KUCHLER TYPE
KUCHLER TYPE: Northern floodplain forest



KUCHLER-TYPE-NUMBER : 
K098

PHYSIOGNOMY : 
Low to tall broadleaf deciduous forest, open to dense, often with
lianas [30].


OCCURRENCE : 
Northern floodplain forest as mapped by Kuchler [31] occurs from eastern
Montana east to Minnesota and south to eastern Colorado and northern
Oklahoma.  The elm-ash-cottonwood (Ulmus spp.-Fraxinus spp.-Populus
spp.) ecosystem (FRES 17), which is largely synonymous with northern
floodplain forest, occurs on the lower terraces and floodplains of the
Mississippi, Missouri, Platte, Kansas, and Ohio rivers, and from the
Dakotas, Minnnesota, and Ohio south through Kansas, southern Illinois
and Missouri, with additional areas in Pennsylvania, New England, and
New Jersey.  The areas of elm-ash-cottonwood in the central states as
far south as southern Illinois, southern Indiana, and Kentucky (not
mapped by Kuchler) are considered to be included in northern floodplain
forest.  Elm-ash-cottonwood east of the Mississippi River Basin includes
other species with distributions more restricted to the eastern United
States and is thus not considered northern floodplain forest sensu
Kuchler [15,23].  In Montana east of the Rocky Mountains the species
present are largely consistent with Kuchler's description of northern
floodplain forest.  Although they are included in the western hardwoods
ecosystem (FRES 28) [15], these forests are considered part of the
northern floodplain forest Kuchler type.

In central North Dakota, the Missouri River floodplain varies in width
from less than 1 mile (1.6 km) to 7 miles (11.3 km) with four major
terrace levels [24].  The streamside forested area in southeastern
Nebraska averages approximately 15 miles (24 km) wide, near Omaha it is
about 5 miles (8 km) wide, and in northeastern Nebraska it is 2 miles
(3.2 km) wide or less.  The extent of cottonwood-willow (Salix spp.)
stands along the Missouri River is usually about one-half mile from the
water's edge, and usually limited to the lowest level ("first bottoms").
Upper terraces ("second bottoms") are mostly grassland with sparse to
absent woody cover [51].  Stack and others [46] estimated that there
were approximately 17,290 acres (7,000 ha) of forest remaining along the
Red River in North Dakota.


COMPILED BY AND DATE : 
Janet Sullivan, January 1995


LAST REVISED BY AND DATE : 
NO-ENTRY


AUTHORSHIP AND CITATION : 
Sullivan, Janet. 1995. Northern floodplain forest. In: Remainder of Citation


Kuchler Type Index
FEIS Home

KUCHLER TYPE DESCRIPTION


PHYSIOGRAPHY : 
Where the upper Missouri River flows through South Dakota, western Iowa,
eastern Nebraska and adjoining areas of Missouri and Kansas, the topography
is mostly rolling to hilly; elevations in this area range from 1,250
feet (381 m) at the Nebraska-South Dakota line to 850 feet (259 m) in
southeastern Nebraska [51].  In southern Illinois, bottomlands are
characteristically flat; slight variations in elevation are associated
with different soils, drainage conditions, and trees species.
Characteristic features include ridges, flats, sloughs, swamps,
sandbars, pointbars, and natural levees [23].

On the Wabash River, Indiana, upper terraces have mostly been cleared
for cultivation; first bottoms are still forested [33].  First bottoms
are subject to frequent flooding.  Second bottoms and upper terraces
formed by older drainage systems rarely experience floods [23].


CLIMATE : 
The elm-ash-cottonwood ecosystem, which includes northern floodplain
forests, crosses several climatic zones.  It is characteristic of moist
sites that either flood frequently, receive abundant precipitation, or
both.  Mean annual precipitation ranges from approximately 10 inches
(254 mm) near the Rocky Mountain foothills to 50 inches (1,270 mm) in
southern and northeastern areas [15].

Central North Dakota:  This climate is described as dry subhumid
mesothermal.  Precipitation is substantially lower than potential
evapotranspiration.  The average frost-free period is 140 days.  The
mean annual precipitation is 16.5 inches (420 mm), 70 percent of which
falls in growing season, 50 percent from May to July [24].  Severe
drought occurs about once every 10 years at the North Dakota-Minnesota
border, and about once every 20 years in Minnesota.  The Red River Basin
has a mean annual temperature ranging from 36 to 43 degrees Fahrenheit
(2.2-6 deg C) on a north-south gradient.  Average annual precipitation
ranges from a low of 16 inches (406 mm) in the west to a high of 25
inches (635 mm) in the east [49].

Western Iowa to eastern Nebraska:  Mean annual precipitation is 34
inches (864 mm) in western Iowa [51].  Eastern Nebraska has a
continental climate with average annual precipitation of 23 inches (584
mm).  The mean annual temperature is 51.6 degrees Fahrenheit (10.9 deg
C).  The average number of frost-free days is 188.  Eastern Nebraska
experiences occasional drought years with annual precipitation below 19
inches (480 mm) [43,51].

Southern Illinois:  The climate is continental with hot summers and cool
to cold winters.  Average frost-free periods range from 180 to 210 days.
The mean annual precipitation ranges from 42 to 47.9 inches (1,066-1,216
mm).  May is the wettest month, July the driest.  Extended periods of
soil moisture deficit are common in mid- to late summer [23].


SOILS : 
Floodplain soils are silts, sands, and clays; soil development is
largely driven by fluvial processes.  The soils under young stands of
cottonwoods and willows tend to be sandy with limited amounts of organic
matter and are low in nutrients and available water capacity.  Available
water capacity is higher in soils under older stands which have
developed higher organic matter content and have higher nutrient levels
due to flood-deposited silts.  Changes in the surface soil texture can
occur rapidly in floodplain communities due to flood-carried siltation
[24].  Soil types and development in the Red River Valley of North
Dakota are discussed in detail by Wanek [49].

