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Introductory

SPECIES: Carex heliophila | Sun Sedge
ABBREVIATION : CARHEL SYNONYMS : Carex pensylvanica Lam. var. digyna Boeck. SCS PLANT CODE : CAHE5 COMMON NAMES : sun sedge TAXONOMY : The currently accepted scientific name of sun sedge is Carex heliophila MacKenz [11,13]. LIFE FORM : Graminoid FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Kathy Ahlenslager March 1988 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Ahlenslager, Kathy. 1988. Carex heliophila. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Carex heliophila | Sun Sedge
GENERAL DISTRIBUTION : Sun sedge's range extends from Ontario to Alberta, south to Ohio, Missouri, Kansas, and New Mexico. Sun sedge also occurs in New York [11]. ECOSYSTEMS : FRES20 Douglas-fir FRES21 Ponderosa pine FRES29 Sagebrush FRES34 Chaparral - mountain shrub FRES35 Pinyon - juniper FRES36 Mountain grasslands FRES38 Plains grasslands FRES39 Prairie STATES : CO IL IN IA KS MI MN MO MT NE NM NY ND OH SD WI WY AB MB ON SK ADMINISTRATIVE UNITS : BADL BICA DETO FLFO GRSA MEVE PIPE ROMO THRO WICA YELL BLM PHYSIOGRAPHIC REGIONS : 7 Lower Basin and Range 8 Northern Rocky Mountains 9 Middle Rocky Mountains 10 Wyoming Basin 11 Southern Rocky Mountains 12 Colorado Plateau 13 Rocky Mountain Piedmont 14 Great Plains 15 Black Hills Uplift 16 Upper Missouri Basin and Broken Lands KUCHLER PLANT ASSOCIATIONS : K012 Douglas-fir forest K016 Eastern ponderosa forest K023 Juniper - pinyon woodland K037 Mountain-mahogany - oak scrub K038 Great Basin sagebrush K055 Sagebrush steppe K056 Wheatgrass - needlegrass shrubsteppe K063 Foothills prairie K064 Grama - needlegrass - wheatgrass K065 Grama - buffalograss K066 Wheatgrass - needlegrass K067 Wheatgrass - bluestem - needlegrass K070 Sandsage - bluestem prairie K074 Bluestem prairie K081 Oak savanna SAF COVER TYPES : NO-ENTRY SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Sun sedge is a codominant or indicator species in many habitat types in ponderosa pine (Pinus ponderosa) forests and plains grasslands. It is a codominant with buffalograss (Buchloe dactyloides), oatgrass (Danthonia parryi), blue grama (Bouteloua gracilis), Gambel oak (Quercus gambelii), creeping juniper (Juniperus horizontalis), mountain-mahogany (Cercocarpus montanus), Idaho fescue (Festuca idahoensis), and needle-and-thread (Stipa comata).

VALUE AND USE

SPECIES: Carex heliophila | Sun Sedge
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : Sun sedge is an important forage at the beginning of the grazing season and after summer rains [13]. It provides abundant early forage, especially in the drier areas of prairie grasslands [21]. In South Dakota and Wyoming, the ponderosa pine/sun sedge habitat type is used as a spring-fall range for livestock and spring-summer range for large mammals [13]. At Wind Cave National Park, South Dakota, a grazing study showed that sun sedge increased in response to grazing by prairie dogs for the first 3 years after introduction of the prairie dogs. This increase may have been the result of sun sedge's tolerance to grazing combined with the decreased abundance of mid-sized grass competitors [4]. PALATABILITY : Sun sedge is rated 50 to 80 percent palatable to livestock [13]. The palatability of sun sedge to livestock and wildlife species in several western states has been rated as follows [8]: CO MT ND WY Cattle Fair Fair Good Fair Sheep Fair Fair Good Fair Horses Good Good Good Good Pronghorn ---- ---- Fair ---- Mule deer ---- ---- Fair ---- White-tailed deer ---- ---- Poor ---- NUTRITIONAL VALUE : Compared with other graminoids, sun sedge is rated fair in energy value and protein value [8]. COVER VALUE : The degree to which sun sedge provides environmental protection in North Dakota during one or more seasons is rated as fair for pronghorn but poor for white-tailed deer [8]. VALUE FOR REHABILITATION OF DISTURBED SITES : Sun sedge was rated for several factors relating to its use in rehabilitation work in several western states [8]: CO MT ND WY Potential biomass Low Low Low Low production Erosion control potential Medium Medium Medium Low Establishment requirements ---- ---- High Low Short-term revegetation ---- Low Low Low potential Long-term revegetation ---- Low Medium Medium potential OTHER USES AND VALUES : NO-ENTRY MANAGEMENT CONSIDERATIONS : The presence of wildlife affects the production of sun sedge [3]. There is also evidence that heavy grazing in areas with sun sedge may reduce the production of this species [14]. The impact of prairie dog activity was studied in a mixed-grass prairie community in southwestern South Dakota. Over time plant species diversity decreased. Sun sedge was one of the "decreasers" [3].

