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Introductory

SPECIES: Cornus racemosa | Gray Dogwood
ABBREVIATION : CORRAC SYNONYMS : Cornus foemina ssp. racemosa SCS PLANT CODE : CORA6 COMMON NAMES : gray dogwood gray-stemmed dogwood panicled dogwood TAXONOMY : The currently accepted scientific name of gray dogwood is Cornus racemosa Lam. [16]. Some authorities consider C. racemosa a subspecies of Cornus foemina [8,10]. Little [17], however, considers it a distinct species. LIFE FORM : Tree, Shrub FEDERAL LEGAL STATUS : No special status OTHER STATUS : NO-ENTRY COMPILED BY AND DATE : Milo Coladonato, July 1993 LAST REVISED BY AND DATE : NO-ENTRY AUTHORSHIP AND CITATION : Coladonato, Milo. 1993. Cornus racemosa. In: Remainder of Citation

DISTRIBUTION AND OCCURRENCE

SPECIES: Cornus racemosa | Gray Dogwood
GENERAL DISTRIBUTION : Gray dogwood's main range is from Maine and southern Ontario; south through New England and Pennyslvania; and west to Ohio, Indiana, Michigan, Illinois, Missouri, Iowa, and Minnesota. Its southern range is from the Appalachian Mountains of Virginia to northern Arkansas. Disjunct populations also occur in North Carolina, Tennessee, Kentucky, North and South Dakota, and Nebraska [2,10,17,30]. ECOSYSTEMS : FRES10 White - red - jack pine FRES11 Spruce - fir FRES15 Oak - hickory FRES17 Elm - ash - cottonwood FRES18 Maple - beech - birch FRES19 Aspen - birch STATES : AR CT DE IL IN IA KY ME MD MA MI MN MO NE NH NJ NY NC ND OH OK PA RI SC SD TN VT VA WV WI MB ON PQ ADMINISTRATIVE UNITS : ALPO APIS BLRI CATO CUVA DEWA EFMO GWCA INDU ISRO PIPE SARA SHEN SHIL VOYA WICR BLM PHYSIOGRAPHIC REGIONS : NO-ENTRY KUCHLER PLANT ASSOCIATIONS : K093 Great Lakes spruce - fir forest K095 Great Lakes pine forest K096 Northeastern spruce - fir forest K099 Maple - basswood forest K100 Oak - hickory forest K102 Beech - maple forest K101 Elm - ash forest K103 Mixed mesophytic forest K104 Appalachian oak forest K106 Northern hardwoods K108 Northern hardwoods - spruce forest K110 Northeastern oak - pine forest SAF COVER TYPES : 17 Pin cherry 19 Gray birch - red maple 20 White pine - northern red oak - red maple 21 Eastern white pine 22 White pine - hemlock 23 Eastern hemlock 24 Hemlock - yellow birch 25 Sugar maple - beech - yellow birch 26 Sugar maple - basswood 27 Sugar maple 28 Black cherry - maple 30 Red spruce - yellow birch 31 Red spruce - sugar maple - beech 32 Red spruce 33 Red spruce - balsam fir 34 Red spruce - Fraser fir 35 Paper birch - red spruce - balsam fir 37 Northern white-cedar 39 Black ash - American elm - red maple 42 Bur oak 43 Bear oak 44 Chestnut oak 52 White oak - black oak - northern red oak 53 White oak 55 Northern red oak 57 Yellow-poplar 58 Yellow-poplar - eastern hemlock 59 Yellow-poplar - white oak - northern red oak 60 Beech - sugar maple 62 Silver maple - American elm 64 Sassafras - persimmon 107 White spruce 108 Red maple 110 Black oak SRM (RANGELAND) COVER TYPES : NO-ENTRY HABITAT TYPES AND PLANT COMMUNITIES : Gray dogwood is one of the dominant shrubs in the oak-hickory (Quercus-Carya) forests of the northeastern United States. Common codominants include maple-leaved viburnum (Viburnum acerifolium) and Virginia creeper (Parthenocissus quinquefolia). Other common associates of gray dogwood include American hazel (Corylus americana), beaked hazelnut (C. cornuta), chokecherry (Prunus virginiana), smooth sumac (Rhus glabra), and red raspberry (Rubus idaeus) [3,23,26].