Wilson [53] described eastern cottonwood (Populus deltoides var.
deltoides) forests along the Missouri River in southeastern South
Dakota.  Over 20 years of eastern cottonwood growth, there was a gradual
change from the initial dry sandy surface soils supporting little
undergrowth to moist clay or silt surface soils with an abundance of
shrubs.


VEGETATION : 
Kuchler [30] lists three dominant tree species for northern floodplain
forest:  eastern cottonwood, black willow (Salix nigra), and American
elm (Ulmus americana).  Cottonwoods (Populus spp.) often occur in pure
or nearly pure stands [15].

Other components of northern floodplain forest include (in alphabetical
order by scientific name) boxelder (Acer negundo), red maple (A.
rubrum), silver maple (A. saccharinum), river birch (Betula nigra),
American bittersweet (Celastrus scandens), hackberry (Celtis
occidentalis), virgin's bower (Clematis virginiana), white ash (Fraxinus
americana), green ash (F. pennsylvanica), honeylocust (Gleditsia
triacanthos), black walnut (Juglans nigra), Virginia creeper
(Parthenocissus quinquefolia), sycamore (Platanus occidentalis) in the
south, plains cottonwood (Populus deltoides var. occidentalis) in the
west, eastern poison-ivy (Toxicodendron radicans), peachleaf willow
(Salix amygdaloides), sandbar willow (S. exigua), bristly greenbrier
(Smilax hispida), coralberry (Symphoricarpos orbiculatus), and slippery
elm (U. rubra) [30].  Garrison and others [15] list sugarberry (Celtis
laevigata) for southern stands.  Streamside stands of trees in North
Dakota include bur oak (Quercus macrocarpa), American basswood (Tilia
americana), green ash, American elm, boxelder, quaking aspen (P.
tremuloides), and paper birch (Betula papyrifera).  These stands appear
to be successionally related to northern floodplain forest as defined by
Kuchler [52].

The following Society of American Foresters cover types appear to
overlap with Kuchler's definition of northern floodplain forest:
cottonwood (SAF 63), black willow (SAF 95), cottonwood-willow, and
silver maple-American elm (SAF 235) [14].  The cottonwood type (which
includes eastern cottonwood, plains cottonwood, and swamp cottonwood [P.
heterophylla]) is a pioneer type.  Cottonwoods occur with or precede
willows, usually sandbar willow, black willow, or peachleaf willow.  In
some areas the willow stage is identifiable as a separate cover type
(black willow).  Early successional associates in the cottonwood type
include white ash, green ash, silver maple, and American elm in the
north, and pecan (Carya illinoensis), sycamore, and sugarberry in the
south.  Cottonwoods are somewhat less flood tolerant than willows; with
frequent and/or extended flooding willows may survive cottonwoods and
dominate the site.  Cottonwood merges with cottonwood-willow (SAF 235)
in the eastern Great Plains [14].

Cottonwood-willow (SAF 235) occurs mostly as gallery forest in
grasslands.  It is a climatic anomaly because it includes species that
occur regularly in cooler areas with abundant precipitation.  It
therefore is considered a specialized type whose existence depends on
groundwater availability.  This type includes other species of
cottonwood in addition to eastern cottonwood and plains cottonwood:
lanceleaf (P. xacuminata), Fremont (P. fremontii), Rio Grande (P.
fremontii var. wislizeni), narrowleaf (P. angustifolia), and black
cottonwood (P. trichocarpa).  Willows are secondary only to cottonwood
and usually include black willow, peachleaf willow, other tree willows,
and many shrubby willow species.  Minor associates include boxelder and
hackberry; other species described for this type occur farther south and
west than northern floodplain forest [14].

Species that commonly invade cottonwood-willow stands in northern
floodplain forests include sycamore, sweet pecan, hackberry, sugarberry,
river birch, green ash, American elm, silver maple, red maple, and
boxelder.  The silver maple-American elm cover type (SAF 62) is most
common in the Ohio, Wabash, upper Mississippi, and Missouri river
valleys and also occurs in the Great Lakes and St. Laurence River
regions of Canada.  It is usually a subclimax type following cottonwoods
and willows; it is a climax type in southern Ontario.

Other community descriptions and classification systems that contain
types approximately equivalent to northern floodplain forest include the
following:

A preliminary classification of the natural vegetation of Colorado [2]
Composition and environment of floodplain forests of northern Missouri [11]
Native woodland habitat types of southwestern North Dakota [17]
Riparian dominance types of Montana [20]
Floristic analysis of the Missouri River bottomland forest in North
  Dakota [26]
A comparison of the plant communities of the South Platte and Arkansas
  River drainages in eastern Colorado [32]
Vegetation and environment along the Wabash and Tippecanoe Rivers [33]
Community analysis of the forest vegetation in the lower Platte River
  Valley, eastern Nebraska [43]
The timber resources of North Dakota [50]


WILDLIFE : 
Birds:  Birds are the most common and conspicuous form of wildlife in
riparian ecosystems, and much research describes riparian bird
communities.  The most abundant breeding birds on 998 census plots in
riparian vegetation (all areas) included red-eyed vireo (Vireo
olivaceus), northern cardinal (Cardinalis cardinalis), common
yellowthroat (Geothlypis trichas), song sparrow (Melospiza melodia),
wood thrush (Hylocichla mustelina), American redstart (Setophaga
ruticilla), Acadian flycatcher (Empidonax alnorum), red-winged blackbird
(Agelaius phoeniceus), European starling (Sturnus vulgaris), American
robin (Turdus migratorius), gray catbird (Dumetella carolinensis),
tufted titmouse (Parus bicolor), house wren (Troglodytes aedon),
mourning dove (Zenaida macroura), eastern wood pewee (Contopus virens),
yellow warbler (Dendroica petechia), rufous-sided towhee (Pipilo
erythrophthalmus), northern oriole (Icterus galbula), indigo bunting
(Passerina cyanea), ovenbird (Seiutus aurocapillus), northern parula
(Parula americana), northern flicker (Colaptes auratus), and blue jay
(Cyanocitta cristata) [8].  Kricher's [29] list of indicator animals for
northern floodplain forests includes belted kingfisher (Ceryle alcyon),
bank swallow (Riparia riparia), spotted sandpiper (Actitus macularia),
green-backed heron (Butorides striatus), wood duck (Aix sponsa),
yellow-throated vireo (V. flavifrons), and blue-gray gnatcatcher
(Polioptila caerulea).