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Carex heliophila | Sun Sedge
GENERAL BOTANICAL CHARACTERISTICS : Sun sedge is a low-growing, strongly rhizomatous, cool-season, nonmychorrhizal, native perennial. Plants grow from slender rootstocks and form tufts or bunches. Culms (stems) range up to 11 inches (3 dm) in height [8,11,13,21]. RAUNKIAER LIFE FORM : Geophyte REGENERATION PROCESSES : Although sun sedge produces seeds, most regeneration is through well-developed, creeping rhizomes [11]. SITE CHARACTERISTICS : Sun sedge occurs in prairies, pastures, plains, and hills from 4,000 to 9,000 feet (1,200-2,600 m) in elevation [11,13]. It usually is a minor component of prairie vegetation. On dry ridges, sun sedge may comprise 5 to 10 percent of the cover, but in some areas it may comprise approximately 30 to 50 percent of the plants present [11]. Elevational ranges in three western states vary as follows [1]: from 4,400 to 9,500 feet (1,300-2,800 m) in CO 3,700 to 6,200 feet (1,100-1,800 m) in MT 3,400 to 8,500 feet (1,000-2,500 m) in WY Sun sedge is found in openings in ponderosa pine forests with sandy loam to sandy clay soils with pH's ranging from 6.0 to 7.6 [14]. The growth of sun sedge on soils from several western states is rated as follows [8]: CO MT ND WY Gravel Fair Fair Fair Good Sand Fair Good Good Good Sandy loam Good Good Good Good Loam Good Good Good Good Clay loam Fair Fair Fair Fair Clay Poor Poor Poor Poor Dense clay Poor Poor Poor Poor SUCCESSIONAL STATUS : Obligate climax species Sun sedge is a climax species in many habitat types [1,5,14]. SEASONAL DEVELOPMENT : Sun sedge is a cool-season species that begins flowering in April in Montana and in June in North Dakota and Wyoming. Blooming ends in July [8]. Fruits mature in late May or in June. After this time plants are more or less covered by grasses [21].

FIRE ECOLOGY

SPECIES: Carex heliophila | Sun Sedge
FIRE ECOLOGY OR ADAPTATIONS : Sedges tolerate fire very well. Fire probably stimulates plants of this rhizomatous species to initiate new shoots at primodial regions of the root system. Although summer grass fires harm warm-season species, they favor cool-season ones like sun sedge. POSTFIRE REGENERATION STRATEGY : Rhizomatous herb, rhizome in soil