VALUE AND USE

SPECIES: Cornus racemosa | Gray Dogwood
WOOD PRODUCTS VALUE : NO-ENTRY IMPORTANCE TO LIVESTOCK AND WILDLIFE : In Missouri, Ohio, and Illinois, gray dogwood is one of the most important forage plants for white-tailed deer [5,19,24]. The seeds and buds are a favorite food for ring-necked pheasant and northern bobwhite in southern Michigan [28]. Gray dogwood thickets provide cover for a variety of birds and mammals [2,6,14]. PALATABILITY : NO-ENTRY NUTRITIONAL VALUE : NO-ENTRY COVER VALUE : NO-ENTRY VALUE FOR REHABILITATION OF DISTURBED SITES : Gray dogwood is well adapted for revegetating disturbed sites. It is easily established by direct seedling and grows rapidly [11,29]. It has been successfully planted for revegetating highway corridors in Wisconsin and coal mine spoils in the eastern United States [11,28,29]. OTHER USES AND VALUES : Gray dogwood has been planted for ornamental purposes because of its showy flowers, fruits, and attractive fall coloring [2]. MANAGEMENT CONSIDERATIONS : NO-ENTRY

BOTANICAL AND ECOLOGICAL CHARACTERISTICS

SPECIES: Cornus racemosa | Gray Dogwood
GENERAL BOTANICAL CHARACTERISTICS : Gray dogwood is a native, deciduous, rhizomatous shrub, usually from 4 to 10 feet (1.2-3.0 m) high. It sometimes becomes a small tree up to 27 feet (8 m) high [17]. It has ascending stems and branches that often form impenetrable dome-shaped clusters or thickets [4]. The leaves are 2.5 to 4.0 inches (6.0-10 cm) long, and the flowers are borne in open, irregular cymes. The individual fruits enclose a single stone and occur in clusters [2,6,14]. RAUNKIAER LIFE FORM : Phanerophyte REGENERATION PROCESSES : Gray dogwood reproduces both sexually and asexually. It begins producing seed at about 4 to 5 years of age and produces an abundant amount of seed every year. Gray dogwood reproduces vegetatively by sprouting from underground rhizomes [22,29]. SITE CHARACTERISTICS : Gray dogwood grows on a variety of sites within its range. It is found in meadows, open woodlands, riparian zones, along roadsides, and forest margins. It grows best on rich, moist, well-drained soils, but will also grow on mineral-rich limestone bedrock and rock outcroppings. In Appalachian oak-hickory forests, it usually occurs on open ridgetops and south- and west-facing slopes [1,10,16]. SUCCESSIONAL STATUS : Facultative Seral Species Gray dogwood is an early to mid-seral species [12,20]. It is most common in understories of mixed, open forests and grows best in moderate to full sunlight [18]. In southwestern Wisconsin, aboveground growth rates of gray dogwood were greater in open habitats than in forest understories [12]. SEASONAL DEVELOPMENT : Gray dogwood flowers from May through July, with fruits maturing from August through October [4,14]. Leaves emerge in early April and abscise in late October [13].

FIRE ECOLOGY

SPECIES: Cornus racemosa | Gray Dogwood
FIRE ECOLOGY OR ADAPTATIONS : Postfire regeneration strategies of grey dogwood are not documented in the literature. It probably survives fire by sprouting from rhizomes. It also produces an abundance of soil-stored seed [23], which may germinate after fire. POSTFIRE REGENERATION STRATEGY : Rhizomatous shrub, rhizome in soil Ground residual colonizer (on-site, initial community) Initial-offsite colonizer (off-site, initial community)

FIRE EFFECTS

SPECIES: Cornus racemosa | Gray Dogwood
IMMEDIATE FIRE EFFECT ON PLANT : Aboveground plant parts are often killed by fire [25,26]. The underground rhizomes probably survive all but severe fires that remove duff and heat the upper soil for extended periods of time. DISCUSSION AND QUALIFICATION OF FIRE EFFECT : NO-ENTRY PLANT RESPONSE TO FIRE : Percent cover of native shrubs, including gray dogwood, decreased following fire in a bur oak (Quercus macrocarpa) savanna in east-central Minnesota [26]. In a study of postfire plant response in four plant communities in central New York, gray dogwood frequency on 17 burned plots averaged 62 percent at postfire year 1. Frequency on unburned plots was 62 percent [25]. DISCUSSION AND QUALIFICATION OF PLANT RESPONSE : NO-ENTRY FIRE MANAGEMENT CONSIDERATIONS : NO-ENTRY