The most abundant lowland forest (assumed to be equivalent to northern
floodplain forest) bird species on the Platte River, Nebraska, include
house wren, mourning dove, American robin, and brown-headed cowbird
(Molothrus ater).  The eastern phoebe (Sayornis phoebe), eastern wood
pewee, Bewick's wren (Thryomanes bewickii), eastern bluebird (Sialia
sialis), blue grosbeak (Guiraca caerulea), lazuli bunting (Passerina
amoena), and rufous-sided towhee occurred in the area only in this
habitat type.  The most numerous species occurring on wooded sandbars
were cliff swallow (Hirundo pyrrhonota) and common yellowthroat.  Other
species on wooded sandbars included belted kingfisher, willow flycatcher
(E. traillii), bank swallow, Bell's vireo (Vireo bellii), northern
cardinal, and indigo bunting [48].  Garrison and others [15] add the
following typical forest edge birds as common in elm-ash-cottonwood
ecosystems:  northern bobwhite (Colinus virginianus), American goldfinch
(Carduelis tristis), yellow-billed cuckoo (Coccyzus americanus), lark
sparrow (Chondestes grammacus), northern mockingbird (Mimus
polyglottos), common crow (Corvus brachyrhynchos), ruby-throated
hummingbird (Archilochus colubris), and Cooper's hawk (Accipiter
cooperii).

Mammals:  Mammals commonly occurring in riparian areas in the northern
Great Plains include eastern mole (Scalopus aquaticus), shrews (Family
Soricidae), raccoon (Procyon lotor), mink (Mustela vison), striped skunk
(Mephitis mephitis), spotted skunks (Spilogale spp.), red fox (Vulpes
vulpes), coyote (Canis latrans), fox squirrels (Sciurus spp.), pocket
gophers (Thomomys spp.), ground squirrels (Spermophilus spp.), beaver
(Castor canadensis), western harvest mouse (Reithrodontomys megalotis),
deer mouse (Peromyscus maniculatus), white-footed mouse (Peromyscus
leucopus), hispid cotton rat (Sigmodon hispidus), northern red-backed
vole (Clethrionomys rutilus), common muskrat (Ondatra zibethicus),
meadow jumping mouse (Zapus hudsonius), cottontails (Silvilagus spp.),
swamp rabbit (S> aquaticus), jackrabbits and hares (Lepus spp.),
white-tailed deer (Odocoileus virginiana), and mule deer (O. hemionus)
[8,15].

Reptiles and Amphibians:  Common reptiles and amphibians in Plains
grasslands riparian ecosystems include snapping turtle (Chelydra
serpentina), mud turtles (Kinosternon spp.), sliders (Trachemys spp.),
cooters (Pseudemys spp.), water turtles (Clemmys spp.), box turtles
(Terrapene spp.), softshell turtles (Apalone spp.), earless lizards
(Holbrookia spp.), spiny lizards (Sceloporus spp.), skinks (Scincella
and Eumeces spp.), whiptails and racerunners (Cnemidophorus spp.), water
snakes (Nerodia spp.), redbelly and brown snakes (Storeria spp.), racers
(Coluber spp.), green snakes (Opheodrys spp.), rat snakes (Elaphe spp.),
bullsnake (Pituophis melanoleucus sayi), eastern massassauga (Sistrurus
catenatus catenatus), pygmy rattlesnakes (Sistrurus spp.), rattlesnakes
(Crotalus spp.), mole salamanders (Ambystoma spp.), spadefoots
(Scaphiopus spp.), toads (Bufo spp.), cricket frogs (Acris spp.),
treefrogs (Hyla spp.), and true frogs (Rana spp.) [8].


ECOLOGICAL RELATIONSHIPS : 
Northern floodplain forest dynamics are controlled by fluvial processes
and moisture availability.  Hydroperiod determines vegetation survival
and nutrition.  The hydroperiod may be coincidental with nutrient pulses
(i.e., silt deposition), but nutrient availability is dependent on
acidity and oxygenation; these are determined in large part by soil water
capacity [13].

The northern floodplain forests within the Great Plains (usually
described as gallery forests) are limited in extent by moisture
availability and by fire.  These forests are usually confined by
moisture requirements to moist, sheltered sites on the bottoms and lower
slopes of draws [7,51]. It is generally accepted that prior to 1855,
woody vegetation in the Great Plains existed almost exclusively along or
near rivers and streams [44].  During presettlement times frequent
prairie fires commonly burned up to and sometimes on to the slope
bordering the river valleys where northern floodplain forests developed
[21,42,43].

In northern floodplain forests, willows are usually the species closest
to water's edge, followed by cottonwoods, elms, boxelder, and ashes.
When a stream develops a floodplain with wide, protecting banks,
large-fruited species such as bur oak, black walnut, hickory (Carya
spp.), and hazel (Corylus spp.) are able to establish and survive
[49,51].  Boggs [6] identified the following community types for
riparian areas along the lower Yellowstone River in Montana:  bare
sandbar, willow thicket, cottonwood forest, green ash forest, and
grassland.

