. Jones, George Neville. 1940. A monograph of the genus Symphoricarpos. Journal of the Arnold Arboretum. 21: 201-253. [13499] 10. Klebenow, Donald A.; Beall, Robert C. 1977. Fire impacts on birds and mammals on Great Basin rangelands. In: [Source unknown]. Reno, NV: University of Nevada, Division of Renewable Natural Resources: 59-62. On file with: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Lab, Missoula, MT. [1348] 11. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 12. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 13. Monsen, Stephen B.; Christensen, Donald R. 1975. Woody plants for rehabilitating rangelands in the Intermountain Region. In: Stutz, Howard C., ed. Wildland shrubs: Proceedings--symposium and workshop; 1975 November 5-7; Provo, UT. Provo, UT: Brigham Young Univeristy: 72-119. [1680] 14. Pelton, John. 1953. Studies on the life-history of Symphoricarpos occidentalis Hook, in Minnesota. Ecological Monographs. 23(1): 17-39. [11957] 15. Plummer, A. Perry; Christensen, Donald R.; Monsen, Stephen B. 1968. Restoring big-game range in Utah. Publ. No. 68-3. Ephraim, UT: Utah Division of Fish and Game. 183 p. [4554] 16. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 17. Stanton, Frank. 1974. Wildlife guidelines for range fire rehabilitation. Tech. Note 6712. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 90 p. [2221] 18. Stickney, Peter F. 1980. Data base for post-fire succession, first 6 to 9 years, in Montana larch-fir forests. Gen. Tech. Rep. INT-62. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station. 133 p. [6583] 19. Tisdale, E. W.; Hironaka, M. 1981. The sagebrush-grass region: a review of the ecological literature. Bull. 33. Moscow, ID: University of Idaho, Forest, Wildlife and Range Experiment Station. 31 p. [2344] 20. U.S. Department of Agriculture, Forest Service. 1937. Range plant handbook. Washington, DC. 532 p. [2387] 21. Vines, Robert A. 1960. Trees, shrubs, and woody vines of the Southwest. Austin, TX: University of Texas Press. 1104 p. [7707] 22. Wood, Benjamin W. 1965. Revision of Symphoricarpos (Caprifoliaceae) in Utah. Utah Academy Proceedings. 42(II): 203-213. [13497] 23. Wright, Henry A.; Neuenschwander, Leon F.; Britton, Carlton M. 1979. The role and use of fire in sagebrush-grass and pinyon-juniper plant communities: A state-of-the-art review. Gen. Tech. Rep. INT-58. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Statio. 48 p. [2625] 24. Young, Richard P. 1983. Fire as a vegetation management tool in rangelands of the Intermountain Region. In: Monsen, Stephen B.; Shaw, Nancy, compilers. Managing Intermountain rangelands--improvement of range and wildlife habitats: Proceedings; 1981 September 15-17; Twin Falls, ID; 1982 June 22-24; Elko, NV. Gen. Tech. Rep. INT-157. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Forest and Range Experiment Station: 18-31. [2681] 25. Zschaechner, Greg A. 1985. Studying rangeland fire effects: a case study in Nevada. In: Sanders, Ken; Durham, Jack, eds. Rangeland fire effects: Proceedings of the symposium; 1984 November 27-29; Boise, ID. Boise, ID: U.S. Department of the Interior, Bureau of Land Managment, Idaho State Office: 66-84. [2692]

Index

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