REFERENCES

SPECIES: Cornus racemosa | Gray Dogwood
REFERENCES : 1. Braun, E. Lucy. 1961. The woody plants of Ohio. Columbus, OH: Ohio State University Press. 362 p. [12914] 2. Brinkman, Kenneth A. 1974. Cornus L. dogwood. In: Schopmeyer, C. S., technical coordinator. Seeds of woody plants in the United States. Agric. Handb. 450. Washington, DC: U.S. Department of Agriculture, Forest Service: 336-342. [7593] 3. Buell, Murray F.; Facey, Vera. 1960. Forest-prairie transition west of Itasca Park, Minnesota. Bulletin of the Torrey Botanical Club. 87(1): 46-58. [14171] 4. Chapman, William K.; Bessette, Alan E. 1990. Trees and shrubs of the Adirondacks. Utica, NY: North Country Books, Inc. 131 p. [12766] 5. Dalke, Paul D. 1941. The use and availability of the more common winter deer browse plants in the Missouri Ozarks. Transactions, 6th North American Wildlife Conference. 6: 155-160. [17044] 6. Duncan, Wilbur H.; Duncan, Marion B. 1988. Trees of the southeastern United States. Athens, GA: The University of Georgia Press. 322 p. [12764] 7. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905] 8. Gill, David S.; Marks, P. L. 1991. Tree and shrub seedling colonization of old fields in central New York. Ecological Monographs. 61(2): 183-205. [14486] 9. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998] 10. Gleason, Henry A.; Cronquist, Arthur. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. 2nd ed. New York: New York Botanical Garden. 910 p. [20329] 11. Harrington, John A. 1989. Major prairie planting on highway corridor to test methods, value of resulting vegetation (Wisconsin). Restoration and Management Notes. 7(1): 31-32. [8069] 12. Harrington, Robin A.; Brown, Becky J.; Reich, Peter B. 1989. Ecophysiol. of exotic & native shrubs in s. WI. I. Rel. of leaf charac. resource availability, & phenol. to seasonal patterns of carbon gain. Oecologia. 80: 356-367. [9241] 13. Harrington, Robin A.; Brown, Becky J.; Reich, Peter B. 1989. Ecophysiol. of exotic & native shrubs in s. WI. I. Rel. of leaf charac. resource availability, & phenol. to seasonal patterns of carbon gain. Oecologia. 80: 356-367. [9241] 14. Hunter, Carl G. 1989. Trees, shrubs, and vines of Arkansas. Little Rock, AR: The Ozark Society Foundation. 207 p. [21266] 15. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384] 16. Landin, Mary C. 1979. The importance of wetlands in the north central and northeast United States to non-game birds. In: DeGraaf, Richard M.; Evans, Keith E., compilers. Management of north central and northeastern forests for nongame birds: Proceedings of the workshop; 1979 January 23-25; Minneapolis, MN. Gen. Tech. Rep. NC-51. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Forest Experiment Station: 179-188. [18087] 17. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952] 18. Medve, Richard J. 1984. The mycorrhizae of pioneer species in disturbed ecosystems of western Pennsylvania. American Journal of Botany. 71(6): 787-794. [8544] 19. Nixon, Charles M.; McClain, Milford W.; Russell, Kenneth R. 1970. Deer food habits and range characteristics in Ohio. Journal of Wildlife Management. 34(4): 870-886. [16398] 20. Olson, Jerry S. 1958. Rates of succession and soil changes on southern Lake Michigan sand dunes. Botanical Gazette. 119(3): 125-170. [10557] 21. Raunkiaer, C. 1934. The life forms of plants and statistical plant geography. Oxford: Clarendon Press. 632 p. [2843] 22. Stickney, Peter F. 1989. Seral origin of species originating in northern Rocky Mountain forests. Unpublished draft on file at: U.S. Department of Agriculture, Forest Service, Intermountain Research Station, Fire Sciences Laboratory, Missoula, MT; RWU 4403 files. 7 p. [20090] 23. Smith, Albert J. 1975. Invasion and ecesis of bird-disseminated woody plants in a temperate forest sere. Ecology. 56(1): 19-34. [15667] 24. Strole, Todd A.; Anderson, Roger C. 1992. White-tailed deer browsing: species preferences and implications for central Illinois forests. Natural Areas Journal. 12(3): 139-144. [19494] 25. Swan, Frederick R., Jr. 1970. Post-fire response of four plant communities in south-central New York state. Ecology. 51(6): 1074-1082. [3446] 26. Tester, John R. 1989. Effects of fire frequency on oak savanna in east-central Minnesota. Bulletin of the Torrey Botanical Club. 116(2): 134-144. [9281] 27. U.S. Department of Agriculture, Soil Conservation Service. 1982. National list of scientific plant names. Vol. 1. List of plant names. SCS-TP-159. Washington, DC. 416 p. [11573] 28. Van Dersal, William R. 1938. Native woody plants of the United States, their erosion-control and wildlife values. Washington, DC: U.S. Department of Agriculture. 362 p. [4240] 29. Vogel, Willis G. 1981. A guide for revegetating coal minespoils in the eastern United States. Gen. Tech. Rep. NE-68. Broomall, PA: U.S. Department of Agriculture, Forest Service, Northeastern Forest Experiment Station. 190 p. [15577] 30. Voss, Edward G. 1985. Michigan flora. Part II. Dicots (Saururaceae--Cornaceae). Bull. 59. Bloomfield Hills, MI: Cranbrook Institute of Science; Ann Arbor, MI: University of Michigan Herbarium. 724 p. [11472]

Index

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