Northern floodplain forests originate with stream channel movement and
after floods.  The light, windblown seeds of cottonwoods and willows
require bare, moist mineral soil for seedling establishment
[12,14,15,51].  Cottonwoods and willows are common pioneers on point
bars, well-drained flats, front land ridges, and newly formed natural
levees [12,14,15].  New stream development is often accompanied by
cottonwood and willow establishment [51].  Distinct zones of
establishment often separate cottonwood and sandbar willow, with sandbar
willow on the more frequently flooded sites [38].

Willows, particularly sandbar willow, have abundant fibrous roots which
help stabilize soils.  Their roots and stems also help catch silts and
clays carried by floodwaters, further adding to soil development.
Cottonwoods exhibit very rapid growth and usually overtop willows,
except at the water's edge where the more flood-tolerant willows outlive
cottonwoods.  Occasionally erosion is rapid enough to remove the
willow-dominated zone to the point where cottonwood stands are in
contact with the river's edge [51].

On the Little Missouri River in western North Dakota, cottonwoods occur
in a series of even-aged stands corresponding to channel migration
events [12].  River meandering and lateral movement of the channel,
deposition on interior curves, collapse of cut banks, and floods ensure
a constant supply of bare sites for cottonwood and willow establishment
and initiation of succession [12,24].  Cottonwood seedlings are often
"flood trained":  Young stems are bent over by floodwaters with the
direction of flow.  These stems are often silted into position and
subsequently sprout, forming small groves of trees from one original
seedling [12].  Young cottonwood-willow forests in central North Dakota
are composed of many small trees from 18 to 36 feet (6-12 m) tall [24].
During floods, stems over 8 inches (20.3 cm) in diameter may break or
become uprooted rather than bend over [12].

In western North Dakota, the herb layer of young cottonwood forests is
mostly composed of annuals including clovers (Trifolium spp.).  Gradual
establishment of shrubs and vines such as elderberry (Sambucus
canadensis), coralberry, American bittersweet, and grapes (Vitis spp.)
is facilitated by frugivorous birds [51].

Old cottonwood stands in central North Dakota consist of trees from 59
to 80 feet (18-24 m) tall that are widely spaced with large, straight,
unbranched boles ranging from 6 to 36 inches (16-91 cm) dbh.  There is
rarely a closed canopy; the open stand allows numerous species of shrubs
and saplings of other species to develop a distinct layer.  Arboreal
lianas are prominent [24].  Older forests contain more perennial grasses
and shrubs than do young stands.  Very few cottonwood saplings and
seedlings are found under cottonwood canopies.  Old trees occasionally
regenerate by suckering [12].  In southeastern South Dakota, cottonwood
reproduction of any kind is rare after 15 to 25 years of cottonwood
overstory growth [53].

The cottonwood-willow stage is followed by river birch and silver
maple-American elm types in the north, and sycamore-pecan-American elm
or sugarberry-American elm-green ash types in southern Illinois,
Indiana, and Kentucky [14,15].  In western North Dakota, replacement of
cottonwood trees occurs from about 30 to 60 years.  Common replacement
species include green ash, Rocky Mountain juniper (Juniperus
scopulorum), American elm, and boxelder.  Old stands usually retain a
few old cottonwoods [12].  In central and southeastern North Dakota,
green ash, boxelder, American elm, and bur oak usually replace
cottonwoods and are predominant in old stands near the edge of the
floodplain [24].  In central North Dakota floodplains, tree species
diversity increases with stand age in the initial stages of replacement,
is maximal in stands with mixtures of intolerant and tolerant species,
and decreases slightly in the oldest stands [24].

Mature cottonwood forests adjacent to grasslands may be replaced by
shrubs and eventually, grasses, depending on moisture availability [6].

KUCHLER TYPE VALUE AND USE
KUCHLER TYPE: Northern floodplain forest



FORESTRY VALUES : 
Cottonwood is valued for sawtimber and pulp and is commercially
harvested on a small scale in central North Dakota [24].  Eastern
cottonwood is the fastest-growing bottomland species in the central
states and black willow grows almost as fast [25].  Eastern cottonwood
can be managed in short rotation even-aged stands [36], and is a good
candidate for plantation silviculture.  Light thinning is recommended at
5- to 10-year intervals beginning at age 5.  Most stands of eastern
cottonwood are harvested by age 50, although stands may remain healthy
until 75 years.  Black willow begins to deteriorate at around 35 years
and commercial value is low by 50 years [25].  Well-drained silt flats
are the most valuble timber and fiber producing areas.  Depressional
swamps do not produce commercial timber or fiber, but silver maple and
willow in these swamps provide cordwood and can be regenerated by
coppicing.  Silver maple is a valuable timber species in the Midwest and
is managed by clearcutting or group selection [36,37].

Other northern floodplain forest species valued for timber and fiber
production include American elm, green ash, sycamore, hackberry, red
maple, and river birch.  Site index curves for cottonwood, American elm,
silver maple, and green ash are available [37].


RANGE VALUES : 
In the Great Plains, many livestock grazing areas include patches of
northern floodplain forest.  These forests provide important shade and
shelter for cattle grazing on adjacent grasslands [16].


WILDLIFE VALUES : 
Riparian forests including northern floodplain forests are important
wildlife habitat, particularly for birds [8,36].  These forests are
complex due to the mixture of many physical and biological features.
There is substantial development of edge habitat at the interfaces
between stream and forest, and between forest and adjacent uplands.
Many species occur almost entirely in edge zones [8].