FIRE EFFECTS

SPECIES: Carex heliophila | Sun Sedge
IMMEDIATE FIRE EFFECT ON PLANT : Sedges generally tolerate fire very well. The season of a fire has the greatest effect on these plants [22]. Fires usually consume dry vegetation to ground level. Because sun sedge is relatively leafy, quite a bit of heat may be transferred to belowground plant parts [22]. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : In central Alberta annual early spring burns for 25 to 30 years resulted in an increase in sun sedge [2]. However, a study of the effect of spring burning on vegetation in Kansas pastures showed that even though the number of sun sedge plants increased after burning, they decreased postfire years 3 and 4. At the same time, the number of plants increased somewhat on unburned areas [12]. On a mesic mixed prairie in western North Dakota with average annual precipitation of 16 inches (41 cm), the effects of three wildfires (May, August, and September) were studied. After a hot, late May fire the frequency of plants was the same or higher as that before the fire, while a fire in late summer harmed plants [9,20]. Since mowing and grazing, as well as fire, defoliate plants, the effects of mowing and grazing are discussed here also. A 3-year study on a mixed-grass prairie in southwestern North Dakota showed a greater increase in sun sedge and threadleaf sedge (Carex filifolia) in an exclosure than on adjacent grazed plots. Upland sedges are considered to be increasers; however the results of this study contradict this. Thus, the reaction of upland sedges to grazing should be reevaluated [3]. On the eastern edge of the badlands of South Dakota in the Pine Ridge Indian Reservation near Wanblee, species composition of a relict mixed-grass prairie on one mesa was compared with that of two mesas which were grazed and mowed. The dominant species on the protected mesa included bluebunch wheatgrass (Pseudorogneria spicata), needleleaf sedge (Carex eleocharis), sun sedge, threadleaf sedge, blue grama, and needle-and-thread [7]. The protected area produced a mixed-grass association dominated by bluebunch wheatgrass and dry-land sedges, whereas combined mowing and grazing resulted in a short-grass association dominated by blue grama grass. The results of this study show that mowing and grazing greatly reduce the amount of sedges [16]. The relationship between prescribed fire and plains rough fescue (Festuca scabrella) and western porcupine grass (Stipa spartea) communities in the aspen parkland of central Alberta was studied. This fire-climax grassland is well adapted to fire. With settlement in the early 1900's and the cessation of fires, trees and shrubs invaded the prairie. Annual early spring burns for 25 to 30 years eliminated a few species, increased the diversity of herbaceous species, and maintained forest cover at the same levels as presettlement. During this period sun sedge increased on the burned areas. Annual burning created a drier microenvironment than normally found on the fescue prairie and favored sun sedge [2,16]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Carex heliophila | Sun Sedge
REFERENCES : 1. Alexander, Robert R. 1986. Classification of the forest vegetation of Wyoming. Res. Note RM-466. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 10 p. [304] 2. Anderson, Howard G.; Bailey, Arthur W. 1980. Effects of annual burning on grassland in the aspen parkland of east-central Alberta. Canadian Journal of Botany. 58: 985-996. [3499] 3. Archer, Steven R. 1983. Plant community structure, competitive interactions and water relations as influenced by herbivores. Fort Collins, CO: Colorado State University. 114 p. Dissertation. [338] 4. Archer, Steve; Garrett, M. G.; Detling, James K. 1987. Rates of vegetation change associated with prairie dog (Cynomys ludovicianus) grazing in North American mixed-grass prairie. Vegetatio. 72: 159-166. [2864] 5. Baker, William L. 1984. A preliminary classification of the natural vegetation of Colorado. Great Basin Naturalist. 44(4): 647-676. [380] 6. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 7. Brand, M. D.; Goetz, H. 1978. Secondary succession of a mixed grass community in southwestern North Dakota. Annual Proceedings of the North Dakota Academy of Science. 32(2): 67-78. [7512] 8. Dittberner, Phillip L.; Olson, Michael R. 1983. The plant information network (PIN) data base: Colorado, Montana, North Dakota, Utah, and Wyoming. FWS/OBS-83/86. Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 786 p. [806] 9. Dix, Ralph L. 1960. The effects of burning on the mulch structure and species composition of grasslands in western North Dakota. Ecology. 41(1): 49-56. [808] 10. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 11. Great Plains Flora Association. 1986. Flora of the Great Plains. Lawrence, KS: University Press of Kansas. 1392 p. [1603] 12. Hensel, R. L. 1923. Effect of burning on vegetation in Kansas pastures. Journal of Agricultural Research. 23(8): 631-644. [1131] 13. Hermann, Frederick J. 1970. Manual of the Carices of the Rocky Mountains and Colorado Basin. Agric. Handb. 374. Washington, DC: U.S. Department of Agriculture, Forest Service. 397 p. [1139] 14. Hoffman, George R.; Alexander, Robert R. 1987. Forest vegetation of the Black Hills National Forest of South Dakota and Wyoming: a habitat type classification. Res. Pap. RM-276. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 48 p. [1181] 15. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 16. Larson, Floyd; Whitman, Warren. 1942. A comparison of used and unused grassland mesas in the Badlands of South Dakota. Ecology. 23: 438-445. [4000] 17. Lyon, L. Jack; Stickney, Peter F. 1976. Early vegetal succession following large northern Rocky Mountain wildfires. In: Proceedings, Tall Timbers fire ecology conference and Intermountain Fire Research Council fire and land management symposium; 1974 October 8-10; Missoula, MT. No. 14. Tallahassee, FL: Tall Timbers Research Station: 355-373. [1496] 18. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 19. Ross, Robert L.; Hunter, Harold E. 1976. Climax vegetation of Montana based on soils and climate. Bozeman, MT: U.S. Department of Agriculture, Soil Conservation Service. 64 p. [2028] 20. Ferguson, Dennis E.; Boyd, Raymond J. 1988. Bracken fern inhibition of conifer regeneration in northern Idaho. Ogden, UT: U.S. Department of Agriculture, Forest Service, Intermountain Research Station. 11 p. [2834] 21. Weaver, J. E.; Fitzpatrick, T. J. 1934. The prairie. Ecological Monographs. 4(2): 111-295. [2464] 22. Wright, Henry A. 1978. Use of fire to manage grasslands of the Great Plains: central and southern Great Plains. In: Hyder, Donald N., ed. Proceedings, 1st international rangelands congress; 1978 August 14-18; Denver, CO. Denver, CO: Society for Range Management: 694-696. [2615] 23. Wright, Henry A.; Bailey, Arthur W. 1982. Fire ecology: United States and southern Canada. New York: John Wiley & Sons. 501 p. [2620] 24. Bernard, Stephen R.; Brown, Kenneth F. 1977. Distribution of mammals, reptiles, and amphibians by BLM physiographic regions and A.W. Kuchler's associations for the eleven western states. Tech. Note 301. Denver, CO: U.S. Department of the Interior, Bureau of Land Management. 169 p. [434] 25. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 26. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 27. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 28. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573]

Index

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