Open stands of northern floodplain forest with heavy herb cover are
ideal waterfowl nesting habitat.  Dead elms and cottonwoods provide food
and shelter for many species of songbirds [36].  Cottonwood plantations
provide excellent wildlife habitat [25].  On the Platte River, extensive
tracts of woodland are of benefit to many species of raptors including
bald eagle (Haliaeetus leucocephalus), peregrine falcon (Falco
peregrinus), prairie falcon (F. mexicanus), American kestrel (F.
sparverius), northern harrier (Circus cyaneus), red-tailed hawk (Buteo
jamaicensis), rough-legged hawk (B. lagopus), ferruginous hawk (B.
regalis), and golden eagle (Aquila chrysaetos) [48].

The Big Bend Reach of the Platte River is an area of great importance to
numerous species of migratory birds using the Central Flyway [47].  This
area is an important stopover site for sandhill crane (Grus canadensis),
which roost in the river and feed in nearby corn fields [56].

Threatened and endangered animals occurring in riparian ecosystems (some
of which are northern floodplain forests) include Indiana myotis (Myotis
sodalis), gray myotis (M. grisescens), whooping crane (G. americana),
bald eagle, peregrine falcon, and Bachman's warbler (Vermivora
bachmanii) [47].  Riparian ecosystems are important to at least
one-third of the species or subspecies listed as threatend or endangered
by the U.S. Fish and Wildlife Service.  Many riparian species are
candidates for future federal listing as threatened or endangered.
These species include Bell's vireo, western populations of yellow-billed
cuckoo, many invertebrates, and many plants [8].

Breeding birds along the Platte River, Nebraska, are diverse.  Some 142
species have been observed nesting in northern floodplain forest and
adjacent ecosystems.  In many areas, decreased flow due to dams and
irrigation has decreased meandering and increased land area.  Species
which nest in northern floodplain forest have increased because of the
extension of woody vegetation into former river channels.  However,
available nesting habitat for species such as least tern (Sterna
antillarum) and piping plover (Charadrius melodus), which nest on
sandbars, has decreased [48].  Breeding bird densities in some riparian
ecosystems (largely equivalent to northern floodplain forest) are as
follows (compiled by Brinson and others [8]):

                                  birds per 40 ha
willow, Colorado                    100
cottonwood-willow, Colorado         525-589
cottonwood-willow, Colorado         225-900
cottonwood, Colorado                319
bottomland forests, Oklahoma        400
riparian communities, Great Plains  137-748


Riparian bird population densities in winter are as follows [45]:

                                                  birds per 40 ha
floodplain forest, Illinois (2 locations)          148;226
mixed habitat-disturbed bottomland, Oklahoma       183
floodplain cottonwood, Colorado                    186
cottonwood-willow riverbottom, Colorado            311


OTHER VALUES : 
Floodplain forests help regulate water level, flow rate and direction,
bank stability, and potential evapotranspiration.  Changes in the
vegetation affect flood characteristics.  Flood damage to bridges and
other streamside structures can be reduced if open forests are
maintained adjacent to these structures [36].


MANAGEMENT CONCERNS : 
Floodplains are a mosaic of cultivated fields, marshes, sand dunes,
sandbars, brushland, and forest [24].  Klapotek and others [28] reported
in 1979 that the potential area of northern floodplain forest was over
17 million acres (7,171,000 ha) but that the actual area covered by
northern floodplain forests was less than 700,000 acres (279,000 ha).
Irrigation and flood control projects have resulted in a 70 percent
reduction in flow of the Platte River in Nebraska.  As a result woody
vegetation has encroached on thousands of hectares of former channel
area, contributing to changes in channel features and altering wildlife
habitat [56].  Petranka and Holland [39], however, reported that there
were only minor differences in plant species composition and structure
between channelized and undisturbed south-central Oklahoma gallery
forests (which contain predominately the same species as northern
floodplain forest but are not mapped as such) of various ages since
channelization.

Heavy use of floodplain forests by livestock can have severe detrimental
effects on the vegetation.  Concentrated use by cattle can result in
sparse stands of low vigor, much dead material on the ground, compacted
soils, and little or no seedling survival.  This type of damage is
particularly common when the forest occupies only a small portion of a
range.  Continuous summer grazing of these woodlands, even under good
conditions, will probably result in some degradation [58].

Riparian ecosystems are dependent on fluvial processes and drainage
density for soil moisture, nutrient availability, and community
stability and structure [8].  Usually, resilience of floodplain forests
to timber harvests is relatively high.  A combination of harvest and
site alteration (i.e., dam construction and irrigation projects) may,
however, increase regeneration time for many forests [13].  Many
riparian woodlands in the United States have been replaced by grasses;
many others are in a state of decline due to a number of factors.  Many
of the stands are exposed to concentrated use by livestock; some are
nearing the natural end of their lifespans and are vulnerable to
insects, disease, and mechanical stresses [7].  Heavy grazing and timber
cutting have reduced some central North Dakota floodplain forests to
brushlands [24].

A lack of eastern cottonwood seedling establishment along the Missouri
River in central North Dakota was attributed to a reduction in the
meandering rate following the construction of dams [24,53].  Lack of
cottonwood seedling establishment in other floodplains is also
attributed to flood control projects [7,24,26].  A decline in the
establishment of silver maple and elms, and a reduction in the diameter
growth rate for silver maple, elms, and ash is apparently related to
flood control which reduces the height and duration of spring flooding
[24,53].  Boxelder and sweet pecan also appear to depend on flooding for
reproduction.  Boxelder can even be used as an indicator of flooding
frequency [3].

Channelization of streams in south-central Oklahoma has contributed to
reductions in bottomland forest area of up to 93 percent.  These forests
are not included in northern floodplain forest maps but consist of the
species listed by Kuchler for northern floodplain forest.  On two study
sites, the combined floodplain areas were 76 percent forested in 1871
and only 10 percent forested in 1969.  Black willow and eastern
cottonwood seedlings are restricted to the channel zone.  Bird densities
were substantially lower on channelized sites than on unchannelized
sites.  Species that occurred only on unchannelized sites included
northern parula, Louisiana waterthrush (Seiurus motacilla), northern
cardinal, Carolina chickadee (Parus carolinensis), tufted titmouse,
brown-headed cowbird, red-bellied woodpecker, indigo bunting, painted
bunting, yellow-bellied cuckoo, Carolina wren, and American goldfinch
[3].

The widespread infection of American elm with Dutch elm disease has
important consequences for conservation, floodwater management, and
resource utilization in northern floodplain forests.  Loss of large
trees decreases bank stability, increases water availability (and flood
level), and decreases timber [46].

Wildlife species that require a combination of riparian habitat features
are more sensitive to alterations than those requiring only one
component [8].

KUCHLER TYPE FIRE ECOLOGY AND MANAGEMENT
KUCHLER TYPE: Northern floodplain forest



FUELS, FLAMMABILITY, AND FIRE OCCURRENCE : 
Fire is rare in northern floodplain forest in humid climates.  Higgins
[21] noted that fire was frequent enough to favor grasslands in the
northern Great Plains, but that areas with rough and dissected
topography were much less likely to experience fire [21].  However,
occasionally fire probably influenced presettlement northern floodplain
forests in drier climates.  In a North Dakota gallery forest, the oldest
trees were only 80 years old, even though the forest had been recorded
in presettlement survey records.  A long-term local resident noted that
prairie fires frequently invaded the forest and sometimes killed some of
the larger trees [27].


FIRE EFFECTS ON SITE : 
NO-ENTRY


FIRE EFFECTS ON VEGETATION : 
Willows (including black willow, sandbar willow, and peachleaf willow)
are usually top-killed by fire.  Severe fire that consumes organic
layers and exposes roots kills most willows.  Most top-killed willows
sprout following fire; however, black willow typically decreases
following fire [57].  Willows are pioneer species on burned sites and
establish from light, windborne seed [54].  Fire easily kills
cottonwood seedlings and saplings.  Trees less than 20 years old may
sprout after fire damage.  Large cottonwoods have thick bark which may
provide some resistance to fire damage, but fire wounds are likely to
create avenues for infection [58].

Mid- and late successional northern floodplain species such as American
elm, silver maple, and boxelder are easily damaged or killed by fire,
particularly when young.  Some species sprout after damage or top-kill,
particularly green ash [16].  Bur oak is more resistant to fire damage;
prairie fires burning into bur oak-dominated stands tend to maintain the
large oaks with a grassy understory [42].

Controversy continues over whether northern Great Plains woodlands have
expanded or declined since settlement and the beginning of fire
exclusion in the late 19th century.  The suppression of fire may have
allowed woodland expansion; but some authors argue that woodland species
are adapted to fire (due to sprouting ability and rapid colonization of
bare sites) and thus may be limited by climate and moisture availability
[58].  In the absence of fire, trees and shrubs may replace prairie
grasses on mesic sites [51].


FIRE EFFECTS ON RESOURCE MANAGEMENT : 
NO-ENTRY


FIRE USE CONSIDERATIONS : 
NO-ENTRY


FIRE MANAGEMENT CONSIDERATIONS : 
Myers and Buchman [37] do not consider fire a management tool for site
preparation or shrub control in elm-ash-cottonwood types.  All species
present are susceptible to fire damage; surface fire readily kills
saplings and seedlings, and wounds larger trees.  Fire wounds lead to
heart rots and decrease timber values [37].  Putnam [41] also made a
general recommendation against the use of prescribed fire for any
objectives in bottomland forests.


REHABILITATION OF SITES FOLLOWING WILDFIRE : 
NO-ENTRY

References for Kuchler: [K098]

1. Aikman, John M. 1926. Distribution and structure of the forests of eastern Nebraska. University Studies. 26(1-2): 1-75. [6575]

2. Baker, William L. 1984. A preliminary classification of the natural vegetation of Colorado. The Great Basin Naturalist. 44(4): 647-676. [380]

3. Barclay, John S. 1980. Impact of stream alterations on riparian communities in southcentral Oklahoma. FWS/OBS-80/17. Albuquerque, NM: U.S. Department of the Interior, Fish and Wildlife Service, Region 2, Biological Services Program. 91 p. [23990]

4. Bell, David T. 1974. Tree stratum composition and distribution in the streamside forest. The American Midland Naturalist. 92(1): 35-46. [10410]

5. Bellah, R. Glenn; Hulbert, Lloyd C. 1974. Forest succession on the Republican River floodplain in Clay County, Kansas. The Southwestern Naturalist. 19(2): 155-166. [241]

6. Boggs, Keith Webster. 1984. Succession in riparian communities of the lower Yellowstone River, Montana. Bozeman, MT: Montana State University. 107 p. Thesis. [7245]

7. Boldt, Charles E.; Uresk, Daniel W.; Severson, Kieth E. 1979. Riparian woodlands in jeopardy on Northern High Plains. In: Johnson, R. Roy; McCormick, J. Frank, technical coordinators. Strategies for protection & management of floodplain wetlands & other riparian ecosystems: Proc. of the symposium; 1978 December 11-13; Callaway Gardens, GA. Gen. Tech. Rep. WO-12. Washington, DC: U.S. Department of Agriculture, Forest Service: 184-189. [4359]

8. Brinson, Mark M.; Swift, Bryan L.; Plantico, Reuben C.; Barclay, John S. 1981. Riparian ecosystems: their ecology and status. FWS/OBS-81/17. Kearneysville, WV: U.S. Department of the Interior, Fish and Wildlife Service, National Water Resources Analysis Team, Eastern Energy and Land Use Group. 154 p. [23991]

9. Cox, Mike K.; Franklin, William L. 1989. Terrestrial vertebrates of Scotts Bluff National Monument, Nebraska. The Great Basin Naturalist. 49(4): 597-613. [11004]

10. DeGraaf, Richard M.; Scott, Virgil E.; Hamre, R. H.; [and others]. 1991. Forest and rangeland birds of the United States: Natural history and habitat use. Agric. Handb. 688. Washington, DC: U.S. Department of Agriculture, Forest Service. 625 p. [15856]

11. Dollar, K. E.; Pallardy, Stephen G.; Garrett, H. Gene. 1992. Composition and environment of floodplain forests of northern Missouri. Canadian Journal of Forest Research. 22: 1343-1350. [19706]

12. Everitt, Ben L. 1968. Use of the cottonwood in an investigation of the recent history of a flood plain. American Journal of Science. 266: 417-439. [25445]

13. Ewel, Katherine Carter. 1979. Riparian ecosystems: conservation of their unique characteristics. In: Johnson, R. Roy; McCormick, J. Frank, technical coordinators. Strategies for protection & mgmt. of floodplain wetlands & other riparian ecosystems: Proceedings of the symposium; 1978 December 11-13; Callaway Gardens, GA. Gen. Tech. Rep. WO-12. Washington, DC: U.S. Department of Agriculture, Forest Service: 58-62. [4355]

14. Eyre, F. H., ed. 1980. Forest cover types of the United States and Canada. Washington, DC: Society of American Foresters. 148 p. [905]

15. Garrison, George A.; Bjugstad, Ardell J.; Duncan, Don A.; [and others]. 1977. Vegetation and environmental features of forest and range ecosystems. Agric. Handb. 475. Washington, DC: U.S. Department of Agriculture, Forest Service. 68 p. [998]

16. Girard, Michele M.; Goetz, Harold; Bjugstad, Ardell J. 1987. Factors influencing woodlands of southwestern North Dakota. Prairie Naturalist. 19(3): 189-198. [2763]

17. Girard, Michele M.; Goetz, Harold; Bjugstad, Ardell J. 1989. Native woodland habitat types of southwestern North Dakota. Res. Pap. RM-281. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 36 p. [6319]

18. Grimm, Eric C. 1984. Fire and other factors controlling the Big Woods vegetation of Minnesota in the mid-nineteenth century. Ecological Monographs. 54(3): 291-311. [22170]

19. Gutzwiller, K. J.; Anderson, S. H. 1988. Co-occurrence patterns of cavity-nesting birds in cottonwood-willow communities. Oecologia. 76: 445-454. [294]

20. Hansen, Paul L.; Chadde, Steve W.; Pfister, Robert D. 1988. Riparian dominance types of Montana. Misc. Publ. No. 49. Missoula, MT: University of Montana, School of Forestry, Montana Forest and Conservation Experiment Station. 411 p. [5660]

21. Higgins, Kenneth F. 1984. Lightning fires in North Dakota grasslands and in pine-savanna lands of South Dakota and Montana. Journal of Range Management. 37(2): 100-103. [1148]

22. Hopkins, Rick B.; Cassel, J. Frank; Bjugstad, Ardell J. 1986. Relationships between breeding birds and vegetation in four woodland types of the Little Missouri National Grasslands. Res. Pap. RM-270. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station. 12 p. [2758]

23. Hosner, John F.; Minckler, Leon S. 1963. Bottomland hardwood forests of southern Illinois--regeneration and succession. Ecology. 44(1): 29-41. [3739]

24. Johnson, W. Carter; Burgess, Robert L.; Keammerer, Warren R. 1976. Forest overstory vegetation and environment on the Missouri River floodplain in North Dakota. Ecological Monographs. 46(1): 59-84. [6313]

25. Johnson, R. L.; Shropshire, F. W. 1983. Bottomland hardwoods. In: Burns, Russell M., tech. comp. Silvicultural systems for the major forest types of the United States. Agric. Handb. 445. Washington, DC: U.S. Department of Agriculture, Forest Service: 175-179. [18953]

26. Keammerer, Warren R.; Johnson, W. Carter; Burgess, Robert L. 1975. Floristic analysis of the Missouri River bottomland forest in North Dakota. Canadian Field-Naturalist. 89: 5-19. [7447]

27. Killingbeck, K. T.; Wali, Mohan K. 1978. Analysis of a North Dakota gallery forest: nutrient, trace element and productivity relations. Oikos. 30: 29-60. [24046]

28. Klopatek, J. M.; Olson, R. J.; Emerson, C. J.; Joness, J. L. 1979. Land-use conflicts with natural vegetation in the United States. Environmental Sciences Division Publ. No. 1333. ORNL/TM-6814. Oak Ridge, TN: Department of Engergy, Oak Ridge National Laboratory. 22 p. [26104]

29. Kricher, John C. 1988. A field guide to eastern forests: North America. Peterson Field Guide Series 37. Boston: Houghton Mifflin Company. 368 p. [22677]

30. Kuchler, A. W. 1964. Manual to accompany the map of potential vegetation of the conterminous United States. Special Publication No. 36. New York: American Geographical Society. 77 p. [1384]

31. Kuchler, A. W. 1975. United States [Potential natural vegetation of the conterminous United States]. Special Publication No. 36. New York: American Geographical Society. 1:3,168,000; colored. 2nd edition. [28322]

32. Lindauer, Ivo E. 1983. A comparison of the plant communities of the South Platte and Arkansas River drainages in eastern Colorado. The Southwestern Naturalist. 28(3): 249-259. [5886]

33. Lindsey, Alton A.; Petty, Robert O.; Sterling, David K.; Van Asdall, Willard. 1961. Vegetation and environment along the Wabash and Tippecanoe Rivers. Ecological Monographs. 31(2): 105-156. [22578]

34. Little, Elbert L., Jr. 1979. Checklist of United States trees (native and naturalized). Agric. Handb. 541. Washington, DC: U.S. Department of Agriculture, Forest Service. 375 p. [2952]

35. Manci, Karen M. 1989. Riparian ecosystem creation and restoration: a literature summary. Biol. Rep.89(20). Washington, DC: U.S. Department of the Interior, Fish and Wildlife Service. 60 p. [11757]

36. Morris, L. A.; Mollitor, A. V.; Johnson, K. J.; Leaf, A. L. 1979. Forest management of floodplain sites in the northeastern United States. In: Johnson, R. Roy; McCormick, J. Frank, technical coordinators. Strategies for protection & mgmt of floodplain wetlands & other riparian ecosystems: Proceedings of the symposium; 1978 December 11-13; Callaway Gardens, GA. Gen. Tech. Rep. WO-12. Washington, DC: U.S. Department of Agriculture, Forest Service: 236-242. [4364]

37. Myers, Charles C.; Buchman, Roland G. 1984. Manager's handbook for elm-ash-cottonwood in the North Central States. Gen. Tech. Rep. NC-98. St. Paul, MN: U.S. Department of Agriculture, Forest Service, North Central Forest Experiment Station. 11 p. [8919]

38. Noble, Mark G. 1979. The origin of Populus deltoides and Salix interior zones on point bars along the Minnesota River. The American Midland Naturalist. 102: 59-67. [6172]

39. Petranka, James W.; Holland, Robert. 1980. A quantitative analysis of bottomland communities in south-central Oklahoma. The Southwestern Naturalist. 25(2): 207-214. [5885]

40. Phillippe, Philip E.; Ebinger, John E. 1973. Vegetation survey of some lowland forests along the Wabash River. Castenea. 38(4): 339-349. [22567]

41. Putnam, John A. 1951. Management of bottomland hardwoods. Occasional Paper 116. New Orleans, LA: U.S. Department of Agriculture, Forest Service, Southern Forest Experiment Station. 60 p. [6748]

42. Reichman, O. J. 1987. Forests. In: Konza Prairie: A tallgrass natural history. Lawrence, KS: University Press of Kansas: 115-124. [4255]

43. Rothenberger, Steven J. 1985. Community analysis of the forest vegetation in the lower Platte River Valley, eastern Nebraska. Prairie Naturalist. 17(1): 1-14. [2031]

44. Rothenberger, Steven J. 1989. Extent of woody vegetation on the prairie in eastern Nebraska, 1855-1857. In: Bragg, Thomas B.; Stubbendieck, James, eds. Prairie pioneers: ecology, history and culture: Proceedings, 11th North American prairie conference; 1988 August 7-11; Lincoln, NE. Lincoln, NE: University of Nebraska: 15-18. [14012]

45. Ryder, R. A.; Ryder, A., eds. 1979. Thirty-first winter bird-population study. American Birds. 33: 18-53. [28292]

46. Stack, R. W.; Jones, A. C.; Harsel, R. 1987. Impact of Dutch elm disease on the forest resource of the Red River Valley in North Dakota and Minnesota. In: Proceedings, 79th North Dakota Academy of Science. Grand Forks, ND. 41: 73. [5718]

47. U.S. Department of the Interior, Fish and Wildlife Service. 1980. Republication of lists of endangered and threatened species and correction of technical errors in final rules. Federal Register. 45(99): 33768-33781. [28293]

48. U.S. Department of the Interior, Fish and Wildlife Service, Northern Prairie Wildlife Research Center. 1981. The Platte River ecology study. Special Research Report. Jamestown, ND. 187 p. [25443]

49. Wanek, Wallace James. 1967. The gallery forest vegetation of the Red River of the North. Fargo, ND: North Dakota State University. 190 p. Dissertation. [5733]

50. Warner, John R.; Chase, Clarence D. 1956. The timber resource of North Dakota. Station Paper No. 36. St. Paul, MN: U.S. Department of Agriculture, Forest Service, Lake States Forest Experiment Station. 39 p. [25442]

51. Weaver, J. E. 1960. Flood plain vegetation of the central Missouri Valley and contacts of woodland with prairie. Ecological Monographs. 30(1): 37-64. [275]

52. Wikum, D. A.; Wali, M. K. 1974. Analysis of a North Dakota gallery forest: vegetation in relation to topographic and soil gradients. Ecological Monographs. 44: 441-464. [28291]

53. Wilson, Roger E. 1970. Succession in stands of Populus deltoides along the Missouri River in southeastern South Dakota. The American Midland Naturalist. 83(2): 330-342. [25441]

54. Zasada, J. 1986. Natural regeneration of trees and tall shrubs on forest sites in interior Alaska. In: Van Cleve, K.; Chapin, F. S., III; Flanagan, P. W.; [and others], eds. Forest ecosystems in the Alaska taiga: A synthesis of structure and function. New York: Springer-Verlag: 44-73. [2291]

55. Zimmerman, Gregory M. 1981. Effects of fire upon selected plant communities in the Little Missouri Badlands. Fargo, ND: North Dakota State University. 60 p. Thesis. [5121]

56. Norling, Bradley S.; Anderson, Stanley H.; Hubert, Wayne A. 1992. Roost sites used by sandhill crane staging along the Platte River, Nebraska. The Great Basin Naturalist. 52(3): 253-261. [20102]

57. Adams, Dwight E.; Anderson, Roger C.; Collins, Scott L. 1982. Differential response of woody and herbaceous species to summer and winter burning in an Oklahoma grassland. The Southwestern Naturalist. 27: 55-61. [6282]

58. Severson, Kieth E.; Boldt, Charles E. 1978. Cattle, wildlife, and riparian habitats in the western Dakotas. In: Management and use of northern plains rangeland: Reg. rangeland symposium; 1978 February 27-28; Bismarck, ND. Dickinson, ND: North Dakota State University: 90-103. [65]

Related categories for Kuchler Type: Northern floodplain forest